Five New Combinations in Gossia (Myrtaceae) from Melanesia

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Five New Combinations in Gossia (Myrtaceae) from Melanesia Author(s): Neil Snow Source: Novon, Vol. 15, No. 3 (Sep., 2005), pp. 477-478 Published by: Missouri Botanical Garden Press Stable URL: http://www.jstor.org/stable/3393498 . Accessed: 15/06/2014 11:14 Your use of the JSTOR archive indicates your acceptance of the Terms & Conditions of Use, available at . http://www.jstor.org/page/info/about/policies/terms.jsp . JSTOR is a not-for-profit service that helps scholars, researchers, and students discover, use, and build upon a wide range of content in a trusted digital archive. We use information technology and tools to increase productivity and facilitate new forms of scholarship. For more information about JSTOR, please contact [email protected]. . Missouri Botanical Garden Press is collaborating with JSTOR to digitize, preserve and extend access to Novon. http://www.jstor.org This content downloaded from 194.29.185.216 on Sun, 15 Jun 2014 11:14:33 AM All use subject to JSTOR Terms and Conditions

Transcript of Five New Combinations in Gossia (Myrtaceae) from Melanesia

Page 1: Five New Combinations in Gossia (Myrtaceae) from Melanesia

Five New Combinations in Gossia (Myrtaceae) from MelanesiaAuthor(s): Neil SnowSource: Novon, Vol. 15, No. 3 (Sep., 2005), pp. 477-478Published by: Missouri Botanical Garden PressStable URL: http://www.jstor.org/stable/3393498 .

Accessed: 15/06/2014 11:14

Your use of the JSTOR archive indicates your acceptance of the Terms & Conditions of Use, available at .http://www.jstor.org/page/info/about/policies/terms.jsp

.JSTOR is a not-for-profit service that helps scholars, researchers, and students discover, use, and build upon a wide range ofcontent in a trusted digital archive. We use information technology and tools to increase productivity and facilitate new formsof scholarship. For more information about JSTOR, please contact [email protected].

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Page 2: Five New Combinations in Gossia (Myrtaceae) from Melanesia

Five New Combinations in Gossia (Myrtaceae) from Melanesia

Neil Snow Department of Biological Sciences, University of Northern Colorado, Greeley, Colorado 80634,

U.S.A. Current address: H. T. Harvey 81 Associates, 423 W. Fallbrook Ave., Suite 202, Fresno, California 93711-6138, U.S.A. [email protected]

5(4). Branchlets 4-angled; leaves very narrowly ovate, apex apiculate ........... G. randiana

5'. Branchlets round; leaves narrowly ovate to ovate, apex acute ..................................... 6

6(5). Stipular "hairs" obscure (or if present then not swollen at base); style hairy at least near base

, . .

........................ ..................... (J. aneztyenszs

6'. Stipular "hairs" prominent and greatly swollen at base; style glabrous ...................... G. floribunda

1. Gossia randiana (Merrill & Perry) N. Snow, comb. nov. Basionym: Myrtus randiana Merrill & Perry, J. Arnold Arbor. XXIII: 237. 1942. TYPE: Papua New Guinea. Central Division, Kubuna, Dec. 1933, L. J. Brass 5642 (holo- type, A; isotype, BRI).

The holotype specimen has two illustrations in pencil, presumably by Merrill or Perry, that illus- trate two characters which in part diagnose Gossia from related genera. The first is the circinate em- bryo, the second is the gelatinous endosperm seen associated with the embryo (Snow et al., 2003).

2. Gossia eugenioides (A. J. Scott) N. Snow, comb. nov. Basionym: Myrtus eugenioides A J. Scott, Kew Bull. 33: 514. 1979. TYPE: Indo- nesia. Manokwari, Vogelkop, Arfak Mts., Angi gita lake, 2800 m, A. Kostermans 2398 (holo- type, K [photos at BRI, GREE]; isotype, L [photos at BRI, GREE]).

The original description by Scott (1979) indi- cates only two collections, both from Irian Jaya. In the key to fertile specimens (Snow et al., 2003: 32), Gossia eugenioides would key out most closely to Gossia shepherdii (F. Mueller) N. Snow & Guymer, but it differs most readily from G. shepherdii by its moderate to densely sericeous hypanthium and ev- ident lack of smooth bark. Further information is needed about the biology and distribution of this specles.

3. Gossia versteegllii (Merrill & Perry) N. Snow, comb. nov. Basionym: Myrtus versteeghii Mer- rill & Perry, J. Arnold Arbor. XXIII: 238. 1942. TYPE: Indonesia. Irian Jaya, 9 km NE of Lake Habbema, 2800 m camp, Oct. 1938, L. J. Brass 10618 (holotype7 A; isotypes, BRI, K, L).

ABSTRACT. Based on characters of the fruit, seed, and embryo, five new combinations in Gossia N. Snow & Guymer (Myrtaceae) are proposed for spe- cies occurring in New Guinea, Vanuatu, or the Sol- omon Islands. These species previously all had been placed in Myrtus L. or Austromyrtus Burret. A lectotype is selected for Gossia aneityensis. A key is provided for the Melanesian species of Gossia.

Key words: Austromyrtus, (:;ossia, Melanesia, Myrtaceae, Myrtus, Solomon Islands, Vanuatu.

Recent revisionary work in Myrtaceae has led to the splitting of Austromyrtus s.l. into Austromyrtus s. str., Lenwebbia N. Snow & Guymer, and Gossia N. Snow & Guymer (Snow et al., 2003). Gossia is most species-rich in Australia and New Caledonia, where together approximately 42 species occur (Snow, 2000; Snow et al., 2003). The author is pres- ently revising species of Gossia for the Flore de la Nouvelle-Caledonie and has had the opportunity to reassess boundaries of all known species of the ge- nus. The following species currently placed in Myr- tus or Aatstroreyrtals are all easily attributable to Gossia and are here transferred to that genus based on the amended descriptions of Austromyrtus, Myr- tus, and Gossia (Snow et al., 2003). Gossia f oribun- da (A. J. Scott) N. Snow & Guymer, which is most common in Australia, also occurs in New Guinea (Snow et al., 2003).

KEY TO THE SPECIES OF GOSSIA FROM NEW GUINEA, THE SOLOMON ISLANDS, AND VANUATU

1. Flowers 5-merous .............................. 2 1'. Flowers 4-merous .............................. 4 2(1). Branchlets rounded .............................. G. eugenioides 2'. Branchlets 4-angled .............................. 3 3(2). Stipular "hairs" prominent, very elongated api-

cally; leaves broadly elliptic7 12-28 mm long; hypanthium densely appressed-pubescent . . .

........................ ........................ G. versteeghii

3'. Stipular "hairs'7 somewhat obscure, only some- what elongated apically; leaves narrowly ellip- tic to ovate, 15-75 mm long; hypanthium gla- brous to sparsely sericeous ......................... C. floribalnda

4(1). Axis of inflorescence with short, spine-like bracts . G. salomonensis

4'. Axis of inflorescence lacking spine-like bracts

............................... ................................ 5

NOVON 15: 477-478. PUBLISHED ON 21 SEPTEMBER 2005.

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Page 3: Five New Combinations in Gossia (Myrtaceae) from Melanesia

478 Novon

(Xtlillclulllill (19tS1: 2,44) in(li(clte(l thclt the ('0111-

1llon ncilile of this sipe(ies is :\is-ie.'' The seleete(l le( tolx pe is the first of tss-o spe(ililelis nielitionecl in the pl otolocgue hie h I hclx e seen cin(l s hieh is -e^)leselitciti-e of the s)ee ies.

A(Aslolzle(lgmerlt*N. Illclllks to Exe l,uccis cit the lto!-cil }30tcinie (^cir(lens. Ke^X- (K). for sending cl dig- ikil ililare of the t!-pe spee imell of G. .s(llomc)nensis; Chlistialle'l'ilel cin(l Ce(ile Aupi( at the Museum Ncitional (I'Histoire INTaturelle (P) for assistanee in the herlcil-ium; and the (uratols of A BRI, K, 1,, \1 k,1,. al(1 P for permission to l)ol los or X ieuT spee iniells. 'rhe (omments of ts-o anollxnious rexiesers an(l Xi(toria Hollos-ell (larifietl the malluseript.

I itelcittile Cite(§

(}ExilllvXillt A. 1931. (Lolltl-it)ltiolls to the flola of the iNew Heti(les: Plulltx (olle(te(t t!- S. Ft Kajessski in 1928 a,(1 1 (329. J. Aw-rlol(l Rl t)ol. 1 2: 2:2(WS64.

S( ott \. J. 19. 9. I\c-Xs spe( ies all(t t olllbinatiorls ill MX I ta(eae flolll 01alesia al(1 .\ust-alia. Kess- Btlll. 33: all- .,1.,.

Sllos. y . ;2()()(). .\ ( onspec tus of Australasiall M) rtinae (01!1-ta(eae). Ke^s- Bull. ao: 647-6tz4.

, (J. P. (J[l\'lllel' &- G. Sas-\-e1. 223. S-stemati(s of All.stron7 1 rtlls. lwn7l ebbifl an(t the Xustralian spe( ies of (,ON.Si(l (M!-l-tateae). S!st. 130t. Mollogl. 6o: 1-9tz.

The stipular X hails 9 of Gossi(l l ersteegtlii aIe unique in the genus. As s-ith the t+-pe Bs stiyoules discussed in Snow et al. (2003: 7)^ the- 11as e a

prominently swollen base but they differ in hax ing a greatly attenuated apex and can be up to 2 mm long.

4. Gossia salomonensis (A. J. Scott) N. SllOU.

comb. nov. Basionym: Mlrtls s(llonlorletlsis A. J. Scott, Kew Bull. 35: 412. 198(). TYPE: Sol- omon Islands. Gizon Islanclt R. Hflllriflsi et fll. BSIP 11740 (holotype, K [seen as digital im- age]).

The species is distinct in Gossifl b! sirtue of the number of short spinose bracts oecurring in the in- 13orescence, which are typically aggregate(l neal- tlle base of the pedicel; other species in the genus t-p- ically have none or only a few such bracts.

5. Gossia aneityensis (Guillaumil) .N. Snos- comb. nov. Basionym: lll-rtlls (lneit-ensis (uil- laumin, J. Arnold Arbor. XII: 2a4. 1931. Alls- tromyrtus aneitAensis (Guillaumin) Burret, -No- tizbl. Bot. Gart. Berlin-Dahlem la: 506. 1941. TYPE: Vanuatu ( = New- He}rides). Anelgau- hat Bay, 23 Feb. 1929, S. J. Kflielxski 810 (lec- totype, designated here, P; photos at BRI. GREE; isotypes, BRI, K [photo], MEL).

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