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Transcript of GEORGE B. JOHNSON Copyright ©The McGraw-Hill Companies, Inc. Permission required for reproduction...
GEORGE B. JOHNSON
Copyright ©The McGraw-Hill Companies, Inc. Permission required for reproduction or display
PowerPoint® Lectures prepared by Johnny El-Rady
19 The Animal Body And How It Moves
Essentials ofThe Living
WorldFirst Edition
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19.1 Innovations in Body Design
Four key innovations have led to the diversity seen in the phylum Animalia
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In radial symmetry, the parts of the body are arranged around a central axis
In bilateral symmetry, the body has a right and left half that are mirror images of each other
Allows different organs to be located in different parts of the body
Ultimately led to cephalizationEvolution of a definite head and brain area
Radial Versus Bilateral Symmetry
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The body cavity supports organs, distributes materials and fosters complex developmentThree body arrangements
Acoelomates: No body cavityExample: Sponges and flatworms
Pseudocoelomates: Body cavity (pseudocoel) forms between the endoderm and mesoderm
Example: Nematodes
Coelomates: Body cavity (coelom) forms entirely within the mesoderm
Example: Mollusks, arthropods and vertebrates
No Body Cavity Versus Body Cavity
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Segmentation is the subdivision of the body into segments
Two advantages result from embryonic segmentation1. Each segment may develop a more or less complete set of adult organ systems
2. Locomotion is far more effective because of increased flexibility of movement
Nonsegmented Versus Segmented Bodies
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Deuterostomes evolved from protostomes > 630 myaWhile both are coelomates, they differ in three aspects of embryonic growth
Protostomes Versus Deuterostomes
Protostomes Deuterostomes
How cleavage forms a hollow ball of cells
Spiral cleavage Radial cleavage
How the blastopore determines body axis
Blastopore forms the mouth
Blastopore forms the anus
How developmental fate of embryo is fixed
Determinate cleavage
Indeterminate cleavage
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TABLE 19.1
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TABLE 19.1
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19.2 Organization of theVertebrate Body
All vertebrates have the same general architecture:
Food flows through a long tube from mouth to anus Tube is suspended in coelom, which is divided into
Thoracic cavity – Heart and lungsAbdominal cavity – Stomach and intestines
Body is supported by a skeleton made up of jointed bones
The skull protects the brainThe vertebral column protects the spinal cord
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TissuesCells of the same type are organized into tissuesTissues form as the vertebrate body developsThree fundamental layers of cells
Endoderm ; Mesoderm ; EctodermThey differentiate into > 100 different cell types
Adult tissues are grouped into 4 general classesEpithelialConnective MuscleNerve
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Fig. 19.1 Vertebrate tissue types
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OrgansStructures composed of several different tissues grouped into large structural and functional units
Organ systemsGroups of organs that work together to carry out an important function
Fig. 19.2 Levels of organization within the vertebrate body
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Vertebrates contain 11 principal organs systemsSkeletal system
Bones, skull, cartilage, ligaments
Circulatory systemHeart, blood vessels, blood
Fig. 19.3
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Vertebrates contain 11 principal organs systemsEndocrine system
Pituitary, adrenal, ductless glands
Nervous systemNerve, sense organs, brain, spinal cord
Fig. 19.3
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Vertebrates contain 11 principal organs systemsRespiratory system
Lungs, trachea, other air passages
Immune systemLymphocytes, macrophages, thymus, lymph nodes
Fig. 19.3
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Vertebrates contain 11 principal organs systemsDigestive system
Mouth, esophagus, stomach, intestines
Urinary systemKidneys, bladder, associated ducts
Fig. 19.3
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Vertebrates contain 11 principal organs systemsMuscular system
Skeletal, cardiac and smooth muscles
Integumentary systemSkin, hair, nails and sweat glands
Fig. 19.3
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Vertebrates contain 11 principal organs systemsReproductive system
Testes or ovariesAssociated structures
Fig. 19.3
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19.3 Epithelium is Protective TissueThe vertebrate body consists of one tube (digestive tract) suspended into another (body cavity: coelom)
The outside of the body is covered with cells (skin) derived from embryonic ectoderm tissue
The body cavity is lined with cells derived from embryonic mesoderm tissue
The hollow inner core of the digestive tract is lined with cells derived from embryonic endoderm tissue
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Epithelial cells are produced from all 3 germ layersCollectively called the epitheliumThree important functions
1. Protect tissues from dehydration and mechanical damage
2. Provide sensory surfaces
3. Secrete materials Fig. 19.4
Gila monster
Tough, scaly skin
Epithelium possesses amazing regenerative abilities
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Epithelial cells are classified into three types according to their shape
Squamous
Cuboidal
Columnar
Table 19.2
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There are three general kinds of epithelial tissueSimple epithelium
Only a single layer thickFound in the lining of the lungs and major body cavities
Stratified epitheliumSeveral layers thickFound in the skin
GlandsInvolved in secretionEndocrine glands secretes hormones into the bloodExocrine glands use ducts to secrete sweat, milk, saliva and digestive enzymes out of the body
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19.4 Connective TissueSupports the Body
Connective tissue is derived from the mesodermThree functional categories
1. Immune systemBody defense
2. Skeletal systemBody support
3. Blood and fat cellsStorage and distribution of substances
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Two principal immune cells are
Immune Connective Tissue
2. LymphocytesMake antibodies
OrAttack virus-infected or cancerous cells
1. MacrophagesEngulf and digest invading microbes
Table 19.3
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1. FibroblastsThe most common kindSecrete structurally strong proteins such as collagen into spaces between cells
Skeletal Connective Tissue
2. Cartilage Collagen matrix forms in long parallel arrays along lines of mechanical stressFound in joint surfaces
3. BoneCollagen fibers are coated with calcium phosphate
Table 19.3
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Includes
Storage and Transport Connective Tissue
Erythrocytes (RBC)Transport O2 and CO2 in the blood
Adipose tissueAccumulates fat
The fluid portion of blood is called plasma
Table 19.3
Contains nutrients, wastes and antibodies
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Bone is a dynamic tissue that is constantly being reconstructed
Outer layer is very dense and compactCalled compact bone
Interior is a more open lattice structure Called spongy bone
A Closer Look at Bone
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New bone is formed in two stages1. A matrix of collagen fibrils is secreted by cells called osteoblasts2. The fibrils are impregnated by crystals of a calcium phosphate mineral called hydroxyapatite
Bone is laid down in thin layers around a narrow central channel
This central canal (or Haversian canal) runs parallel to the length of the bone
A Closer Look at Bone
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Fig. 19.5 The structure of bone
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Two cell types are responsible for bone “remodeling” during growthOsteoblasts
Deposit boneOsteoclasts
Secrete enzymes that digest the bone matrix
OsteoporosisExcessive bone lossUsually associated with aging Fig. 19.6
Equal osteoblast and osteoclast
activities
Excess osteoclast activity
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19.5 Muscle TissueLets the Body Move
The distinguishing characteristic of muscle cells is the abundance of contractible protein fibers
These microfilaments (myofilaments) are made up of actin and myosin
Muscle contraction occurs when actin and myosin slide past each other
The vertebrate body possesses three different kinds of muscle cells
Smooth Skeletal Cardiac
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Cells are long and spindle-shapedEach contains a single nucleus
Cellular microfilaments are loosely organized
Smooth Muscle
Power rhythmic involuntary contractions
Table 19.4
Sheets of cells
Found in the walls of blood vessels, stomach and intestines
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Produced by fusion of several cells at their endsThis creates a very long muscle fiber that contains all the original nuclei
Skeletal Muscle
Microfilaments are bunched together into myofibrils
Found in voluntary muscles
Power voluntary contractions
Table 19.4
Striated
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Fig. 19.7 A muscle fiber, or muscle cell
Modified ER
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Composed of chains of single cells, each with its own nucleus
Chains are interconnected, forming a latticework
Cardiac Muscle
Each heart cell is coupled to its neighbors by gap junctions
Table 19.4Allow electrical signals between cells
Cause orderly pulsation of heart
Striated
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19.6 Nerve TissueConducts Signals Rapidly
Nerve tissue is composed of two kinds of cells
1. Neurons Specialized for the transmission of nerve impulses
2. Glial cellsSupport neurons with nutrients, support and insulation
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19.6 Nerve TissueConducts Signals Rapidly
Neuron plasma membranes are rich in ion-selective channels
These maintain a voltage difference between the cell’s interior and exterior
Depolarization is the temporary loss of this voltage difference
It results in a wave of electrical activity, or nerve impulse
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Each neuron consists of three parts
1. Cell body – Contains the nucleus2. Dendrites – Bring nerve impulses to the cell3. Axon – Carry nerve impulses away from the cell
Fig. 19.8
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Neurons are separated by tiny gaps termed synapses
Neurons communicate by passing neurotransmitters across these synapses
Note: A nerve is made up of the axons of many neurons
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TABLE 19.5
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19.7 Types of SkeletonsThere are three types of skeletons in animals
1. Hydraulic skeletonFluid-filled cavity encircled by muscle fibers
Found in soft-bodied invertebrates like jellyfish
Fig. 19.9
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2. ExoskeletonRigid hard case surrounding the bodyFound in arthropods
3. EndoskeletonRigid internal skeleton to which muscles attachFound in vertebrates and echinoderms
Fig. 19.10
Rock crab
Exoskeleton made up of
chitin
Fig. 19.11
Endoskeleton made up of
bone
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Made up of 206 individual bones
Axial skeletonSupports the main body axisSkull, backbone (spine) and rib cage80 bones in all
Appendicular skeletonSupports the arm and legsPectoral and pelvic girdles126 bones in all
The Human Skeleton
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Fig. 19.12 Axial and appendicular skeletons
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19.8 Muscles and How They Work
The major human muscles include the following
Fig. 19.13
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19.8 Muscles and How They Work
Skeletal muscles are attached to bones by straps of connective tissue called tendons
Bones pivot about flexible joints pulled back and forth by attached muscles
The origin of the muscle is the end attached by a tendon to a stationary bone
The insertion is the end attached to a bone that moves during muscle contraction
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Muscles in movable joints are attached in opposing pairs
Flexors retract limbsExtensors extend limbs
Fig. 19.14
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Limb movement is always the result of muscle contraction
Never muscle extension
There are two types of muscle contraction
IsotonicMuscle shortens, thus moving the bones
IsometricMuscle does not shorten, but it exerts a force
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Myofilaments are made up of actin and myosinActin filaments consist of two chains of actin molecules wrapped around one anotherMysosin filaments also consist of two chains wound around each other
One end consists of a very long rodThe other consists of a double-headed globular region or “head”
Muscle Contraction
Fig. 19.15
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An ATP-powered myosin head-flex mechanism allows the actin filament to slide past myosin
This causes myofilament contraction
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Fig. 19.16 How actin and myosin filaments interact
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In vertebrate skeletal muscle, contraction is initiated by a nerve impulse
Nerve fibers are embedded in the surface of the muscle fiber forming a neuromuscular junction
When a signal reaches the end of a neuronThe neuron releases acetylcholine into the gap between neuron and muscle
This causes depolarization of the muscle cell
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When a muscle is relaxed, attachment sites for myosin heads are blocked by tropomyosin
Role of Calcium Ions in Contraction
For the muscle to contract, tropomyosin must be moved by another protein called troponin
The troponin-tropomyosin complex is regulated by calcium ion concentrations in the muscle cell
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Absence of Ca++ Muscle is relaxed
Muscle contracts
Fig. 19.17 How calcium controls muscle contraction
Presence of Ca++
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Muscle fibers store Ca++ in the sarcoplasmic reticulum
Role of Calcium Ions in Contraction
Nerve activity causes the release of Ca++ and ultimately muscle contraction
Fig. 19.18