Coloquios de Palcontologia, 2002,53: 33-68 An annotated...

Coloquios de Palcontologia, 2002,53: 33-68 ISSN: 1132-1660

An annotated check-list of genera and species of carpoids

Lista anotada de los géneros y especies de carpoideos

Patricio DoMíNcuEz ALONSO1’2, Richard RS. JEFItERIES1 & Dolores GIL CID2

Ábstract: Tisis ciseek-list includes liso al presoní valid names of genera and species of liso carpoids. It is a provisional atíempí lo stabili-ze tse names of MI fornially described carpeids up te date and includes mosí of Ihe combinations of genera and species that bayo beenused in tSe seicotifie literatsjre, It sisould be especially useful re tisose roading oidor papers in wbicis awide variety of genera and spe-cies Sayo beco used, and te tisose iriterested in biodiversity, evelutionaiy studies or isistory of the palaeontology. Tise ciseclc-list is divi-ded mío five sections: 1. índex ofgenera; II. índex of species; Hl. índex of valid speciflc epithets; IV. Tables of stratigrapisical occurrenceand y. Tiso mosí complete and up-to-date lisí of references en Ihe taxonomy of carpoids.Key words: ciseck-lisl, index nonrioorrrnr, Seluta, Cornuta, Mitrata, Ctenocystoidea, Cineta, calciciserdates, isomalezoans, stylopisorans.

Resumen: Esta lista recoge los génerc>s y especies actualmente válidos de carpoideos. La lista incluye la práctica totalidad de las cera-binaciones de género y especie que San sido empleadas en la literatura científica. No es una lista exhaustiva de sinonimias pero se ofre-ceo referencias clave para elaborarlas. Este trabajo debería sordo utilidad para aquellos lectores interesados en obras antiguas donde unaconfusa variedad de formas son descritas o referenciadas y para aquellos interesados en paleodiversidad, procesos evolutivos o isistoriade la paleontología. La lista se encuentra dividida en cinco secciones principales: 1. Índice de géneros; II. índice de especies; III. Indicede epítetos específicos válidos; IV. Tablas de distribución estratigráfica y V. la más completa y actssaalizada relación de referencias biblio-

gráficas sobre taxomía de carpoideos.Pajabras clave: indice, índex norniooru,n, Soluta, Comuta, Mitrata, Ctenocystoidea, Ciocta, calcicordados, Homalozna, Stylopisom.

INTRODUCTION

The carpoids are an informal grouping of Palaeo-zoic fossils wisich hayo a skolelon ofecisinoder¡n-likecalcito buí Iack radial symmetry based 00 tSe watervascular system.

They aro isighly controvorsial being regarded bysorne as aberrant echinoderms and by others as mem-bers of liso stem groups of many of Ihe major sub-groups of deuterostomes (craniales, tunicates, acra-niales, chordates and conceivably of tSeechinodem~s, isemichordates, ambulacraria and por-haps of Iho deuterostomes lhemselves).

TSe aims of Ihe prosoní work are bibliographicalso we do nol consider the stem-group position of tSevarious subgroups of carpoids. Instead, we use asgroups liso Jaokolian orders plus liso ctonocystoidsand within tiseso groups wo arrange tSe genera andspecies in alphabetical ondeE

In Ihis chock-list a total of 179 species of carpoidsaro alphabetically rocorded by Classes (including 22solutos, 52 cornutos, 75 mitrales, 24 cinctans and 6ctenocystoids). It is a provisional attempt lo stabilizebe names of alí forrnally described carpoids up lodate and includes mosí of the combinatioras of gene-raand species lhat hayo beon usod in bibliography. Itshould be especially useful for tisose reading oidorpapers in which a wide variety of genera and spociesSayo been usod, and lo thoso interested in biodiver-sity and evolutionary sludies. When placing speciesin particular gonera we nocessarily oxpress our ownpersonal opinions. No attempt at pisylogenetic orde-ring has been made. TSe sequence is puroly alpisabe-tic. TSe check-list comprises five main sections:

1. índex of goneraII. índex of speciesIII. índex of valid specific namesIV. Tables of stratigraphical occurrence

¡ Departmení of Palaoontology, TSe Natural History Museuni, London SW7 SBD, 1.3K. [email protected]; rpsj@?nhmacukDepartamento de Paleontología & Instituto de Geología Económica (UCM-CSIC), Fac. CC. Geológicas. Universidad Complu-

tense de Madrid, 28040 Madrid, Spain. [email protected]; [email protected]

33

/‘esíiioi’s í)or.5551555c: Alsossto & ¿5:[]. Xis sistroraled check-lisl of gersera asid species of carpoids

‘e,‘5,

31

505 55 90 50 909209 9409 09c01975519990515520®

Fietsre 1- Cusnulaíi»e nuortrbcr of al proseo] das salid described

sísec¡es ssf carpoids por sean./-eQisres 1,— :Vsose’ro eseíonu!¿it/ss dc copes ‘es así ro/mcta le ió!ie/csss/’ eosj’oielc:os s/sas 54<55 usi ¿sises.

TSe ciseck-list is based típon valid iaxenoniic lite-roture necords ond is arranged taxonomicaíly by Class,ISeo alpisobetically by Gonus and Spoeios witbin oacbClass. Entiles o liso hijeo ofGenero (Section 1) inclu-de: Genus; A utisor(s) wiso desenibed liso genus; dato ofliso original pníblication; type species (using uSe origi-nal spelling, ciseck So Índex of .spec:icfl: ouínber ofspecios: included specios (¡So syrnbel e”’» denotes lisolype species); stratigrapisical occurrenco; goegrapisicaldistnibiruion; kev relerenees. Entries in the Índex oJSpeeics (Seclion II) includo: combination of genusondspecicsúscientific binomir.a);Autisor(s} wisodescnibedriso species (in pa¡entiseses if liso specros rs new ossre-ned lo a clifforetñ genus tisan wison desenibed bv aul-

Sor): date r>f Ibe original pubí ication; autisor(s) of tSenew combí o<tlron oric~ínol name; occuirronco, distnibu—tion. kev ictorenccs Also included ~500 Indo» oJspe-

‘sanie 5 (Suclion III), wisore liso val id spocifrcriamos of oíl e arpords are lisled and referred lo lisogonera. Some spo<rlic epitiscís Sayo beco used in seso-rol gesiora. No separatron oto classes Sas been attemp-lcd i o liso hrelc of 5pc llj!c~ nonios wisicis is lisereftreenrorely alpisabeiical.

Supragenerie taso aro oot considered Soro. excepílisose lraditionally considered as classes (j.c. tSe for-mor JaeLel ion ordens plus tSe alen doscribed cíe-r>ocvstoids), TSe recomniended nomos are sisown inboU type. TSe oíd nomos (incor-recí spollings. rice-ireer goríder. no/nielo sano. nonhilso dubio, juniorsviasuytas. etc.) are sbown in squaro brackers «1TSe symbol «~ss indicates SoIS a synonyns and a crossreference inside liso indos; picoso cisock ilIon furtisor

Gsñrsc¡sócss lIs’ /‘os)esnílolrsgíss

Ficure 2.- Valid described species of carpoids per decado (iSicklino) and cutnulati»e percenlage sf »alid described species pordecade (lisio I¡ae).figura 2.- Espee:ics válidas dc earpoideos descritas por décadas(en liesea gruesa) y isoreeniaie acumulado de especies válidasele’scritas por clúcades (línea tina).

dala. Tho syrnboí sc >» associated with a namo emp-isasizes tSe eross-reference íisat musí be followed.

Tubbs oJsírodgraphieol occíctrence (Sectien IV) listte spocies ordeíing tisom by suratigropisical oceurrenceand ISeo, witisin caeS suage, o alpisabetical orden Dala[br tSe stages aro based en Fo.ssil Record 2 BENTON,i993 in orden lo relato oursuratigrapisical dala witb equi-valeor doto of otisor groups witiseut preposing a newworldwide siraligrapisical correlation. Re nomos of lisosíagos are also reforrod lo in Soctiens 1 and II. Recentlydoscribod spocios are usually not diffrcult te assign lo aparticular siage. Olderspocieshíwcvcr~ especiahlytho-se based 00 Sistorical colleclions with only a generallecation in tSe bibliegrapisy or in Iho labels accompan-ying tSe specimens, are diffrcull lo assign stratigraphi-cally. lo tiseso casos liso species oppoar in liso labIo witha quoslion mark «7», or in grey stipple.

Tisis índex was cempilod from eur ewo notos ondbibliograpisie records and subsequently completodand cisocked by using external databases sucis asGeoRcf and Zoelogical Record (boris from Silvor-Plalter Information N.y.) and general soareises en lisoInternet. We did nol make an exisaustivo search in lisocomplote printod version of tSe Zeological Record.

lo addition te lisis papor version. tisis cisoek-Iist italso ovailable al liso web sites of our instituticos alDepartctmeole do Paleontología, UCM, Madrid<hltp://www.ucm.os/iofo/palee/carpos> and TSeNatural History Museum <Sltp://wwwnhm.ac.uk/palaeenlology/carpoids> wisoro it will be periodi-calíy updalod.

Typical eníries in tSe cisock-list retad os fellow:

50

540

a30

a20

5

2 10

100%

9080

70

605040

30

20

10

o~‘II

0000 0 0000 0000

<lItIo 5 35-61=34

Patricio Domínguez Alonso & el al, .Nn annotaied ciseck-iist of gerrera asid spocies of caxpoids

Example 1, a genus nanie:

Lagynocystis JAnKEL, 1918TYI’e 5PECiES: Ano,nalocystites pyrarnidalis (BARRANDE,

1887)NUtrSABER OF srecies: 1INCLUDEI) seecios: Lagynocysíis pyrarnidalis

t (BAima~oe,

1887)STRATiGRAPHicPJ. OCCURRENCE: Lower-Middle Ordovician(Llanviru)GEocítAprircAL DiSIRiBUTiON: Europe (Bohemia, France,Spain)REFERENCES: BAi&4y0e, 1887; CHAUvEL, 1941; CHAUvEL &NíeN, 1977; JAEKEL, 1918; JEFFI5RrFS, 1973; PAR5LEY, 2000;UBAcris, 1991

JAEKEL, 1918 proposedthe genus Lagynocystis forAnomalocystites pyramidalis (BARRANDE, 1887)(which incidentally became type specios bymonotypy). Lagynocystis JAEKEL, 1918 is in boldtype, showing a valid genus natne; thore is only onovalid species doscribed (we inusí search in Ihe Check-

lisí ofthe species naenes); te specios includod is Lagy-nocystis pyramidalis (BARRANDE, 1887), te symbol *

indicates tal Ihis is tho typo species of te gonus; teslratigraphical occurrence and liso geographieal distri-buhen are given. Finally, somo roferonces are usted.

Example 2, a salid species name:

Sucocystis melendezi (SCHROEDER, 1973);FRIEDRIcE, 1993

[=Gyrocystis?,nelendezi SCOROEDER, 19731l=Troclíocinctus pardail/sanicus TERMíER & TErr=drER,1973]STRATicIRAPHrcAL OCcURRENCE: Middle Cambrian (St.David’s)GEooRAPurcAL DisTRinurrioN: Europe (Spain. France)REitRENCES: ScFrRoEnere, [973;F~rorwrcu, 1993

SCHROEDER, 1973 described Gyrocys/is? melendezi;

but lator FRIEDRICH, 1993 proposed te new combina-tion Sucocystis melendezi. Troehocinetus pardailhani-

cus TERMIER & TERMIER, 1973 is a junior synonym.Since Sucocystis melendezi (SCHROEDER, 1973) isconsiderod valid, it is lo bold type in tho check-Iisl;however bolh synonyms appear in squaro brackets«[ ]». The symbol «=» previous te Iho names indicateste synonym as well as a cross in te check-Iist.

Example 3, a species name no Ionger used:

[DendrnqstitesVldnIi NeRss, 19351=Minervaecys/is vidail (TIIORAL, 1935); UnAcris & CASFER,

1967 —>

REFERENCES: THORAL, 1935b; UBACHS & CASTER in CASTER,

1967

THoI¿AL, 1935 described a new Soluta as Den-drocystites 1/idali, a combination no longer in use(and Iherefore quoted in square brackels «[1») sinceUBÁGES & CASTER, 1967 ira Ihe Treatise on Inverte-

brate Palaeontology (CASTER, [967) proposed Ihogenus Minervaecyshs for this soluto and so Iho newcombination Minervaecystis vidalí (THORAL, 1935)became available. The symbol «=» before Minerva-

ecystis vidalí thus indicales Iho currently usedsynonym arad acts as a cross roference lo the cisock-lisí where moro information is available. The symbol«-~» emphasizes thaI cross roferonce. Note also diocapitalised «Vidali» as originally describod.

CHECK-LIST OF CARPOIDS

1. íNDEX OF GENERA

Cineta JAEKEL, 1918

Asturicysds SDZUY, 1993TYPE sPECrEs: Asturicystisjaekeli Sdzuy, [993NUMBER OF 5PECiE5~ 2INCLUDED spEctEs: Asturicystisjaekeli* SO/UY, 1993; A./íavliceki FATRA & KORDULE, 2001STRATIGRAPOICAL OCCURRENCE: Middle Cambrian (St.Davids)GEOGRAPHíCAL DiSTRIBUTiON: Europe (Spain: Asturias;Boisemia)REFERENCES: SDZUY, 1993; FATKA & KORDULE, 2001

[Cepí’íalocínctusTERMIER & TonseíioR, 1973]=Elliptocinctus TERMíER & TERMíER, [973 >

REFERENCES: TERMIER & TERMIER, 1973; FíuroDr<íCH,[993

[Cyclocinc/usTERMíER & TERMIER, 1973]parima =Elliptocinctus TERMíER & TERMiER, 1973

lElliprocincrus barrandei (MIsNíER-CHALMAs & BERGE-RON, 1889)Jpartimn =7 Sucocystís CABraEL, TERMIER & TER>AíER,1959 [Sucocystis?enaroccana (TERIIíER & TERMíER,1973)1 -9<

REPERENCES: TERMíER & TERMIER, 1973; FRrEDiaicH,1993

Davidocinctus FRIEDRICE, 1993Tv~o 5PECiE5: Davidocinctr¿s pembrokensis FrucoRieR, 1993NUMBER OF spoci~5: 1INCLUDED svrserrss: Davidocinetus pe,nbrokensis* FRirsoRien,[993

Coloquios de Paleontología2002, 53: 33-68

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Patricio Domínguez Alonso & ci al, An asisrosated chcck-lisí of genera and species of carpoids

STRATic,rcAi’urcÁr occi:RRENce: Middle Cambrian (St.David s)Gí¿:ocrowrsícAt, DIsTRiBUTiON: Europe (UK: Wales)Rer:ertoscms: FRreoRrcH, l 993

[DecacestisGíSLÉN, 1927]

=Gvrocystis JAF.KíuI., 1918ReFeRísNí:rzs: FrsieríRicrr. 993; GíSLÉN, 1927

Elliptocinetus TERMtER & TtiRMlriu, 973lyrt sríscres: Trochoeystitcs Rarrandel MUNíER-CHALMAS &BERGERON, 1889NoMBus OF SPEeiLs: 2lNr:LUoEo sPEerEs: Elliproí:inc/tas barrondei

9< (MUNrER-CírAr.-MAS & BIsRe;rsRor=.1889); EIlip/oeincvus vizcainoi FRreoRrt:ír,[995STRxnrovssPísrcAL oCc?IJRRENI7IS: Middlo Cambrian (St.David’s)Geot;RAroicÁs. Di5TRiOUTiON: Europe (Spain, Franco); Nerí-Seos AIrica (Morecco)REFERENcEs: Mí;srEa-CHAI,MAS & BERGERON in BCRGERoN,[889;TísuMrízíe & TI<RM[ER, 1973;FuiEngícu, 1993

Gyrocystis JAEKEL, 1918Typn spízcíes: Ge roeystis platessa JAEREL, 1918NuMiseR OF 51’FCib5: 6INCLUDID 5PECi[5: Gírocysris badalesiensis Fuirmnuico,1993: G. eruzae GIL & DoMÍrse,Uez, 1995; G. erecta FRíE-bRiOl, 1993; G. hispan/ea (GisrÉN, 1927), [dubiousspe-cies]; G. plales.set* JAEKEL, 1918; G. íestudíformis FRrE-i)RitAi, l993STrtsric.wwoicAí. O(CURRLNCE: Middle Cambrian (St.David’s)GÉdcít&14fi¿ÁL bétRiñÚTriYp¿ EúfoÑ (S¡SaiitFraoce)REFERENGES: JAGKtSL. 1918; FRieonJcii, 1993; Go. Cro &DOMÍNGUEZ, [994

Ludwigicinctus FRIEDRIcO, 1993tít 5PECiE5: Ludwigiei,rctas trancatos FRíeDRíco, 1993NísMnízR 01’ 5FEcrE5: 1lNctuJrsFo sítcííes: Ludwig/cinctus lruncatus* FrirEoRreo,1993STresriuRAurriesí. OGCISRRENCE: Middle Cambrian (St.David s)GEoGiZÁPHrCAL OiSTRiOiiTiON: Europe (Germany)RereRrNcu.s.FurLoRicH, 1993

Progyrocystis FRILDRtCH, [993Typiy smzcies: Progvrocvsíi.s disjaneía FRreríí<ieií, [993NI:srírr¿R Dr’ sPisc:iiss:iNCLUDED 5Pr¿CrF5: Progyrocystis disjancza9 FrerEoíercH, 1993STreATicgApmcesi. OccsjRRízNcE: Middlc Cambrian (St.David s)GeoGuAerírcAt. DiSTRiBsJOoN: Europe (Spain: Asturias)REFERENCES: FRrEuRrt:H. [993

Coloquios de Pe¿/eontología

Sotociuctus Síztsy, 1993Tyr’u sr’ociss: Sotocioctas aboghsi SDZUY, l993NUMBER OF .Sí’ECiE.Y 1INcLUOLO sr>ecItis: Sotociucías ubaghsi9< Sozev, 1993:STRATiIRAFHiCAi OCGURRENCE: Midd[e Cambrian (St.Davids)GeoGRAPuicÁl. DiSTRiBUTiON: Europe (Spain: Asturias)Ru¡r:íLNeLs, Suar’», 1993

Sucocystis CABIBEL, TERMIER & TERMIER, 1959Toc srEciF~: Sí¡co<vstis ¿lícrooensis CABíBEL, TERMiER &TCRMIER, [959NUMBER OF 5PEGiF.5~ 7lNcttJDF.D SPFLiE.S: Sueocys¿is acrofera FREORicH, 1993; 5.bretooi Fnííeoíucfl, 1993; 5?? ,naroeeaoa (TERMíER & Tau-NIrER, [973);5?. ,nelcodezi (Seelr=oLDr¿R, 1973): 5. quads-ieor-nata Fuier,uico, [993;5. theronensist CAurisEl,, TERMíER &TERMíER, 1959; £ unja/a FRrísr,Rrco, 1993STRAííGrsxí’rjrcsxL OGGtSRREN(?E: Middle Cambrian (St.Davids)GeoGuÁPoicÁL DiSTRiBUTiON: Europe (Spain; Franco); Non-Seos Africa (Merocco)REFERENCES: CAumer. eta!., 1959; FuisuRicH, 1993

[Troehocysti.snorra. van.]— Troehocvstiíes BARRANDE, 1887

Trochocystites BARRANDE, 1859TYPE SPEeií’s: Tros:hocvsri/es bohemicus BARRANDE, l887NUMrsER OF 5PECiii5: 1INcLsSOED sPeciEs: Troclrocystiícs bohemicus * BARRANDE,1887STRATíGRAPHíCAL OGCURRENGE: Middle Cambrian (St.flavids3

GEoGRAFHicM. Oi5TRiBUTiON: Europe (Boisemia)NOTE: Mosí early descriplions of Ciocta were given as Tro-elíocy.stitcs or TroclrocystisRrserssENcts: BARIG\NOE, 1859, [887;JAEREL, 1918; TErzMir¿u& TERMíER. 1973; FRiFDRico, [993;DOMíNGUEZ, 1999

TrochocystoUes JAEKEL, 1918Twr. SPEctr5S: Trochocys¡oides parvas JAEKEL, l91 8NUMBER OF 5PEGitS: 2INCLUDED SitúEs: Trochocysíoides parias> JAEKEL, 1918; Tplanas SDZUY, 1993STRxriGrtAprircAr, OcCURRENCE: Middlc Cambrian (St.Da»ids)GEOGRÁrtiCAL DiSTRiBUTiON: Europe (Boisemia, Spain)ReFaReNeas: JAEREL, 1918; Fuíeourco, ¡993; Soso», l993

Ctonocystoidea RonísoN & SPRINKLE, 1969

Ctenocystis RehuseN & SríiiNr=tE,1969TYPFi spaciíss: Ctenoc}’stis ulahensis RoisísoN & Sí’riiNKr±,1969

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Patricio Domí,rgrsez Alonso & 01 al. An arinotated check-iist of gorrera and species of caspoids

NUMBER OF sPEciEs: 4INCLUDED sPEciF.s: Cienocystis eolo don UBAOIIS & RouísoN,1988; C. jagol JELL, BURRET & BANK5, 1985; C. smithiUDAGOs, [987; C. utalíensis> RoísrsoN & SPr¿iNiwe, 1969STRATíGRAPHIIIAL OCCURRENCE: Middle Cambrian (St.David’s)GEOcrtár’uicsL OiSTRiBUTiI)N: Nertis Amorica <USA: Utah,Idaho); Australia (Tasmania); Europe (France)REFERENCES: RonisoN & SPRíNKLE, [969

Etoctenocyslis FATKA & KORDULE, 1985Tvít 5PECiE5: Etoctenocystis boheosica FATRA & KORDI5LE,1985NUMBER OF SPECiES~ 1INCLUDEO sPEeieS: Eloctenos:v.sui,s hoheenica> FATKA & KoR-DULE, 1985STRATIoRAPISrcAr. OCCItRRi5NCIÁ: Middle Cambrian (St.David’s)GEoGRAPurcAL DiSTRiBUTiON: Europe (Boisemia)REFERErsCes: FAEKA & Kor=ixss±,985; DoMiNríUez, l999

Jugoszovia L»íK & OaLowsKi, 1995TYPE 5PECiE5: .Iugoszosna usehueoeyathoides D~ii< &OruowsKi, 1995NUMBER OF 5PECiB5: 1INCLUDED 5PECiE5: Jugoszovio arehaeocvarhoides> D~iK &Caxowsra, [995STRATiGrosr’oicAL OCCURRENCE: N4iddle Cambrian (St.David’s)GEOGRAPOWAL DiSTRiBUTiON: Europe (Poland)REFERENCES - D2iK & DRaowsKr, [995;DOMÍNGUEZ, 1999

Soluta JAEKEL, [901

Belemnocyslites MILLER & GrsRLr±y,1894TYPE 5PEI?iE5: Bclemnocysíites weíherhyi MILLER & GURLEY,1894NUMBER OF 5P~cics: 1INCLUDED sPísciEs: Relemnocv.síi¿es weeherbyi* MiLLER &GURLEY, 1894STRATiGruxi’í]icÁL OCcURRENCE: Uppor Ordovician (Caradoc)GEoGRÁPHicA¡. DISTRiBUTiON: Nortis America (USA)REFERENCF5: MILLER & GURLEV, [894; PAR5LEY, [972;KOLxíA cl al,, l977

Castericystis UBACES & RoBísoN, 1985TvPE 5PECiE5: Gasiericystis vali UBAGH5 & Ronrsors, 1985NUMBER risc SFECiE5: 1INCLUDED 5FECiE5: Castericysíis valí> UBAGHS & Ronrsors,1985STRósiGasporcAL OCCURRENCI]: Middle Cambrian (St.David’s)GEOGRAPHíCAL DiSTRiBUIrON: Nortis America (USA)REFERENciES: UnAuHs & RonísoN, 1985: DALEY, 1996

Coleicarpus DALEY, 1996TYPE 5PECIES: Castericystis? sprinklei UBAGHS & RoarsoN,1988NUMBER OF 5PEIZiEs: 1INcLUDEY sPECIE5: Coleicarpus sprinklei> (UBAGHS & Rení-sON, 1988)STRATIGRAPIIICAL OCCURRENCE: Midd[e Cambrian (St.David’s)GEOGRAPHíCAL oI5TRiBUriON: NorIl, America (USA)REFERENCES: DALEY, 1996

Dehmicystis CASTER, 1967TYPE sPEciE5: Dendrocystiies (Dendrocystoides?) globalasDeusí, 1934NUMBER OF SPF.crí&5: 1INCLUDED SEECiES: Delrmicystis globalus> (DEosr 1934)STRATiÚízÁPurcsL OCCURRENCE: Lower Devonian (Emsian)GEOGRAPOICAL DiSTRIBUTrON: Europe (Germany)REFERENCES: DEOM, 1934; CASTER, 1967

[Dendrocystisnom, van.]= Dendrocystites BARRANDE, 1887 -4

Dendrocystites BARRANDE, 1887TYPE 5PECiE5: Cystidea sedgwicki BARRANDE, 1867NUMBER OF SEECiES: 2INCLUDED 5PECiFS: Dendrocystites barrandei (BATHER, 1913);D. sedgwicki> (BARRANDE, 1867)STRATrorcAí’tsrcAL Ot?cURRENcE: Europe (Bohemia)GEOGRAPHICAL DiSTRiBUTiON: Midd[e Ordovician (Llandeilo)Raí’ErwNcEs: BARRANDE, 1887; BATHER, 1913; Jsrva, 1918

Dendrocysto ¡des JAEKEL, 1918TYPF 5PECiES: Dendrocystis scotica BATHER, 1913NUMBER oc sPFCirtS: 1INCLUDED SPECIES: Dendrocysíoides scoiicus* (BATHER,1913)STRAnGRAPoIcÁí. oCCURRENCE: Upper Ordovician (Asisgilí)GEOORAPHíCAL DisTRiBUriON: Europe (UK: Scotland)RerosaNcEs: BATHER, [913; JAEKEL, 1918 JEPFERIES, 1990

Girvanicyslis CASTER, 1967 [1968]Ts~e SPEIíE5: Girvanicyslis baiherí CASTER, 1967 [19681NUMBER OF 5PECiE5: 2INCLUDED SPECiE5: Girvanicystis bat/ter? CASTos, 1968; G.casterí DALEY, [992STRATIGRAPIIICAL OCCURPENCE: Upper Ordovician (Ashgill)GeooRAPíircAL DiSTRiBUTiON: Europe (UK: Scotland; SouthWales)REFERENCES: CASTER, 1967; DALEY, 1992

lleckericystis GiL & CASTER, 1960TYPE SPECiE5: Dendrocysti/es kackersiana HECKER, 1940NUMBER OF SPECiE5~ 1INCLUDED Sí’ECiES: Heckericvstis kackersiana* (HECKER,1940)

37 Coloquios de Paleonlología2002, 53: 33-68

Patricio Dssnsíííguez A/o,ísss & O al

STítxTíc;aÁs’criíÁi, oeetsRRisNr:E: Middle Ordovician (Llandeilo)GEoGR’rííícesi, DrsTRinrsTioN: Ecropo (Russia: St. Pelersburgarea)RísnízaisNci.s, HíseKísa. 194(1; Gui, & CAs-res, 1961)

Jowacyslis THOMAS & LADD, 1926Ts~o suecias: lesrs’aeystis sagitiaria THOMAS & L>rno, [926NrssrrsoR ob sPtsr:sn.s: 1lNeLr:rSEL) 5PFtT7iE5: Ioss’aeystis sagittaria * TuoMÁs & LADO,¡926STaxrií;RsF’elrt:Ás, (Sec:r;RRENI:F:: tipper Ordovician (Caradoe-Asisgilí)GcOGRÁPí]rcst. t}r5TRiBUTrON: Nortis Amorica (USA: Iowa)REFERENCES: THoMAs & LADD. 1926; PxRsrísy & CASTER,[965;RelATA es al., [977

Maennilia RozílNov & JEFFERIES, [996fyros srrzerr:s: Maevínilia eslotiica Roztsr*ov & ifappesies,996

rÑsJMBER 1SF SFEciE5: 1[NCLitOiiOSPttiES: fiel oen,iilia cstooica * ROZHNOs & JH±E-Ríes, l996STrovrrc.ÑesPr riCAL OCCISRRíZNCU: Upper Ordovician (Caradoc)Geooíosrtuicxt, oisTRiuuxros: Europe (Estonia)RereRísr=cLs:RoztiNov & JEFFERiE5, 1 996

Minervaecystis UBAGHS & CÁSTISR, 1967TYI’E 5Fi&iLS: Dendrocystiíes Vidalí TíroRÁs., 1935NUMBER OF SFECiE5: 1INCLSJOLi) 5PEeiE5: Mincrvaeevslis vidali> (THORAL, l935)Srí=ÁriovsAi’Hícxr.sísIcIsRRLNcE: Lower Ordovician (Trema-doc)CieOGrtxruicAi, DiStRiisUTiON: Europe (France)RnswrerNcrs: TíroaÁr, 19355; UBAGHS & CASTER io CASres,1967

Myeinocystites SrRIMPÍw, [953Tíos swcríss: Mvei,soeysíites traías STRirviPLE, 1953NUMOFR oc surscssys: 21 set LOEn suecias: Mveinocvstííes erossmani KOLATA, STairs—rin & tEvORSON, 1977; M. ,íatus> Sír<iMPíe, ¡953SirsxricrsAi’h]ic:xe oCGURRENGE: Upper Ordovician (Caradoc)Geísc;R,xphireAe t)i5TRiBttTiON: Nortis America (tISA: Oklaho-ma)RFFERENSTES: STRíswrís. 1953 RorArA st al., 1977

Ratroclypeus WirHryRs, 1933Tyl’L SPiLCiL5: Rutrocíspeas jaeíori WrTrusns, [933NS:Mrsr:s 1SF 5PI5GiFS: 3Iscruoro sv’eca?s: Rutroclvpous janori> WiTunsRs, [933; R.vií¿oriae Grí.u & CAstas, í0~o: k nilhe,rsiGí:ú& CASTER,

[960Sísxsrorsxr’Hrcxs occUreseNee: Lower Devonian (Emsian)Gr:OouÁprircsí Di5TRSBUTiON: AustraliaREFERENCES: WsTrirRs, 1933: CASTos, [967

Ata annotated check-iist of genera and apecies of carpoids

Scalenocystites KOLATA, 1973TYPE SPECiE5: Scalenocystites striníplei K0ATA, 1973NUMBER OF 5PECiES~ 1INCLUDED SPECiES: Scalcnocysthes strimplei> KOLATA, 1973STRATíGRArHíCAL OcCURRENcE: Upper Ordovician (Caradee)GEOGRAPHíCAL Oi5TRiBtJTiON: Nortis America (USA: Minne-sola)RUFERENCES: KOLATA, [973a, 19735; ROLATA el al., [977

Syringocrinus ErLi.tNús. 1859Ter’r¿ sueCias: Syringocrinus paradoxicas BíLLíNOS, 1859NtsMrnsR or sPF.crtis: 1INCIUDISD sPíwrF.S: Syringocrinus paradoxicus> Brí.LrNGS,[859;5?. sinclairi PARSLEY & CASTraR, 1965STRsxTIGRÁroícAL oCI?URRENcE: Upper Ordovician (Caradoc)GF.OORAI’HíCAL Di5TRiBSsTrON: Nortis America (Canada)REresErsí:es: BiLLiNGs, 1859; PAR5LEY & CASTOS, 1965; CAs-tEle, [967

Comuta JAEKEL, 1901

Acuticarpus SLsINIRALL, SPRINKLE & GUENSBURG, 1997Tvua SuralEs: Acaticarpus delticas SUMRALL, SPRiNELE &GUEN5BI5RG, [997NUMBER 1SF SPBCiBS: 2INCLUDEO suecias: Aculicarpus delticus> SUMRALL, SPRiNKLE& GUEN5OURG, 1997; A.? repablicensis SUMRALL, SPRireK¿.E& GUFNSBIsRG, 997SrRÁTrc;RAPOíGAL OcCURRENGE: Upper Cambrian (Merionetis)GEoCRAPricAL DiSTRiBUTiON: Nortis America (USA)REFERENCES: SUMRALL etat, 1997

Amygdaiotlrrra IIRA<IIIS 1969¡YPE 5PFE1E5: Amygdalotheca gr~ffei UOAGH5, 1969NL’MaisR OF 5PECrLS: 1INCLUDrII) SPF.Crtt5: Aniygdalotheca grdjei’ UBAGOS, [969SrRAsicRAuurcAL OCIZURRi5NCE: Lower Ordovician (Arenig)GeooRAuíircAL DiSTRiBUTiON: Europe (France)RaaizRrzNerss: UBAGHS, 1969

Ampelacarpus LEFEBvRE &VízcAiNo, 1999Tyi’e srecirss: Ampeloeurprs lundeyranen.sís LEFEBvRE

&VízcANo, 1999NísMBers OF 5PBciE5: 1INcLusoeD SPECiES: Avnpelocarpas íandeyranensis* LEFEBX’RE& VizcAÍNo, 1999STRÁricRAuoicÁl. OcCURPENCE: Lower Ordovician (Arenig)GISOGRAPHíCAL Di5TRiOUTiON: Europe (France)Reresrasces: LEFEBvRE & VizcAíNo, 1999

Arauricyslis LEFÉBVRÉ & ViZ¿AiÑo, 1999TYPF suecies: Col/turnocystis prisnaeva THORAL, [935NUMBUR OF SPeeiE.5: 2Iscuoneo SuECiES: Araaricystis primnaeva 9< (THORAL, [935);A. occitana (UBAGIis, 1994)

38Coloquios cíe Paleoís:o/ogkí2002. 53: 33-68

Patricio Domínguez Alonso & el al. Ata annorated check-iist nl genera atad species ol carpoids

STRATIGRAPHICAL OCCURRENCE: Lower Ordovician (Arenig)GEoORArrrrcAL oIsTRiBUrIoN: Europe (France)REFERENCES: LEPEUsRE & VizcAíNo, 1999

Archaeocothurnus SUMRALL, SPRINKLE & OUENS-BURG, 1997

TYPE SPECiES: Co//taníocystis? b<flda UBAGH5 & RonisoN,1984NUMBER or’ sPEcrEs: 2INCLUDED SPECiES: Archaeocotharnas bjfidus> (UBAGHS &RoBísoN, ¡988); A. goshutensis SUMRALL, SPRiNiwE &GUENSEURO, 1997STRATíGRAuHíCAL OCIZURRENCE: Middle-Upper Cambrian (St.David s-Morionetis)OBeoRAur-ricÁL DiSTRiBUTiON: Nertis America (USA)REFERENCES: SUMRALL et al., [997

Babinocystis ROZHNOv, 1990TVPE 5PECiE5: Babinocystis dilabidus RozoNov, 1990NUMBER OF 5PECiE5: 1INCLUDED SuEnES: Babinocyseis dilabida> RoZHNos, 1990STRATíGRAPHrcAt. OCCURRENCE: Lower Ordovician (Arenig)GBoOo.áuHicAL DisTRIBUTiON: Europa (Russia)REPERFNcF.5: RozoNes, 1991)

Beryllia CRIPPS & DALEY, 1994TirE 5PECiES: Be,yllia miranda (?r¿íns & DALEY, 1994NUMBER OF 5PECiE5: 1INCLUDED suEciEs: Beryllia miranda> CRiPPs & DALEY, [994STrtÁrrc,RÁuriicAL OCCURRENCE: Middle Ordovician (Llandei-lo)GEoGRÁPHrcAL DiSTRiBUTiON: Europe (France)REFERENCES: CRíuus & DALEs. 994; RUTA, 1999b

Bohemiaecystis CASTER, 1967[partimn=T/toralicystis CA5TF.R & UBAGH5, 1967; LEFEBvRE& VizcAíNo, 1999][partiní=Scotiaecystis CHAUVEL, 1971; LEFEBsRE & VizcAÍ-NO, 1999]TYPE 5PECrES: Ro/temiaeczstis bouceki CASTOR, 1967NUMBER OF SPEC¡ES: 3INCLUDED 5i’ECrES: Bo/temiaecystis bouceki> CASTER, 1967;B. jefferiesi GIL CID, DoMiNrnísr¿z, SíLVÁN & EscRIBANo,1996; 8. guillor¿i (LÉr’EBvRE & VizcAÍNo, [999)NOTE: LBFBBVRE & VizcAíNo, [999 did nol consider thaIBolíemiaecystis CAsTRE, 1967 was a useful genus. They rofe-rred Bohemniaecystisjefferiesi Gis, CID, DoMíNGUEZ, SíLvÁN &EscRIBANo, 1996to Scoíiaecysíis CHALO/EL, 1971; Bohemia-eeyseis boaceki CASTER, 1967 lo Tlroralicystis CAsTER &UBAC,Hs, 1967 and placed Scoliaeryslis gr¿illoui LEFEBVRE &VizcAíNo, 1999 in Scotiacysxis CIIAUvBL, 1971. We considertisis synonyíny not justiflod. Botis genera sisare some charac-lers sucis as liso same number and distribution of marginal pía-les of liso head and a similar type of salve in liso branchialslits. However, among olber cbaracters, ihe sisape of tSe Sead(in outline and in dorsoventral convexity), liso types of oto-

gutnentaiy platos, liso length of Ihe anterior appendicos andliso ventral spikos of liso marginal trame are veiy difrerení inliso two genera.STRATíGRAPIUCAL OCtIURRENCE: Middle-Upper Ordovícran(Llanvim-Caradoc)GEOGRAPHICAL DiSTRIBUTiON: Europe (Bohemia, Spain, Fran-ce)REFERENCES: CASTRE in UBAGHS, 1967c; GiL Cío el aL, 1996;LErnusRE & VízexiNo, 1999

Ceratocystis JAEKEL, 1901TePE suFInrES: Ceratocysíis perneri jAEKEL, 1901NUMBER OF SPECiES: 2INCLUDED SuBerEs: Ceratocystis perneri> JAEKEL, [901; C.vizcainoi UBAOHS, 1987STRATIGRAí’HíCAL OCCURRENCE: Middle Cambrian (St.David’s)GEOORAPHICAL DisTRIBUTrON: Europe (Bohemia, France)REFERENCES: JAEKEL, 1901; UBAGHS, 1967b, 1967c, 1987;JEFEERiES, 1969, 1986

chauvelicystis UBAOHS, 1969TYPE SPECiES: Chauvelicysíis spinosa UBAGHS, 1969NUMBER OF SPECiES: 3INCLUDED SPECiES: Chauvelicysiis spinosa> UBAGOS, [969: C.ubaghsi (CHAP/EL, 1966); C. r’izcainoi DALEs, 1992STrtÁricRxur4rc.xí. OCCURRENCE: Lower Ordovician (Arenig)GEOGRAuHICAL or5TRiBUTroN: Northeos Africa (Morocco),Europe (Franco)REFERENCES: UnÁtsus, 1969; CHAP/EL, 1971; DALEY, 1992

Cothurnocystis BArRER, 1913TYPE SPECiES: Cot/tarnocystis elizae BATOER, 1913NUMBER or suEcros: 3INCLUDED suocios: Cot/tarnocystis courtessolei UBAGH5,

[969: C. elizae* BATHER, 1913; C. fellinensis UBAGHS,1969STRATIORAPHíCAL OCCURBENCE: Ordovician (Arenig-AsSgil[)GEOGRxí’HrcAL DiSTRiBUTiON: Europe (UK: Scotland, France)REFERENCES: BATHER, 1913, [926; JEr’rEítiízs, 1967, 1968,¡986

Domfrontia CRIPPS & DALEY, 1994TYPE SPECiES: Domfroniia pissotensis (CHAP/EL & NioN,1977)NUMBER or SuEciaS: 1INCLUDED 5PECiFS: Donífrontia pissoteasis> (CHAUvEL &NíON, 1977)STRATIGRAPH!CAL OCCURRENCE: Middle Ordovician (Llandei-lo)GEOGRAPHICAL DiSTRiBUTiON: Europe (France)REFERENCES: Creíns & DALEY, 1994

Drepanocarpos SMITH & JELL, 1999TYPE sPECIES: Drepanocarpos australis SMrru & JELL, [999NUMBER OF 5I’ECiE5~ 1

39 Coloquios de Paleontología2002, 53: 33-68

I’assie(ss I)essníssg’sív: Alssn.ss, & ci al. An arsnotascd chcck-list ol gencra aud spccics ol carpoids

INGitSDED SuElES: Dí-epaoocarpos auzcí culis> SMITH & JELL,

[999Srre.xrrcRAuhrrcÁs ocnt7rsRRFNriz: Upper Caníbrian (Merionelis)Giness:,roxpsriexL Di5TRSBISTiON’: AsísíraliaRSSIERIiN(r:1S: Svrrrs & liii.. [999

Flabellicystis Mxíeri MUs, 2002lyi’L Si’ LUí LS. Hsibellicvstis rushtoni MARTÍ M liS. 2002Ns’Mlt:rs OF 5t’F(iFS: 1¡NUI AS 1)111) 5 Pce ir:s~ Elote/líe: vstize sushtooi* MARTÍ Mlis.20(12Ssssxrsz;seÁ¡’r-sscxr, OCtURiCLNULi Lower Ordovician (Tronra-dsse)(3szsss;re,ss’r=ss:,xr,r:sssTs{rBs:rsssN: Esísope EsiglandíRvr r sise rs NI veis Mus, 21)02

Galliaecvstis UHAGOS. 1969lyui: suscrts: Gal/¡escss.slis li~,s Cre s/ U sAe; r rs, ¡969Nusíssr~se 01: SPFdrliS:

1 N(t.tiOEt) Sl’iSSii Fis: Geilliací v51is ligo ov’si 9< FisAs;ítS, 1969SrrexTrc;sÁrurcAs ssccs :RRFNcI: losvor Orslovicizo (Arooig)O iitSGRN sir íes u j >55*5511 ‘Irse=:Etí rope (FrancoRs i tsr se ¡ 5: Uss st ,os, ¡969: Cssir’s’s. [91=9

Hanusia CRIr’i’s, [989TvPH 5 PU JETS: Hsrrs ‘Niel prelepenxex CRss’FS. [959Nr:xsssí:se 5sf: s~l«[r:s 3lNe:[s:DITD SPI:crI:s: liasrsssics obtsssa Csrr’s’s. 1989: H. pci/e—penses CRisiS. 191=9:11. sorks’n.sis Cserr’s’s, 1989SrR,errcIexps1 ross. 5 seN r:sRiiNcI~: Midd lo tjrdssv ei:50 Sl jan -

si ri>—Llandei lo>GL:oe:re,sps titAs. 055TR555L’TSON: Etsropc lBoSersriaRs, Esí ‘si s Cu¡s’r’s. [989

Jullaecarpus Rí’r - [999Tvr’ts SF5055: .Juliaesísrpir.s oíilocrcrico Rsnr,s. [999Ns’uísíiíe oc SIFUiES: 1

1 Nere¡9145 .sPF.csr.s: iulieiccacpscs os iloes-oreo,s* RI:n:Á, 1999STw’]ss;usrístÁI. occrsI{RFNIF: Uppor Ordovician (Asisuilí)GresÑíÁs’wscv isssTRrlse:rloN: Nssí-tSern Africa (MoroecosRsí-nur’cr s Re r is. 1 999b

Lobocarpus tií}AGHS, 998Txs’r: sr’ssr rs /.ssbosesspír sv/zcssiooíL’tsÁesus, ¡998Nsevícs OF sl’I:crrzs: ¡IxeruoFIs sr’rxir:s: Lesbsssesryrus sizeoiooi> L’Fs,scniss. 1998Sise<tisis Al liCAs (Se SL RRUNCL: L

tpper Cambri sss (Mcrismnelis)Grx;seA 1’F 11(\t 5)1 Si R~t ION: Earope (France)Rssosrs’e 55 t4s Ns lIs ¡998

Lyricocarpus lii; y u’, ¡994Te~s SIte tUN Ls sse ocaspas soac/cssoloi títAGS 15. 1 994Nr:xíos:s OF 55t cts s 11 set í~DF:D5 sss se s 1 trIs osesrpus eoustessolei 9< tiissc;us, 1 994STRÁTIe;rexsírenAs OCCUteieLNUi¿: Lower Ordovician (Arenig)GI:oGss¡’esrcAs UrsrssBe’ísoN: Europe (Frasree)RI:rcrerzNctis: tJBÁGHS, 994

(esloqshsss sIc Psils’O,s teslss5’ísi

Milonicystis CHAUVEL, [986Tvun surcrrs: Milooicvstis kerfornei CHAUvEI., [986Ns>sroue OF SPLtIIES: 1lsci í DPi) SRFCIES: ¡í4ilooisvsiis kccJt cocí> CwsUvizv, 1986STle’sTIGR’pcsIc.sl OtIslISRREN(U: Middlo Ordovician (Llaodei-lo)GEoeauÁs’ctscxL DisrRIBuIrioN: Europe (France)Rrvrarsers: (‘russsvní - 1986: Caruus & Dsr.ss, [994

Nanocarpus UBAGHS [991TYPE SuElES: Naooezrr-pus dolavobíi tirs,scrrs, l 991Nr SM RtiR 5SF 5 upicsses:

1 NSTIl nEc) surcrrs: Neioosac~su.s eloleimbu 9< ti BAGHs, 1 991S-FICATs;ÑssuHrcÁs esvcutI<REMr::: Lowcr OrdovicianGiboeUsur [CALUIsTRIBUTsON: Europe (France)

Rvnrsvsus’s: thssous, 1991: Jt+prrzíirs, 1986

Nevadaecystis UHAGHS, ¡967Tyt>s síscies: Cothur,socvstis? aoiecisaoa UBAGH5, 1963Nsvrrsra 01: SuFs5r5: 11 NUlO 950 suszcies: ¡Verade¡esv.stis omnoricana 9< (UBAIIIiS,

¡963)Srssxrrssss.xr’Hrs:Ái, O5ULJIRFNUE: típpcr Cambrian (MerionetS)Geesosexr’HIUÁL, 055iRIBLT ION’: Norlis Arnerica (USA)REIoreENcr±s:UBAGH5, [963, 1967c: Ju’r±srt¡s,1969, 986;JSLFI’LRsLS el sil., ¡987

Phyllocystis lItORAL, 1935lvrc 5 [tUtES.Phvi/ores/it B/sssacsTHORÁL, ¡935Nssmnssu sic sr’ccrszs: 3IN(:s<:s)Er)sr’cercS: Ph ví/ocestís Ola veis í9< Toes a., 1 935; 1’.ecassiinarsinalss (TETORAL, 1935); P Y salaicíca Dtzssroi Osis.

[985STsCÁTIGSCAIsrieÁu oe:CURRENSTE: Lowei-Middlc Ordovician(lromadoc-L[andoilolGtzoeRAzrsss-AI. DiSTRIBI;TION: Europe (France): Asia (Siberia)Rrs±scNcris:TsíoRsí, 1935a; IIBAs¡Hs. 1963: Ltzwuvresz &VizcAíNo, ¡999: DusBÁrcuovÁ io RosesvÁ, Rosov & DUBATO-roe’, 191=5

Ponluulocarpus SUMRALL & SERIORLE. 1999Tvss SPECIES: Pssoiis:oloearpus cobí.soni SuSMRALL & Susís-ELE. 999Nusstrsse OF 55’LCiLS: ¡INS:LS]DED suE(:sEs: Po,síículocarpus resbíso,rí Ss:MRss 1. &Srrsíwmc. [999STRÁTIGRÁr’wcÁI occs’RRPNFR: Middlo CarobrianGísoea.Ns’ss rUAr. OrsiR 1 RUT Isis: Arnerica USA: FíaS)Rr-s RISC rs, SeMiextí. & SPRINFQLE, ¡999

Prochauvelicystis Dm UY, 1992Ivus strciss: Prssehauveíicysris sensespsnosa ¡JALEe, ¡992NI:MBBR oc SPFC?1H5: 1IN(:LI:rsszD srizcuzs: Proehaucclíeis’svís seínispinosa 9< Date,[992STRATIGRAPHICAI. 51t51CURRENCE: Lower Ordovician (Trornados)

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GEOGRAPHIcAL DIsTRIBUTION: Europa (UK: England)REFERENCieS: DxtEe, [992

Procothurnocystis WooDs & JEFEEmES, [992TYPE sí>btIIiss: Procotlrarnocysti.s owe,ísi WoODS & JEEFERIES,

1992NUsrímísR (SET SPEIiES: 1INCLUDED SPEcIES: Procoíbunrocystís oveensi> Weons & jiep-

FERIES, 1992STRATIGRAIHICAL OtICIJRRENCE: Lower Ordovician (Arenig)GEOGRAPHICAL DISTRIBUTION: Europe (UK: Wa[os)Rí±FERISNcEs:Wooos & JEFETERIES. [992

Progalliacystis CRIPPS, 1989TYPE SPIJCIES: Progallíasysti.s ubaghsi CRIuus, 989NUMBER 01’ SuElES: 1

INCLIsDED SFECiI5s: Progalliacvstí.s abag/tsi> Cssír’r’s, 1989STRATJGRAPHILAL OCCURRENCE: Lower Ordovician (Arenig)GEOGRAPHICAL DISTRInIJTIoN: Europe (France)

REFERENUES: UBAGHS, 1969: Cnsi’r’s, [989

Prokopicystis CRIPPS, 1989Tepie srieciES: Prokopicysíi.s mes~g/i CRíuPs, [989NUMBER OF 51’EtiIES: 1INcLUDEI> SPECIES: Prokopicvstis voergíi”’ Crapr’s, [989STRxrioRxrHICAi, OCCURRENUL: Middle Ordovician (Llandel-lo)GEOGRAFHICAL DiSrRiBISTIoN: Europe (Boisemia)REFERENCUS: CRIPPS, [989

Proscotiaecystis UBAGIIS, 1994TePE suiesnsies: Cot/turnsscvsli.s ,nelchiori UBAOHS, 1983NCMBER OF SPETUIES: 1[rsctUDr’.DSPECiETS: Proscotiaec:vslis orelchiori> (UBAOHS, 1983)STRAT5GRAPHICAL oCCURRENcI:: Lower Ordovician (ArenigíGeOGíuAr’íIIcAs, DiSTRSBUTION: Europe (France)REETERLN5iIJS: UFSAGES, 1994

Protocystites HICKS, 1872TePE sue<ii~s: Prolocyslitcs mencíensis HscKS, 1872NUNIBER oc sítú~s: 1INCLUDED siecries: Protocvsliles ,ne,seveosís* HícleS, 1872STRATIGRAPHICAL OIZcURRFNSti Middle Cambrian (St.David’s)GEoGRAPIIIUXL DISTRIBUTION: Europe (UK: Wales)REFERENCES: Hieies o HIcKs & iONES, 1872; JEFEERIES et al,,[987

Reticulocarpos JEFEERIES & PRoKOP, 1972TePE spieciies: Reliculocarpos lsanusi JErFERIES & PROKOP,1972NUMUER OF 5i’ECiE5: 1INCLUDED suEciEs: Revicuíocarpos hanasí> JEEFETRiEs & PRO-KOP, 1972STRATIGRAIHICAL tIisCcURRENCET: Middle Ordevician (Lían-virn-Llandeilo)

GEOGRAPHICAL DISTRmUrioN: Europe (Bohemia, Franco)REFEREN5IE5: JEFFERIIJS & PROROr’, [972; JEFFERIE5, 1986

Scotiaecystis CASTER & UBACHS, 1967Tyurs SPECiE5: Coí/turnoc-istis carta/a BATHER, l913NíssrBER OF SPECIES: l

INCLUDED SPECiES: Scoíiaecystis collapsa CRIPr’s, 1988; £curva/a

9< (BATHER, l913)STRATIGRAPHICAL OCCURRENCE: Upper Ordovician (Asisgilí)Geograpisical distribution: Europe (UK: Scotland, NortiseosIreland)

REFERENCES: BATHER, 1913; CASTER & UBAGOS in UBAOHS,[969; JEPFEFSIES, 1967, [968, [986; CRíuPS, 1988

Thoralicystis CHAUX’EL, 1971TePE 5PF.CIFS: Scotiaecisss/is grrffei UBAGFS5, 1969NUMBER OF SuECiaS: 4INCLUDED SuElES: Thoralicystis grrffeí* (UBAGHS, 1969); 7?ubaghsi LEETEBVRE &VIzcsiNo, 1999; 7? zagoraensis CHAU-VFL. [971STRATIGRAI’HICAL OcCURRF.NCE: Lower Ordovician (Arenig)GEOGRAPHICAL DISTRIBUTION: Europe (France), NortiseroAfrica (Morocco)REFERENCES: UBAGHS, 1969; CHAUvEI., 1971

Trigonocarpus UBAGES, 1994TePE suuinS: Trígonocarpus singularis UBAGHS, 1994NUMBER OF SPECIE5~ 1INCLUDED suEciEs: Trigonocarpus singularís* Uaxous, 1994STRATIGRAPOICAL osXURRENCE: Lower Ordovician (Arenig)GEOGRAPIII5?AL DISTRIBUTION: Europe (France)REETERENCES: UBAGH5, [994

Mitrata JAEKEL, 1901

Adoketocarpus RUTA & JELL, 1999Tear’ sPEcíE.s: Adoketocarpus oc/reron/icus RUTA & JELL, 1999NUMBER or’ sp~cries: 2INCLUDED SPECIES: Adoketocarpus acheronticas> RUTA &JELL, 1999; A. janeae RUTA & JELL. 1999STRATIGRAPHICAL OCcURRENeSE: Upper Silurian (Piido[i),Lower Dovonian (Lociskovian)

GEOGRAPHICAL DiSTRIBUTION: AustraliaREFERENCES: RUTA & JELL, 1999a

Állanicytidium CASTER & GILL, 1967TePE Suades: Alla,íiczvtidíamflemingi CASTER & GILL, 1967NusMutsrs 5SF SPECIES: 1INCLUDED sPF.CIFS: Allanicvíidiumflemiogit CASTER & GILL,

1967STRATIGRAPHICAL OCCURRENCE: Lower Devonian (Emsian)GETOGRAPHICAL DISTRIBUTION: New Zea[and (South Island)

REFERENCES: CASTER jo UBAGHS, 1967c

4natifopsis BARRANDE, 1872TePE suEcíEs: Anatífopsis bohevníca BARRANDE, 1872NUMBER OF suieCiEs: 18


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Patricio Domínguez A/miso & el al. An annorared checle-lisí of gesrera and species of caspoids

[NCi.UDFDSFECIES: Ana/sfopsis acato BARRANDE, [872; A.ancora DOsrÍNGos-iz & GLTIÉRREZ-MARCO. 1 990; A. arProri-tana (CHAUVEL, ¡941); A. ba/claíclsíensí.s RitEn, 1940; A.borrondci (Círsinvar.. 1941); A. bohemíca> BARNANDL /872;A. cocaban KouxvxsHi, 1960; A. escandei escandei THORAL,

¡935; A. escande/Y t/rorali (CHAP/EL, [966);A. gaicheííensis(CuxisvEi., 1941): Aootí¡ópsis longa BARRANDE, [872; A.minuto (CI]xísvcL, [941): A. popillata (BASSLER, 943); A.panctata (BAsSIFR, >950): A. spínosa IJBAGHS, 1979; A. Ira-pezisfor,nis Triouxr , 1 935; A. travea.so/ensis (CrsAhsvcr.,

[941);A. lrsorcoto Kowxs’xsrsí, 1960SnArIGRArIiíc::sí. IXSCURRENCE: Middle-tipper Ordovician(L¡anvinm-AsSgill)Guoc;axuHIcxt. DIsTRIBsJTION: Europe (Boisemia, France,Spain, Sweden, 11K), NorIS Amorica (tISA), Asia (Korea),Nertisorn Africa (Morecco)REFERENCES: BARRANDF-., 1872; JrsI’rwwíEs, [986; DoMir=oL+z& GUTIÉRREZ-MARCO, ¡990; PARSLIIY & PRoKou, 1999

[AnatiferocystisCHAI5vETL, l94¡ 1=Anaofrpsís BARRANDF, 1872 -‘-

REFERUNCES: CHAUVEL, ¡941; UBrserís, 1979: DOMÍNGUEz &GL’TIÉRRFY-MARCo, 1991)

[KirkocvstísBA5SLLR, 1951)]=Ana/ífrpsis BARRANDE, [872TYFE SuadEs: Enoploaro? papilla/a BASsI,aR 1943REpERENCES: BÁsSs,tiu, 1950

Ánomalocystites HALL, 1859[=BasslesocystisCASTre. [952(junior synonym)]TYPE SPEcIES Aoo,nalocz’stites dispar/lis HALL, [859NLMIsER OF SPiSCiIi5: 2INCIsIDIzD SFECIES: Anovoalocvstiíes corros/as HAll., 1859; A.dispar/lis> HALL, [859SNí=xTioÑxrriíext.OCCURRENCE: Lower Devonían

GETXíRAPHiCAi, DISTRIBItTION: Nortis America (USA)REFETRENCES (‘ASTER, [952, Kos NrA & Jor trE. [982

Aspidocarpus UBAGUS, 1979TYFE SPEIiiES: Aspidocarpas ho/re Inicas UOAGOS, l979NUMBER OF SutiCIr¡S: 2tmjr’ ITrST¡rscI,r:c-TLC A ,,,J,,-,,.,,, kt.,,.,i,,,9< Itn,.-’ns, iO’7O

A. riadaneuísis (Crrrss¡vrr., 1937)STRATIGRAPIIICAL OCCURRFNCI:: Upper Ordovician (Caradoc)Gaocuxuííwzxi, DISTRIRI:r ION: Europe (Bohemia)

RErFRETNCFS, UssÁt3tIs, 1979

Ateleocystites BILLINGS, 1858zAtelcocvsíis LINDSTROM, 1888 (noin. van.)]

[<X/elecystísBesTueR. [889(nom. van.)][=Atelocysti.s¡IAECREL, [896(oom. van.)]

Tvr’c ss±cías:A¡eleocvs¡iíes /rnoJevi Bii.tsscs, [838Nusvmarc OF SuECIr¡S: 2INCI,IsDETD SPECIES: Atcleocysti/cs haxíesí> BILLINGs, 1858; A.guttenbcrgensis KOLATA & Joitie, 1982

S1rsxriGí*xuHicu OCCURRENCE: Middle-Upper Ordovician(Llandeil[o-Caradoc)GEOGRAPHICAL DISTRiBUTiOiÑ , Nortis America (USA)REFERENCES: HAECKEL, 1896; CASTER in UBAGHS, 1967c;KOLATA & JOLLiE, 1982

Australocysñs CASTER, 1954lira SPECIES: Ausíralocystis laogeí CASTER, 1954NUMBER OF SuEcIES: 1INCLUDED 55’iSCiES: Austraíocysiis langei> CASTER, [954STRxsIGRAPIIWÁI. OCCURRENCE: Lower Devonian (Emsian)GFoGIssptssCÁr, DISTRiBUTiON: AustraliaRFFERF\CES (ASTER, 1954a

Balanocystites BARRANDE, 1887L=Sagitlac~yatis PARSLEY, PROROr & DERSrLER, 2000]TYPE sFE(isis: Balonocysíites lagenala BARRANDE, 1887NUMBER 01 SiECIES: 2INcIIsoF.D SPECIES: Balanocyslííes lagenala> BARRANDE,

887: B. primas (BARRANDE, [872)S,íaxrioísxuHiexL otiCL’RRENCE: Middle Ordovician (Lían-viro)

GeocRAuuícAL Di5TRiBItTION: Europe (Beisemia)RUrERENCES: BARRANDE. [887: Jocs’crsírs, [986; PARSLE’I &PRoKou, 1999

Barrandeocarpus UBAGOS, 1979TYFET SPECIEN: Barrandeocarpas jaekoli UBAGHS, l979NUMBER OF SPECIES: 2[NCLIsDI:DSirCIES: Barrandeocarpus jaekeli* UBAOHS [979;B. oon’egisa.s CRA5KE & JISFrERIES 1989STRATiGRxrHiCáL OCCURRENCE: Upper Ordevician (Caradee-Ashgill)GEoGRxuIIIc¡sL DISTRIBUTION: Europe (Bohemia)REFERENCES: tJnxc,u,s, l979; CRASRE & JEPIERiES, [989;RUTA, 1997d

rBasslerosvs/is CASTre, 19521=Anomalor-s’s//tes; HALL 1859 —~

REFERENtES: CASTER, 1952; KOLATA & JOLLIE, [982

Bokkeveldia RUTA & TFIERON, 1997LXITlEflá5JN,1fl5

NFsMBER OF SPECIES: 1INCIUDEI) SPLCiES: Boklcevcldia oos/haízení> RísTss & THE-

RON, [997SrssÁTiGRAPIiiCxL OCCLtRRENCE: Lower Devonian (Emsian)GrOoRAs’r-sií:Ai DISTRIBurTION: Soutisom Africa (Republie ofSoutS Africa)REFERENCES: RUTA & THERON, [997

Chau velia Cr<tpps, 19901yi’E Sr’ETCIES: Choavelia discoidalis CRIPPS, [990NUMBER OP SiEciEs: 1lNCiUI)Eo Suecies: G/tauvclio discoidalis> CRIPPS, 1990SrrassoitxnsrcxL. IXCs’RRFNCE: Middle Ordovician (Llandeilo)

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GEOGRAPHíCAL DISTRIBUTION: Nortiseos Africa (Morocco)

REFERENCES: Cruí’ps, 1990; LEFEvURE, 2000b

Chinianocarpos UBAGES, 1961Tvpn SPECIES: C/tiniatrocarpos lsíroroli iJnxcus, 1961NUMBER OF SuEciEs: 1INCLUDED SPECIES: Chinianocarpos t/íorali> UHAGHS, 1961STRATICRAPEICAL OCCURRENCE: Lewer Ordovician (Trema-doc)GEOORAPH¡CAL DrsrRiBUTioN: Europe (France)REFERENCES: UBAGHS, 1961, 1967c, 1969; JEFrERIES, 1986

Dalejocysús PROKOP, 1963TvPE SPECIES: Dalejocystis casteri PaoKou, 1963NuMMER OP SPECi~S: 1INCLUDED SPEiES: Dalejocystis coste rl PROKOP, 1963STRATiGRAPHIcxL OCCURRENCE: Middle Devonian (Eifelian)GEOGR.APHICAL DiSTRiBUTiON: Europe (Bohemia)REFERENCES: PRoKoP, 1963

Diwnphñliocystis KOLATA & GUENSBURG, 1979TYPE SPECiES: Diamphidiocyslis drepanon KOLATA & GUENS-

EURO, [979NIR,IBER OF sPEcrES: 1INCLUDED SuEciaS: Diamphidiosyslis drepanon* KOLATA &GUENSEURO, l979STRATíGRAPHíCAL OCcURRENCE: Upper Ordovician (Ashgill)GEOGRAPHICAL DiSTRIBOrION: Nortis America (USA, Franco)REFERENCES: KOLATA & GUENSBURO, [979; CHAUN’EL, 1982;LEFEvBRE, 2000a

Enoplonra WETHERBY, 1879Tira SPECiES: Anomalocyslites (Ateleocystiees?) balanoidesMEEK, 1872NUMBER OP sPEcrES: 5?INcLUDED SI’ECiES: Enoploara balanoides WETHERBY, [879[MEEK,EB. 18721; Enoploura craslacea (Hxrcia, 1896);Enoploara meeki CASTER, 1952; Enoploura popei CASTER,

1952; Enoploura wet/terbyi CASTER, 1952Nora: TSe genus Enoploara needs revision. Mosí of tSe des-cribed species musí be restricted te ihe types because lSeylack diagnostic characters.Srí<ATiGRAPKIrxL oCCIJRRENcE: Upper Ordovician (Caradoc)GEOGRAPHICAL DiSTRiBUTiON: Nortis Aniserica (USA)RaraínsNcaS: WETHERBY, ¡879; HAECREL, 1896; CASTER,1952; PARSLEY, 1991

Eumitrocystelia BEISSWENGER, 1994TYPE SPECiES: Eu,nio-ocystello savilíl BEiSSwENOER, [994NUMBER OF SPEciES~ [INCLUDED SPEiES: Ewnitrocys/ella savilli* BEISSWENOER,

1994STsríATioísAPHIcAL OCCURRENCE: Middle Ordovician (Llandei-lo)GEOORAí’HICAL DiSTRIBUTION: Nortisera Africa (Morocco)REFERENCES: BEISswENGER, 1994

Guichenocarpos CHAUVEL, 1981TirE SPECiES: Guiehenocarpos nioni CHAtO/EL, 1981NUMBER OF SPECIES: 1INCLUISED SPEciEs: Gule/tenocarpos nioni* CHAUVEL, 1981STRATIGRAPInCAL (XncURRENCE: Middle Ordovician (LI andeilo)GEoORAPHICsa. DISTRiBUTiON: Europe (France)

REFERENCES: CHAUVEL, [982

Jaekelocarpus KOLATA, FREST & MAPaS, 1991TYPE sPECiES: Jaekelocarpas okla/romensis Kou.xrx, FREST &MAPES, 1991NUMBER OF SPEciES~ 1INCLUDEIS SuElES: Jaekelocarpus okla/tomensis* KDLATA,FREsT & MAPaS, 1991STLATIGRAPBICAL OCCURRENCE: Upper Carbonifereus (Mes-

covian)GEOGRAPHíCAL DISTRIBUTION: North America (USA: Oklaho-ma)REFERENCES: KOLATA el aL, 1991; DOMíNGUEZ el al,, 2002

Kierocystis PARSLEY, 1991TYPE sPECíEs: Kieroeystis inserta PARSLEY, 1991NUMBER OF 5PECiES~ 1INCLUDED SuECIaS: Kierocystis inserta* PARSLEY, 1991STRATiGRAPHICAL OCCURRENCE: Upper Ordovician (Caradoc)GEOGRAPHICAL Di5TRIBISTiON: North America (USA: Ken-tucky)REFERENCieS: PARSLEY, 1991; RUTA, 1999a

Kopflcystis PARSLEY, 1991TypE SPECIES: Kopfieystis kir4fieldi PÁIIsLEY,1991NuMBER OF si’~Cia.s: 1INCLUDED SuEllEs: Kopficystis kirkfieldi* PARSLEV, 1991SNATIGRAPFECxí. OCCURRENCE: Upper Ordovician (Caradoc)GE(XSRAPHICAL DíSTRIBUTION: Nortia Aznerica (Canada: Ontario)

REFERENCES: PARSLEY, 1991; RUTA, 1999a

Liigynocystis JAEKEL, 1918TYPE SuECIaS: Anomalocystites pyramidalis BARPANDE, 1887NUMBER OF 5PEc1ES~ 1INCLUDED SuECIaS: Lagynocystis pyramidalis

t (BARRANDE,l887)STRATiGRAPHICAL oCCURRENcE: Middle Ordovician (tlanvirn)GEOGRAPHICAL DISTRIBUTiON: Europe (Bohemia, France,Spain)REFERENCES: BARKANDE, 1887 JAEKEL, 1918; CHAUVEL,[941; CHAUVEL & NiON, 1977 JEFrERIES, 1973; UBAGHS,1991; PARSLEY, 2000

Milonicystis CHAUVEL, 1986(Somotinises descriised as mitrala, see Milonicyslis in lIso Cor-nula genera lisí “)

Mitrocystella JAEKEL, 1901Tyí’c SuEciaS: Anonralocystites incipiens BARRANDE, 1887NUMBER er’ suEcias: 2 (and a subspecies)

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Patricio Doosíoguez Alonso & es al. An annosased check-iiss of genera and species of carpoids

INCi,511)FO SUEllES: Mivrocssíclla barrandel JAEKEL, 19<)!; 41.inc/picor> (BÁRRxNros, [887);41. incip/ens osiloni (BARRAN-

DC, ¡887; CírAu:vEr., ¡941)STRATIGRAPH5CAL OCCiSRRFNCF.: Middle Ordosician (Lían-vira-Llandoilo)GETOt;RAulrrUAs. DISTRiaUrioN: Europe (BoSemia, France,Spain>RF.Fr:RISNI:rs: BANRANDE. [887; JArRo,, [91)1; UEesc;us,1967c: Jszrrsieies, [967, [968. [986: PAR5I.EY. [994

Mitrocystites BARRANDF~, ¡887Tvssz Si’tiUiLS: Mi/rotrw/res mi/ra BARRANOr. 1887NI’Mnízsz 0v Sus7Cies: 1 (and one doubtfolly altributed te ISisscíeous)1 NCtiT>EIs Sl’ECiETS: Mitrocvslites mitro> Ii ARR ANDE, 1887:«,14 /tsosí’stiie.s» stvlo/detss DEHM, ¡934

Srvsxní;RApizicxi, OCCiIRRiiNI?F: Middle Ordovician (Lían-“ini)GEoGIIxiHSUÁL DISTRIBITION: Europe (BoSeíoia)

Reí:rRíz:Ncrs: BARRANDE, [887: iNFREL, [918; UBAGIIS,1967c; ]ETI:FETRIvSS, ¡967, [968, 1986; PNRSI.ssx, [994

Mirroívsli.s BÁíRáíiiE, ¡887; CrssssvFs, 1941. incorrecí spe-lIingj

=¡Viitroessritcs BARRANDE, [887RPFERRNUF:S: BARRANDE, ¡887: ChrAUva. 1941

Mongolocarpos RozHNov, 199<)TYPET SPECIb:s. Moogolocaipos minzhiíri RozFrleov, 1990Niza nr-a or SFEUiLS:

INULL’DLD SFEUIFS: Mongolocas-pos oíioz/rmni> Rozusov, 1990STÑATIGRÁFII cAí 5 sCsiIJRRENCF: Si jurian (Ludlow)(ieocssÁuríícÁí. 0sSrRiuuTioN: Asia (Mongolia)Rtzr’ekLrsccs: Rtszrassov, 1990

Notocarpos I’uríuíp, 1981Tvus: SPFictcS: A’orccaspos gas-raId Ponto. [981Nisswra ot~ SItúES: 2INCI,U 1)5715 St’LCSES: Noeoceu-pcs garratti> Pr isíru, ¡981; A’. ~>ra—oor (O tAiMAN, [91)4>SraÁTrcmAurrícAr. CsCCLJi<RISNUET: Upper Silurian (Ludlow>GFOGRÁi’FiiCAI. DiSTRsniJTION: Australia (Victoria)

RczrrzssL’swEs: Purrí>’, [981

Occultocystis 1-lAUDE, 1995Tvua suvzcr í::S: Occu/toctsr/a kocoe,r 1 HasiriE. [995NF:MBER OF SFETCIt:S: 1INci .1 ?DFTI SuEZ res: Occullosvsr/s /coctseni 9< H ‘sí np. ¡995STRxTIoícArurrÁI. ttt?t?l;RRENCE: Lower Dovonian (Emsian)

GrocreutireAL iIsIsTRmIisro\: SoutS Amorica (Argentina: Sao

Juan>Rrr FIONCES HAtiusis, ¡995

Ovocarpus UBAGHS, [994Tvie sueesas: Or’ocaspas monccreti UBAUSOS, [994Ni¡vIat¡R (511 SEFUIES: 2

Coloquios dv Palessssssslogíer21)02, 53: 33-68

INCL5JDED SPETCiETS: Ovocarpas? circuías-ls UBAcrís, ¡994; 0.,nons:ereíi> UBAGOS, 1994STRAOGRAPIOCAL OCCURRENCE: Lower Ordovician (Arenig)

GLoGRAPIIICAI. DISTRIBUTiON: Europe (Franco)

RIFERENCES: UBAGHS. [994

Paranacystis CASTER, 1954Tyi’r-~ sueelEs: Parouracvsris pctrii CASTOS, 1954NL:MBETR OF SFECIFS: 2lNUIi~DFt) SUrCiES: Paranacyslis petrii> CASrER, 1954; E.s,moneae RIsTA, 1997STRPrsc;RAuuIcs.s. 0t?CURRENCE: Lower Devonian

GFOGII.áíticAL DrsTRinUTII)N: SoutS America (Brazil: Paraná)R5?FFRPNInETS: CASrER, ¡954b: UnAUrIS, 1967c

Peltocystis THORAL. 1935TirE SImUlES: Pcltocvst/s cornuto IÁioRÁL, 935Nt;xioszu OF SUECiES: 1INCJ1;OUD SETEUrES: Peiiocvstis coníala

9< THORAL, 1935SrRArsssÑAr’rrICAv, OCcí¡RRENcET: Lower Ordovician (Troma-doc)GBOGRAPHíCAL DIsTRrBI:TIoN: Europe (France)

RFYLRLNCES: TíloRAr. ¡ 935b: UBAc;íís, [969:JlzrrssRsrzs, [986

Placooystella RENNIE. 1936TYPET SuEllES: Placocvsí/s africana RaED, 1925NUMBER OF SueCias: 1INCLIsOFO SusICIES: Placoc-vsrcllo africana> (REED, [925)SraxíiGRxt’oIt:Ai, OCCURRENCE: Lower Devonian (Ernsiaim)GEOGRÁPFiICÁL DISsRiBs)TJON: SoutSern Africa (Ropublic ofSoutb Africa)REFERENCES RENNIE, ¡936

IP/otee ‘,stss O arr sri 1 SOGí 1 noris van

par/ros Pías ocx sri/es oc Korssst:ie, 1869partan Pías oc ;wcila RENNiE, 1936

Placocysfltes DE KoNrNcí<, 1869[=PiacocystisHAECREL, 1896 (floro, van.)]TYPE SUElleS: Ploctwvsv/tes forbesianus DE KoNswa, 1869NUMBER >45 SPECIF.S: 1INULUDED SiETUsETs: Placoívscites ftsrbesiosías> DF. KoNINCK,

[869STrcxTicsRApvUíCÁI, OCCURRENCET: Silurian (Wenlock)GEOGRAI’H5UAL DrSTRIiSIsTrON: Europe (UK: Eng¡and)RIirr:Rt:Ns rS: JF~ETFERíES & LEwis, 1978: RUTA, ¡998

[PromivrocvsritesLETFETBN’RE, 2000]—,1eIiOocyste/ia JArRES., ¡901REFERINCí s, JNETREL. 1918: LEFEvBRí:, 2000b

Protocyñdiun, RUTA & JELL, 1999TirE SuEllES: Protocvtidis¿vo elliorrac Ru:rx & JELL, [999Ns¡MvsL:R 05: 55r-<riiS: 1lieci.isnizn sí’seics: Protocvtidiaío cIlio/toe> RUTA & JEIL, 1999STRATICRAFíIIUAL OUCI?RRENCE: Upper Ordovician (Asisgilí)

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Patricio Domínguez Alonso & el al, Aa annotated cbeck-iisr of genera auíd species of carpoids

GEOGRAPHICAL DISTRIBUTION: Australia

REFERENCES: RUTA & JELL, 1999b

Pseudovictoriacystis RUTA & JELL, 1999TYPE 51’ECIES: Pseadovictoriacvstis problematica RLrrA &JELL, [999NUMBER OF suECIas: 1INCLUDED SPECIES: Pseadovictoriacvslis problematica* RUTA

&JELL, 1999SrvsricRAuuicxL OCVURPENCE: lxwer Devonian (Ixchkeviarí)GEOGRAPHICAL DISTRIBUTION: Australia (Victoria)

REFERENCES: RUTA & JELL, 1999d

Rhenocystis DEHM, 1932TYPE SPECIES: Rhenocystis latipedanculala DEaIM, 1932NIJMBER OP SPECIES~ 1INCLUDED SPECIES: Rlrenocystis iatipeduncr¿lata* DEKM, 1932STRATIGRAI’HICAL OCCURPENCE: Lewer Devonian (Emsian)

GEoGRxPHIcAL DiSTRiBIYIioN: Europe (Germany)REFERENCIeS: DEHM, [932; RuITA & BARrEES, 1998; SUTcLIr’-FE et al., 2000

[Sagittacystis PARSLEY, PROKOP & DERSTLER, 20001=Balanocystites BARRANDE, 1887 4

REFERENCES: PARSLEY, PROKOP, & DERSTLER, 2000

Spern¡acystis UBAGHS, 1967[=AnomocystisJAEKEL, 1918]T’íPc SPECiES: Anomalocys/i ¡es ens~fer BARRANDE, 1887NUMBER OF SuElES: 1INcLUDED SPECES: Sperníacystis ensífer* (BARRAr-iDE, 1887)STRsTiGRAr’urCsL OCCURRENCE: Middle OrdovicianGEOGRAPUICAL DISTRIBUTION: Europe (Bohemia)REFERENCES: BARRANDE, 1887; UBAGLIS, 1967c

Tasnuanicytidium CASTER, 1983TirE SPECIES: Tasmanicytidiam barre/ti CAsTER, [983NUMBER OF SuPCIES: 1INCLUDED SPECIES: Tasmanicytidiam barre/ti> CASTER, 1983STRATIORAPHICAL OCCURRENCE: Lower Silurian (Llandovery)

GEOGRAPUiCAL DISTRIBUTiON: Australia (Tasmania)

REFERENCES: CASTER, 1983

Victoriacystis GILL & CASTER, 1960Tirc suEcíEs: Victoriacystis willcinsi GILL & CASTER, 1960

NUMBER OP SPECiES~ 2INCLUDED SPECiES: Viclorracystzs /roltnensoran, RUTA & JELL,

1999; 1? wilkins? GILL & CASTER, 1960STRATIC,RAuHíCAL OCCURRENCE: Upper Silurian (Ludlow)-

Lower Devonian (Lochkovian)GEOGRAPHICAL DiSTRIBUTION: Australia (Victoria)

REFERENCES: GILL & CASTER, 1960

Vizcainocarpus Rris’rA, 1997TirE SPECiES: Vizeainocarpas den/iger RUTA, 1997NUMBER OF SuElES: 2

INCLUDED SPECIES: Vizcainocarpus dentiger> RUTA, 1997; 1Krutai LEFEBvRE, 2000STRATIGRAPHICAL OCCURRENCE: Lower Ordovician (Trema-doc-Arenig)GEoGRAr’HicmL DISTRiBLrI1oN: Europe (France, England)REFERENCES: RUTA, 19975; LEFESBRE 20005

Wilinzanocystis KOLATA & JOLLIE, 1982TYPE SPECIES: Willunanocystis denticulatas KOLATX & JOLUIE,

1982NUMBER OP SPECiES~ 1INCLUDED SuEcias: Willtna,rocystis denliculala* KOLATA &JOLLiE, 1982STRAooRxumcxt. OCCURRENCE: lJpper Ordovician (Caradec)GEOORAPHICAL DISTRIBUTION: Nortis America (USA: Wiscon-sin)REFERENCES: KOLATA & JoLLIE, 1982

Yachalicystis HAIJDE, 1995Teue SPECIES: Yac/talicys/is triangularis HAUDE, 1995NUMBER OF sunC~S: 1INCLUDED SPECIES: Yaclaalicystis triangularis* HAUDE, 1995STRATIGP.APHíCAL OCCURPENcE: Lower Devonian (Emsian)

GEOC,RAPH1CAL DISTRIBErHON: South America (Argentina: SanJuan)REFERENCES: HAUDE, 1995; LEFEvERE, 2000a

II. íNDEX OF SPECIES

Cincta JAEKEL, 1918

Asturicystisjackeli SDZUY, 1993STRATIGRAPHICAL OCCURRENCE: Middle Cambrian (St.Daviris)GEOIIiRxrKicxL oiSTRiBUTiON: Europe (Spain)REI’ERENCES: SDZUY, 1993

Asturicystis havliceki FArRA & KORDULE, 2001STRATIORAPIIICAL OCCURRENCE: Middle Cambrian (St.David’s)GEOCRAI’KICAL DISTIuBUTiON: Europe (Bohemia)

REFERENCES: FATRA & KORDULE, 2001

[Cephalocinctuslabiatus TERMIER & TERMIER 1973]=Efliptocinctas barrandei (MUNIER-CHALMAS & BEROERON,

1889REFERENCES: TERMIER & TERMIER, 1973; FRIEDRICH, 1993

[Cyclocinctussabeircalaris TErwraíí & TERMIER [973]=Ellip/ocinctas barrandei (MUNIER-CHALMAS & BERGERON,1889)REFERENCES: TERMIER & TERMíER, 1973; FRiuosICH, [993

LCyclocinctas nraroecanus TERMIER & TEIO.4íER, 1973]=Sueocystis? maroccana (TERssiER & TERMIER, 1973) —>

REFERENCES: TERMIER & TERMIER, 1973; FRIEDRICH, 1993

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Patricio [)ombigssezAlonso & el al. An anírorared clsecle-lisí of gesrera and species of carpoids

Davidocinetus pe,nbrokensis FRtEDRICH, 1993STRATrGRxpírscAí. c>ccLSRISETNcE: Middle Cambrian (St.David‘s)GFOGRAPHICAI, DISTRIBUTION: Europe (UK: Wa¡es)RFFERFNCES FRrrrorcH, ¡993

[Oec:acvstis/r/spanic:a GísLÉs, 1927: dubious species]= Gs¿ro e vs//.s /rispon/ca (GalÉs. [927);dubieus spocies—‘ Gvrocvs/i.s piascsso JAEKEL. [918 -->

STRÁTsGRÁPIIIcÁr OCCLiRRENCET: Midd[e Cambrian (St.David s)GrsOcaÁuírIcÁL DiSTRIISUTIoN: Europe (Spain)

REFERENCES: JxEKETI., 1918; GísrÉN, 1927: FarEaRícíl, 1993;GIL Cío & DosrNcustzz, 1994

[Dec:acysí/spraeeiiiptic:us TE5zMIER & TERMíER, l 973]= Gyrocx’stis plares.sa JAEREL. [918REFERENCI;S: FI>IEoIIIcH, [993

Elliptocinetus barnandei (MUNIER-CHALMAS &Bí¿RGr-zrzOÑ, 1889); TeaMuza & TERMIER, 1973

Socxrrcacxs’Híc:xs. tti?t:tJRRFNCE: Middle Cambriarí (St.David s)GecxsrcxI’HrexL DisrRiBlsTit)N: Europe (France)

RSFLRLNCEs: MLNíER-CHALMA5 & BeRc;eRors iii BEROFRON

[889; CaRMeN & IERMIER, [973;FRíEDRICH, 1993

Elliplocinetus vizcainol EniEDRicH. 1995STRArIGRAIHICAL OCcISRRENCE: Midd[e Cambrian (St.Davids)GEoCRxumcxt. DiSTRiBI;TION: NortSern Africa (Morocco)

REFETRENCE.S: FRIETiDrsICr-s, 1 995

Gyrocystis badulesiensis FízíísDRieH, 1993SmxrscuAuríIcÁI OCCURItENCE: Midd[o Cambrian (St.David siGEOGRAPHICAL DISTRÍBUTION: Europe (Spain: Aragón)

Reecaeseas: FareDRice, 1993; Go, Cro & DOMÍNGUW, 1994

Gyrocystis cruzar GIL & DOMíNGIsEz, 1995Nora: seo Noto for Gvrocsxstis platcssa JÁstREI., [918 >

SraxrIGRxI’HICÁL OCCURRENCE: Middle Cambrian (St.David s)Gnix;RAr’FsiCAL DISTRiHUTION: Europe (Spain: Asturias)

Rr.rszRs:rsceS Gis, CID & DOMÍNOUEZ, 1 994

Gyrocystis erecta FRIEDRICFr, 1993Snxris;íeAI’r-sicAu, OccIsRRENCE: Midd¡e CamSrian (St.David s)GEOGRAPHICAL DISTRmU1iON: Europe (Spain: Aragón)

REFszRETNCES: Farntsaicrt, [993: GIL CID & DOMÍNGUEZ, [994

Gyrocyslis hispanica (GISLÉN, [927), dubieus spe-cíes, new combinatien heroin

[=Deeac}’.stis/rispanicus GísrÉN, 1927][=2Cicroc}’stis plotesso JAEREI.. 19181

NOTE: Formally Decacvslis hispanica is restricted lo tSeisolotype. FaíEDarcu 1993 considered it as a probable seniorsynorxym of Gyrocysíis platessa JAEREL, 1918. However, alt-Songis tSe altribulion lo Gyrocystis is obvious, liso atíributionlo Gvrocv.stis pla/essa JAENEI., l918 is deubíful. (Seo note en(iyroty.sl/.s plates.sa JÁFREI,, 1918)STRAT5GRAPHICAI. OCCISRRENu7E: Middle Cambrian (St.David’s)GacscRxrnuiC=wDiSTRIBusTIC)N: Europe (Spain: Castilla-Loán)REPETRENCES: JAEKEI., 1918; GIsLÉN, 1927; FRIEDRIcH, 1993;Gil. Cii) & DoMÍNcsusEz, [994

Gyrocyslisplatessa JAEKEL, 1918[?=Decacystishispan/ca GIsLÉN 1927; dubious species]

Decacvs/is praeellip/icas TERMIER & TERMIER, [973]NOTE: In cur opinion Gyroczstis platersa JAEREL, l91 8, asrevised by FRIEDRIcH, 1993, is a complex of fossils, [listen-cally, any Ciocta witS ten marginal platos was described oit-Seras Gyrocysíis platessa JAEREL, [918oras Decacysl/s his-¡mao/ca GísÉÉrs 1927 ( >). A revision of liso materialdetemiined by FRiEDRrCH, 1993 allowed swo of tSe preseníautisors lo propose Gzrocystis cruzoe GIL & DoMÍNc;oEz,1995 ( O. Seo Sowevor FRIEDRiCO, 1995Srrsxricsitxi’uicxr, OCIIURRENIIE: Middle Cambrian (St.David’s)GizoGreAuriicAi. DISTRiBiJTION: Europe (France, Spain)

REFERENCrS: JAEKETI, 1918; FaíEDaícH, 1993: GIL CID &Dosrissjuszz, [994

Gyrocystis testudiformis FRIEDRIcH, 1993SircÁríorcái’HicXL OCCURRENCE: Middle Cambrian (St.David s)GFOGRAPHICAI. DiSTRIDUTION: Europe (Spain: Aragón)

RBr-FR11NU11S: FRrtzDrzrCH, [993:Grí. Cro & DOMiNGUEZ, [994

Ludwigicinctus truncatus FR¡EDRtCH, 1993STRÁTIGRÁPHIcsí, OtXiURRENCE: Middle Cambrian (St.David s)GEoGRxPHicM. DiSTRiBLTiON: Europe <Geosraaoy)

REFETRENCES: FarErsRrce. 1993

Progyrocystis disjuncta FR¡EDRICH, ]993SraxrrcsrsxPHrcAL OCCnURRENCE: Middlo Cambnian (St.Davids)GEíx;RxuííícsxL DiSTRiBIJTION: Europe (Spain: Asturias)

REETERENCES: FarEDRícír, 1993

Sotocinctus ubaghsi SDzuív, [993STRATIORAPHICAL ()CCURRENCE: Middlo Cambrian (St.David ‘s)GEoGRAPIiICÁi, I5iSTRIBUTiON: Europe (Spain: Asturias)

REFETRENCES: SDZUY, 1993

Sucocystis acrofena FRItnRicu, [993STRxrícRxuísrrcxL OCCItRRFsNCIS: Middle Cambrian (St.Davids)

46Coloquios de Paleouísslogia21)02, 53: 33-(sX

Patricio Domínguez Alonso & el al, An anriolated check-iist of genera and species of carpoids

GEOGRAPHICAL DiSTRIOUTIoN: Europe (France)

REFERENCES: FRIEDRICO, 1993

Sucocystis bretoni FRIEDRICH, [993STRATIGRAPHICAL OCCURRENCE: Middle Cambrian (St.David’s)GEOGRAPHICAL DISTRIBUTION: Europe (France)

REFERENCES: FRIEDRICH, 1993

Sucocystis? maroccana (TERMIER & TERMIER, 1973)[=Cyelocincíusnraroccanas TERMIER & TERSIIER, [973]STRATIGRAPHICAL OCCURRENCE: Middlo Cambrian (St.David’s)GEOCRAuHICAL DISTRIBUTION: Nortisoro Africa (Morocco)

REFERENCES: TERMIER & TERMIIsR, [973; FRIEDRICO, 1993

Sucocyslis melendezi (SCHROEDER, 1973)[=Trochocinctuspardalíhanicas TETRMIER & TERMiER, 1973]STRATIGRAPHICAL OCCURRENCE: Middle Cambrian (Sí.David’s)GEOORAPHICAL DISTRIBUTION: Europe (Spain, France)REFERENCES: SCHROEDER, 1973; FRIEDRICH, 1993

Sucocystis quadricornuta FRIEDRICO, 1993STRATIGRAIHICAL OCCURRENCE: Middle Cambrian (St.Davids)GEOGRAFHICAL DISTRIBUTION: Nortisem Africa (Morocco)REFERENCES: FRIEDRIcH, 1993

Sucocystis theronensis CABIBEL, TERMIER & TER-

MIER, 1959STRATIGRAPHICAL OSICURRENCF: Middle Cambrian (St.David’s)GEOORAPHICAL DISTRIBUTION: Europe (France)

REFERENCES: CABIBEL el al., 1959; FRIEDRICH, 1993

Sucocystis undata FRIEDRtCH, 1993STRATIORAFHICAL OCCURRENCE: Middle Cambrian (St.David’s)GEOORAPHICAL DISTRIBUTION: NertSern Africa (Niorocco)

REFERENCIeS: FRIEDRICR, [993

[Trochocinctaspardail/ranicas TERMIER & TERMIER, 1973]=Sacocystis melendezi (SCHROEDER, 1973) —9<

REFERENCES: SCHROEDER, 1973; TERMIER & TERMIER, 1973;FRIEDRICH, [993

[Troc/tocystitesBarrandei MUNIER-CHALMAS & BERCERON,

[889]=Elliptoeinctas barrandel (Mu’NiISR-CHALMAS & BERGERON,[889)REFERENCES: MUNIER-CHALMAS & BERCERON in BERGERON,

1889; FRIEDRICH, 1993

[Troc/tocystisbohevnica]=Trochocystites boheenicus BARRANDE, 1887 —>

REFERENCES: HAECKEL, 1896 FRrEDíuícH, 1993

Trochocystites bohemicus BARRANDE, 1887STRATIORAPHICAL OCtIURRENCE: Middle Cambrian (St.David’s)GEOGRAPHICAL DISTRIBUTION: Europe (Boisemia)

REFETRENCES: BARRANDE, 1887; JAEREL, 1918; FRIEDIIiCH,

1993; DoMÍNGUEZ, 1999

[Troc/rocysíitesIon gífossatus JxEKEL, 1918]=Troc/toq’stites bo/temicus BARRANDE, 1887 —~

REFERENcES: JAEKEL, 1918; GISLÉN, 1930; FRIEDRicFt, 1993

[Trigonocystisovalis HAECREL, 1896]=Troe/rocystites boheinicus BARRANDE, 1887REFERENCES: HsWCKEL, 1896; FRIEDRICH, l993

LTrigonocystis ¡rigona HAECKEL, 1896]=Troc/tocystites bo/temicas BARRANDE, 1887 —>

REFERENCES: I-[AEcKEL, 1896; FRIEDRICH, 1993

Trochocystoides parvus JAEKEL, 1918STRATIGRAPHICAL OCCURRENCE: Middlo Cambrian (St.David’s)GEOGRAPHICAL i)iSrRIBUTIoN: Europe (Boisemia)

REFERENCES: JAEKEL, 1918 UBAOIIS, 1967a; FíxaEDííícu,

1993; SDZUY, ¡993

Trochocystoides phanus SDZUY, 1993STRATIGRAPHICAL OCCURRENCE: Middlo Cambrian (St.David’s)GEOGRAPHICAL DISTRIBUTION: Europe (Spain)

REFERENCES: Sr,zrsy, 1993

Ctenocystoidea ROBISON & SPRINKLE, 1969

Ctenocystis cotodon UBACHS & RoBísoN, 1988STRATIC,RAPHICAL OCCURRENCE: Midd[e Cambrian (Sí.David s)GEOGRAPI4ICAL DISTRIBUTION: NortS America (USA: Utah)REFERENCES: UBAGHS & RoBísoN, 1988; DOMíNGUEZ, 1999

Ctenocystisjagoi JELL, BURRET & BANKS, 1985STRATIGRAuHICAL OCCURRENCE: Middle Cambrian (St.Davids)GEOGRAPOICAL DISTRIBUTiON: Australia

REFERENCES: JELL ci al., l985; DOMÍNGUEZ, ¡999

Ctenocystis smithi UBAGHS, 1987STRATIORAFHICAL OCCURRENCE: Middle CambrianDavid ‘s)GEOGRAPHJCAL DISTRIBUTiON: Europe (France)REFERENCES: UBAGHS, 1987; DoMÍNGuEz, 1999

(St.

Ctenocystis utahensis RoBísoN & SPR¡NKLE, 1969STRATIORAPHICAL OCCURRENCE: Middle Cambrian (St.David’s)


Patricies l3sssuíngs¡ez Alonso & es al. An ansmotated check-Iiss ol genera and spoc’es ol csrrpoids

GEcXRAPIIICAI, DISOSIIIUTION: Nortis Amorica (USA: Utah,IdaSo)REETI?RF.NCETS: ROsSISON & SuRíNarE, [969;UBAGH5 & ROBS-SON, [988a;DOMÍNGUEz, [999

Etoctenocystis bohemica FATKA & KoRDUIri, [985STrsxsrc.rvxurrrCAs. OCCIIRRENCE: Middle Cambrian (St.i)avids>GEcscRxuI iICAL DiSTRIi)IJTION: Europe (Boisemia)

REFERENCES: FATKA & KORDr’LE, 1985; DsMÍNGS.s,z, [999

Jugoszor’ua archaeocyathoides IWiK & ORaewsKt,1995

STRATIoRAuHICAr. OCCs)RRENCE: IVliddle Cambrian (St.D;rvids)(ieOc;rsAuHIuc DisrRirsUrioN: Europe (Poland)

REFERENCES D~rie & ORtOwsKr, ¡995; DOMÍNGUEZ, [999

SoRata JAEKEL, 1901

Betemnocystites wetherbyi MtLLER & GIJRI±Y,1894

Srax-IIGRAPHIcAI. OCCURRENCE: 1Jpper Ordovician (Cara-

doc)

GETíKRArrIIeái, DISTRrIIUTION: Nortis Amorica (USA)REFERENCES: Mír.siyR & GIsRIEY, 1894: PARSIEY, [972;KoIArss et aL, 1977

Castericystis vail UBAGOS & ReisísoN, ¡985SraArIoaAioicxr OCCI¡i<tENCe: Middle Cambrian (St.David s)GeooIsxí’tiit:AL DisTRIBUTION: NoriS America (USA: Utah)P.ETFeRLNcr.Ñ 1 litasais & RnloÑesN 1 OSS QieR- FThs ev 005

JEFrETRIES, 1997

[Gaster/czsíis? sprinkíe/ UBAOH5 & RonísoN 19881= Coleicarpus sprinklei (UiaAGriS & ROBíSON, 1988)RFFEISENCES: UBAGOS & RouísoN, [988;DAs.ey, 1996

Coleicarpus sprinklei (UBAGES & RouísoN, 1988);DALES’, 1996

[=Gasrerievst/.sYsprinklei llusutís & RonísoN, [988]SrRsricrtspírsc:xL OCCIJRRENCET: Midd[e Cambrian (St.Davids)GEoGRAFIIICAr. DrSrRrrsLsrrON: NorIS Arnerica (USA)REFERENCUS: Unxcvrs & ROHIsON, [988: DALIiY, [996:JEETFE-

RíES, 1997

Dehmicystis globulus (DEHM [934); CASTER, [967L=Dcndrocysliles gb/ralas Disus> 19341STsexrreRAr’urcx¿ OCCURRENCE: Lowcr Devonian (Emsian)

GIIOGRAr’HiCsL DISTRiBOrION: Europe (Gornrany)

REI±acNceS:DeriM, [934; Grír & CASTER, [960; CASTCR,1967: Ks’Tsí:ursa, 1975: BARTETLS er al - 1998; HAisDis,191>3

Coloquios de Palers,srssbo~ía

Dendrocyslites barnandel (BATHER, 1913)[=liScndcos:ystisbaícavrdei BATIIER, 19134 nom.van.]STRATIGRAPHICAL, OCCURRENCE: Middle Ordovician (Llandoi-lo)GEOGRAPHICAL DISTRIBUTION: Europe (Boisemia)

REFERETNCES: BATIIER, 1913

Dendrocystites sedgwicki (BARRANDE, 1867);L3ARRANDE, 1887

I=Cystidea sedgwicki BARRANLSE, 18671STresTIGRxuurCxí. oCCI;RRENCE: Middle Ordovician (L[andei-[o)GF.oGRAPHICAs, DISTRIBUTION: Europe (Bohemia)

REFERENCES: BARRANDE, [887

fDendrocysliteze V,dali TIJORAL, ¡935]—Mioe,vaecystis vidaíi (TíroRxs., l935); UBAGLIS & CASTER¡967 -~

REFERENCES: TíroRAr.. 1935b Unxc;íis & CASTER in CASTER,

[967

Dendrocystoides scoticus (RAnIER, 1913); JAEKEL,

1918I=Dendrocvsíis scotica BATHER. 19 l31STRATIoaxvniICxL OCCURRENCE: Upper Ordovician (AsSgill)GEOGRAPHICAL DiSTRiBUTION: Europe (UK: Scotland)REFERENCES: BATHETR, 1913 JAEREL, ¡9l8

«Dendrocys tites» rossicus JAEKEL 1901[=Dendrocysíitesross/caNOTE: tSe genoric attribution is doubttul because only tailessicles from Sistorical col¡ections are known,Siíasricrsxr’HiCxL O(:CUi<RENCC: Lewer-Middle Ordovician(Á¿dí~~LláíiVirb)GeOGRAí’iiíCxi. DrSrRIBrJIION: Europe (Russia, Estonia)

REFERENI:ES: JAEREI.. 1901; BxrursR, [913;HFCRER, [940;RozriNov & JEFFERIiS5, [996

Girvanicystis batheni CASTER, 1968STRATIGRxr’urcAL OCCURRENcE: Upper Ordovician (Asisgilí)

GEOGRAPIIICAL DIsTRiBIrrION: Europe (UK: Scotland)REFERENCES: CASTER, 1967; DALEV, 1992

Oirvanicystis casteni DAI.EY, 1992STrtsriGaxuHiCxL OCCURRENCE: Upper Ordovician (AsSgill)GEOGRAPHíCAL DiSTRiBUTION: Europe (UK: South Wales)REFETRENCES: Dxrtsv, 1992

Heckenicystis kuckersiana (HECKER, 1940); Gui. &CASTER. 1960

=Dendrocvst/tes /eackersianus HECRER, 1940]SrRxsIGRArriícxi. OCCIJRRENCF.: Ordovician (Llandeilo)

GEOGRAPH5CAL DISTRiisUTION: Europe (Russia)

REETERENCES: HECKER, 1940; GILE & CASrER, 960; RozriNov& IEETFERIES. [996

2002, 53: 33-6848

Patricio Domínguez Alonso & es al. Arr anísmotaled check-iist of genera sud specÍes of carpoida

Iowacysds sagittaria THOMAS & LADD, 1926STRATIGRAPHICAL OCCURRENCE: Upper Ordevician (Caradoc-

Asisgilí)GEOGRAPHICAL DISTRIBUTION: Nortis Ameíica (USA: Iowa)ReraleaNCes: KoLsTsetaí., [977; PARSLETY, 1972; THOMAS &LADi), 1926

MaennUia estonica Rozonov & JEEFERtES, 1996STRATIORAFHICAL OCCURRENCE: Upper Ordovician (Caradoc)GEOORAPHII:AL DISTRiISUTiON: Europe (Estonia)

REFERENCES: RozHNov & IEPFERíES. 1996

Minervaecystis vidaui (THORAL, ¡935); UBAOHS &

CASTER 1967[=DendrocystilesVidaíi THORAL, l935]STRArIGRAPHICAL OCCURRENCE: Lower Ordevician (Trema-

doc-Areoig)GEOGRAPHICAL DISTRIBUTION: Europe (France)

REFERENCES: THORAL, 1935b; UBAc;r-rs & CASTER in CASTER,

1967

Myeinocystites crossmani KOLATA, STRtMPLE &LEVORSON, 1977

Myeinocystiles natas STRIMF5F,, 1953 (pars); in KOLATA,1975: p. 15; pl.

2 figs. 810]STRATIORAPHICAL OCCURRENCE: Upper Ordovician (Caradoc)

GEOGRAPHICAL DiSTRIISUTION: NoriS America (USA)REFERENCES: KOLATA, l97$ KOLATA et al., 1977

Myeinocystites natus STRIMPI±,[953STRATIcRxuuícAL OCCURRENCr’: tlppor Ordovician (Caradoc)

GEOGRXPHICAL DISTRIBUTiON: Nertis A,neiiea (USA: Oklahoma)RFPERENCETS: STRIMPLE, 1953; PARSLEY, ¡972; KOLATA, 1975;KoLATAetO/., 1977

Rutroclypeusjunori WIWERs, 1933NOTE: Described in WITHERs, 1933 as a xiphosuran Dr kingcraSSTRAriGaxuviICxL OCCURRENCE: Lower Dovonian

GEOGRAuHICAL DISTRIBUTiON: Australia

REFERENCES: WITHETRS, 1933; JEIí. & HOLLOWAY, 1982

Rutroclypeus victoruae GILI. & CASTER, 1960NOTE: GiLL & CASTER, 1960 doscribed tSe firsí carpoidsrecorded from Australia, inc[uding Vicloriacystis wilkinsi,Ratroclypcas victoriae and Ratroctípeas withersi, They alioproposed a new classifrcation for tSe Carpoidea invo[vingíwe new subclasses and íwo new superorders.

STRATIGRAPHICAL OCt?LtRRENCE: Lower DevonianGeograpisical distribution: Ausiralia (Victoria)REFERENCES: GILL & CASTER. 1960 JELL & HOLLOWAY, 1982

Rutroclypeus withersi Gui & CASTER, 1960NOTE: seo note for Ratroclypeas victoriae GILL & CASTER,1960

SrRATIGRArHicxL OCCURRENCE: Lower Devonian

GEOGRAPHICAL DISTRIBUTiON: Australia (Victoria)

REFERENCES: GILL & CASTER, 1960; JELL & HOLLDWAY, 1982

Scalenocystites strimplei KOLATA, 1973STRATIGRAPFIICAL OCCURRENCE: Upper Ordovician (Caradoc)

GEoGRAPHICAL DISTRiBUTION: Nortis Aanerica (USA: Minnesota)REFERENCES: KOLATA, 1973a, 19735; KOLATA cí al?, 1975:Kolissrx e/al,, 1977

Syringocrinus panadoxicus BtLLINGS, 1859STRATIORAuHICAL OCCURRENCE: Upper Ordovician (Caradoc)GEOORAFHICAL DisTRmtríoN: North America (Canada: Que-

bec, Ontario)REFERENCES: B[LLINGS, 1859; PARSLEY & CASTER, 1965; Cxs-TER, 1967

Syringocninus sinchaini PARSLEY & CASTER, 1965STRATIGRAPHICAL OCCURRENCE: Midd[e-Upper? OrdovicianGeooraAuHícxL DISTRiBUTiON: NorIl, America (Quebec)

REFERENCES: PARSLEY & CASTER, 1965; CASTER, 1967

Comuta JAEREL, 1901

Acuticarpus delticus SUMRALL, SPRtNKLE & GUENS-BURO, 1997

STRATIGRAPHICAL OCCURRENCE: Upper Cambrian (Merionelis)

GEOGRAuHICAL DISTRIBUTION: North America (USA)REFEPENcES: SUMRALL eíal., 1997

Acuticarpus? nepublicensis SUMRALL, SPR1NKLE &GUENSBURG, 1997

STRxríGRAuorCM. OCCURRENCE: Upper Cambrian (Merionetis)GEOGRAuHICAL DISTRIBUTION: Nortl, America (USA)REPERENCES: SUMRALL e/al., ¡997

Amygdalotheca gnffei UBAGHS, 1969STRATIGRAPHICAL OCCURRENCE: Lewer Ordovician (Arenig)

GEOORAPHICAL DISTRIBUTION: Europe (France)

REFERENCES: UBAGHS, 1969; Cr¿íuus, 1989

Anupelocarpus handeyranensis LEFEBVRE & VIzCAÍ-NO, 1999

STRATIGRAPHICAL OCCURRENCE: Lower Ordovician (Arenig)

GEOGRAPHICAL DISTRIBUTION: Europe (France)

REFERENCES: LEFEBvRE & VIzCAÍNO, 1999

Anauricystis pñmaeva (THOiáL, 1935); LEFEBVRE &VízcAiNo, 1999

[=Coíharnocystispe/inaeva THORAL, 1935]STRATIGRAPHICAL OCCURRENCE: Lower Ordovician (Arenig)

GEOGRAPHICAL DISTRIBUTION: Europe (France)REFERE.NCES: THORAL, [935a; UBAGHS, 1969; LEFEBVRE &VIZCAiNO, 1999


Patricio Desnsíng’uez A Ososo & cl al, As anírosated check-liss of gesrera and species of caepoids

Arauricystis occilana (UBAGHS, 994>; [±FEBVRE &VIZCMNo, 1999

1 =Cot/suroescssíis occilano Usxuy i~, 1 994]STRMIc;wxr’srrcAt OceuRRENCE: Lowor Ordovician (Areoig)(jevsv¡RsrtliU¡sL OrSrRsBUtroN: Europe (France)RevewizNces: Usssí}ts, 1994: t.sIs45x11r- & Vrzcsi~o, ¡999

Archacocothurnus bifidus (UBAGLIS & ROBISON,[988); SísMaÁri., SPRINaIrs & GUENSBURG, ¡997

VAsz hoecscoí/ss,rssus bifle/a (UIsNGirS & RoarsoN, 1 981=):Su:ssaArL. St’aiNÑs.e & Gtsesssr:rac, 997 (incorrecí gonder)]

Coí/rsrrnoevsíis Y 01//e/a UISAGHS & Roersoa. ¡988]SrasTss;saAurlrzÁr OUUURRI:NUiL: Middle Casnisrian St.

Dsrvid síGEToovsAuitsc:sL DISTRIEI:Tr(sN: Norih America (USA)Rssr’s:tsscr’s: 1 Jnxe;srs & Rrsssrscss. [988: Suwirexs±et aL,¡95>7

Archaeocothurnus goshutensis SIJNIRAI.t,, SÍ’iz[Ní<Lui

& GUENSHURG, 1997St RAittRS Ai’it >5 ‘Xi EZí IR RENí E~ Upprr (arr Sriarr MeriorretS)Gtls(IRAr’rttt;.Ni s)rsrrSrrtI[TI(tN: Nortis Arríerica (USA)E IYtSRU:NCUS: .1=ti M RAIL er al, - 1 997

Beryllia miranda CRIPES & DALiIY. [994Srrs.Níss;rs,xpssscxL t)cUs:RRs:NUF:: Middle Ordosician (Llandol—

GvocaAs’wcsr. uSSTSSiEL’TSON: Europe (France)Rt¡rersETszcES: Ciesuus & DAr Ev [994: RL’EX, 19995: LEFEN-Bise, 20(>t>a

Babinocystis dilabida RohfiNON, 1990- St R\rsGRáuI5sCAi (5CUURRETNS?E:Lower Ordovician (Arenia>

(¡FOCRNFIttt’Ai DíSiRiissznroN: Etíropa Rsrssia: Sainí Peters-

bourgtRETI-Crss-Ncrs Ro’rssov, ¡990

Bohemiaecystis houceki CASTER. [967[4/ro ses/ics’sli.s bossce/zi CASTER, [967;LrozvsvRE & VrzcM-NO, [999]Nota: See note i o tSe ersrtry of Bohc’vnisaeevsris Cxsrers. 1 967ooder liso secliosr «Check-list of Generas’ ( eSnrsArst¡ÑÁr’iaseAI, tsCUs:’RRJ¡NUs:: Middle Ordoviciarí (lían-s’im-t.Iarndei lo?’>Gt¡o;íAuísscAí ursrierBt:Ti(sN: Europe (Bohemia)Rerarzse=crs:Cssruslo UaAcrts, 1 967c: GIL Cruel al,, 1996;LEFras-vssr: & Vr/cANo, 1999

Bohemiaecvstisjefferiesi GIL Cíu. DoMÍN;íw:z. SIL-VAN & EscRluxNo. [996

[=Seociaesysri,s¡etjériesi (Grr Ciu, DtsMiNc;UEZ., SsLvAS &Esciss}ssrscs. 1996>: LEeizrsvko & VizcAÍNo. [999]Nontz: See note rs tSe entra of Bohemiaecsstis CASTER, [967sínsier tSe section ss(’Seek-list of Cerrera”STPsrse;Rsrr 55UAL O(cs’RRRNCs:: L

tpper Ordevician (Caradoc)

<?o/ssqssisss dc Psr/eesnte,lesgíss2002. 53: 3368

GEOGRAPHICAL D5STRiBUIiON: Europe (Spain: Castilla-LaMancha)RETFs¡RETN(sS, Gsi. Crí, ce al., [996; LETFEON’RE & VizeXÍNo,1999

Bohemiaecystis guilloal (LEI:UBVRE & VIZCAÍNO,1999); new combinalion Serein

[=Sc:oliaecvslisgai/íoai LEFEONRE & VIZCAÍNO, 1999]NoTE: LEETEBVRE & VIZcAÍNo, 1999 described Ibis speciesunder Ssot/oeev.stis CrssTri & URACIIS, [967( 9. Seo note oliso ontry br Bo/rernioec-rsíis CASTCR, [967under liso section«Check-list of Genera” ( 9SrssxrrGkss’uresL ocUURRETNCET: Midd¡e Ordovician (Llandeile)GEoGRsursscAr. DISTRtuLsrrísN: Europe (France)RISFERETNCES: LEETEBVRa & VszCMNo, 1999

Ceratocystis pernerí JAEKEL, 1901STreArscsssurrsí:AI. csct’u:RRE\cF: Miádlo Cambrian (Sr.1)avids>G5¡osiaXnnrc,x¡, r)ss5RIBUrioN: Larope (Bohemia)Rshi’orwNctss: JACaEL, [901; UBAOHS. 19675, 1967c; JFFFE-RíES, 1969, l986

Ceratocysti?~ vizcainol UHAGHS. [987Ssaxriciesrttrexú tXxtiRRETNct: Middle CamSrian (St.Davids)GIzoGRAvIsscxr DiSTRtBUTION: Europe (France)

Retrzkrtsees: UBAGES. 1987

Chau velicystis spinosa UBAGIIS, 1969pos-lico =Chaavclis:vstis s’izcoiooi i?1.xí rtv, 1992STíxrrGRAuHrexL OCVURReNCET: Lower Ordevician (Arenig)Gwoe;RsrwrIcss. i)sSTRIRI:TIoN: Europe (Franco)

RFFRRFNUFS tli)NU,HS, 1969, 1983:0MEv. 992

Chau velicystis ubaghsi (CHAUVEL,. 1966); UBAGHS,

[969[=Coíhurnocs’stisabaghsi CírsovEs., 1966]SrRxrrGRAuHsexi. OCCURRENCE: Lower Ordovician (Arenig)GLOGiSsPtaseAL DiSíRsuUrroN: Norlhem Africa (Merocco)RtzFessisrecls:CIom’vu,. 1966a, [971;UBAGHS, 969; DALeY,[992

Chauvelicystis vizcainoi DALEY. 1992STssxrrc,RÁumíIcXt, OCcURRS>Nciz: Lower Ordovician (Areoig)GE5GRAI’NíCAL OiSrRiOtflION: Eursmpe (France>

RLSLRENCLS: UBAGrsS, 983; Dss.n’, [992

[=Cothartrosvs/i.sY americano URAc,HS, ¡963]Nevae/aetyscis americana (URAGHS, [963);UBAGHS, 1967 >

REFERETNCET5: UBActís, 1963, 1967c; Jvr’r’EssrrzS, 1986; Jeara-RíES e> al., 1987

[Cotlrus-noc:zs/isYbrflda Urn’sc,us & Ronssors, [988]—Arc/raeocothus-,ras bífidas (UBAGOS & ReniSoN, 1988);

50

Pa/ricio Dominguez Alonso & el al. An arsuotated check-IisI of genera arsd species of carpoids

SUMRALL, SPRINKLE & GísENsnrsRc, [997 9<

REFERENCES: UBAGHS & RotrísoN, 1988; SUMRALL el al.,1997

Cothurnocystis courtessolei UBAGHS, 1969STRATIGRAPHICAL OcCURRENCE: Lower Ordovician (Arenig)

GEOGRAPIIICAL DISTRIBUTION: Europe (France)

REFERENCES: UBAOHS, 1969

[Cot/turnocvstiscurvata BATIIER, [9131=Scoíiaecystis cas-vala (BATHER, [913); CASTER & UBAOHS,

1967 ‘

REFERENCES: BATHER, [913; CASrER & UBAGHS in UBACTHS,

[969

Cothurnocystis elizae BArBER, 1913SrRArícRssruícAL OCCURRENCE: Upper Ordovician (Asisgilí)

GeocRAutucAL DiSTRii)LtTIoN: Europe (UK: Scoiíand)REFERENCES: BATHER, 1913, [926: UDAGOS, l963; JEEFERIES,

¡986; JEEFERIES etal., 1987; WooDs & JEFEERIES, 1992

Cothurnocyslis feilinensis tJBAC~HS, 1969STRATIGRAPHICAL oCCIsRRENc:E: Lower Ordovician (Arersig)

GEOGRAPHíCAL DISTRIBUTION: Europe (France)

REFERENIES: Uuxcus, 1969; JrzrpssierEs el al,, 1987

LCoí/racnocystis melc/riori UISAUSHS, [983]=Ps-osí:otiaes:vslis melchiori (Uuscsrís, 1983>; UBAGHS, 1994[=T/ros-alysy.stisurelch/ori (UBAGII>,1983)]REPERENCETS: UBAGHS, 1983, [994; CRIPPS, 1991; GIL Cro etal., 1996

[Col/rurnocystis occitana UBAGHS, 994]= Araaricystis occitana (Uusc;íís, l994), LEPEBVRE & Víz-CAtNO, 1999 ‘

REFERENCES: UBAcJHS, 1994; LEFERvRE & ViZCMNo, 1999

(Cot/turnocystis owensi (Wooos & JEPEERIEs, 1992), LEFEBNRE &VIZCAÍNO, [999]

=Procotlras-nocystis owensi Wooos & JEí’rEaIas, 1992 —>

REFERENCES: WOODS & JEEFERíES, [992;LEFEBvRE & VIZCM-NO, 1999

[Cot/rurnocyslisprirnaeva TIIoRAL, [9351=Arauricvslis prinaaeva (THORAS., 1935), LEPEBvRE & VIZ-CAÍNO, 999 9<

REFERENCES: THORAL, [935a: UBACHS, 1969; LEFEBNRE &VIZcAÍNo, [999

[Cotharnocvstisabag/tsi CHAIsvEL, l 966]=C/raavelicvstis abag/rsi (CrtsísvEL, 1966) —>

RPFERENCES: CHAUX’EI., 1966a; CHAUNEL, 1971; DALEY, l992

Domfrontia pissotensis (CHAUVEL & NIoN, 1977);CRtPPS & DALEY, 1994

[=Rel/calocarpos pissoíensis CHAISvEL & NioN, 1977]

STRATIC,RAPHICAL OCCURRENCE: Middle Ordovician (Llaodei-

lo)GEOGRAuHICAL DISTRIBUrION: Eurepe (France)

REFERENCES: CHAUVEL & NION, 1977; CRIPPS & DALEY,1994; LEFEvBRE, 2000a

Drepanocarpos australis SMITH & JELL, 1995STRATIGRAPHICAL OCCURRENCE: Upper Cambrian (Merienetis)

GEOURAFHICAL DISTRIBUTION: Australia

REFERENCES: SMITH & JELL, 1998

Fkzbellicystis rushtoni MARTÍ MUS, 2002STRATIGRAPHICAL OCCURRENCE: Lower Ordovician (Trema-

doc)GEOGRAPHICAL DISTRIBUTION: Europe (England)REFERENCES: MARTÍ Mos, 2002

Gafliaecystis lignieresi UBAGES, 1969STRATIGRAPHICAL OCCURRENCE: Lower Ordovician (Are-

nig)GEOGRAPHICAL i)ISTRIOUTIoN: Europe (France)

REFERENCES: Uuwris, l969; CRípus, 1989

Hanusia obtusa CRtPPS, 1989STRATIGRAPHICAL OCCURRENCE: Middle Ordovician (Lían-viro)GEOGRAI’HICAL DISTRIBUTION: Europe (Bohemia)REFERENCES: CRípus, 1989

Hanusia prilepensis CRIPPS, 1989STRATIGRAPHICAL OCCURRENCE: Middle Ordevician (Llandei-lo)GEOGRAPHICAL DISTRIBUTION: Europe (Bohemia)REFERENCES: CRIPPS, 1989

Hanusia sarkensis CRIPPS, 1989STRATIGRAPHICAI. cXYCURRENCE: Middle Ordovician (Lían-viro)GEOORAPHICAL DISTRIBUTiON: Europe (Bohemia)

REFERENCES: CRIPPS, 1989

Juliaecarpus milnerorum RUTA, [999SrrexrIcRAuHICsxI, OCCItRRENCE: Upper Ordevician (AsSgill)GEOGRAPHICAL DISTRiRUTION: Nortisero Africa (Morocco)REFERENCES: RUTA, 1 999b

Lobocarpus vizcainoi UBAGHS, 998STRATICRAPHICAL OCCURRENCE: Upper Cambrian (Merionetis)GEOGRAFHICAL DiSTRIIsUTION: Europe (France)

REFERENCES: UBAGHS, 1998

Lyricocarpus courtessolei UBAGES, 1994&rR*TíuRAí’uícAL OCCURRENCE: Lower Ordovician (Arenig)GEOGRAI’HICAL DISTRIBUTION: Europe (France)

REFERENCES: UBAOHS, 1994


Patr(s:io Dsss,síngucz.ilonso & es al, An annotated checic-liss of gesrera and specics of carpoids

Milonicystis kerfornei CHAUVEL, [986SisaiiorvXrHIc:ÁL OCCISRRENCE: Middle Ordovician (Llandei-lo)GtzoorssuríreAL i)IsTRInIJTIoN: Europe (France)Res LRLNCES: Críss’vr¡,, 986; CRur’s & DXs.EY, 1994

Nanocarpus dohambii UDAGHS, 1991STEsslcRwítscXL OCeURReNUL: Lower Ordovician (Aronig)GEOGRAPISICAL DiSiRiELTION: Europe (France)

REFCRLN Es: [BytEs. 1991

Nevadaecystis americana (UBAGHS, 1963); UBAGHS,

1967[=Coe/rarnoe:s’sei.sY amecicaoo UEXc.us, [9631STRTIGRXuuIeAs, OCCURRENíSE: Llpper Cambrian (Meno-

noIS)GaosjRAi’ííseXt DiStRIBUTsttN: NoríS America (USA)R>’raasz~cr:s: UBAGES, 1963, 1 967c; JEFFERIETs, 1986; JEFFE-

RíES ev al? - 1987

Phytlocystis blayaci Tíiortxí., 1935SrR’srrí;rtus]ress. OCCL’RRETNCE: Lowor Ordovician (Trema-doc-Arenig)GEocRAFHsCsí. i)ISTRiBUISON: Europe (France)REFCR11NUF.S: THORAL, 1935a; Grc;oí:T, [954; UnAGHS, [963,1969

Phyllocystis crassimarginata (TnoRÁu.. 1935)PhY’líotx’sC~s Ría :ssú y aricIé írassima vgioa va Tooieex

19351SrrsírrorvXuvrrCXú Oc:CURRiZNUrY: Lower Grdovici an l15ernsr-doc-Arenig)GLoGRAIosiU¡sL DISTRIBUTION: Europe (Frailee)

REFETRENCES: TUORAL, 193.Sa: UBAUSIIS. [969

Phyllocystis? salairica DuíBxrOeovA, [985L=Phvlloevse/.s Y salais-/cas DLBATOLONA, 1985 (incorrecí gen-der)]STRArIGRÁi’tsieAL ocCsSRRETNCET: Middlo Ordovician (Llassdei-[01Gszor;RÁPWCxi, Ot5tRIBIJTSON: Asia (Siborian Russia: Salair)

Rnr-sizsvLS- Dr axtOtOvA it, RosovA, Rosox & DUa.NroLo-“A. 1985

Ponticulocarpus robisoni SUMRALL & SI’r¿iNKIE.1999

STRxriGreADvrrísss. OCCIjRRENce: Middle CambrianGeooRvihIc:sí, rsísTRiSoiboN: Arnerica (USA: Utah)RETFFRINUFS: SIMRALL & SuarNKsE, 1999

Prochauvelicystis semispinosa DALEY, ¡992SrvsxTrszRXpvrre:..u OCCURRENeL: Lover Ordovician (Trema-

doc>GEoGRXí’uiCXs. i)ISTRIBU1ION: Europa (11K: Englané)Rv.rvursrn: Dsr.p:y. [992

Coloquios sil Pesle-esoeología

Procothurnocystis owensi WooDs & JEEFERIES,[992

[=Coehas-oocv.sei.sosve,asi (Wooos & JEFFERIES. 992>,LuTaNRE & VIzCAíNo, ¡999]SiRXsitsRXI’uic:sr, oCci;RRENcE: Lower Ordovician (Arenig)GEOGRAPH5CAL OISTRSSSLsIIoN: Europe (12K: SoutS Wales)RETFFRETNCETS: Wooos & Jer-r-euri-s, 1992. IFETEUNRE & VszcM-Nt), 1999

Progau¡iecystis ubaghsi CRíuPs. 1989STÑsrroRsurIICxI. OCCURStETNUET: Lower Ordovician (AronigíGETOGRAPHICAL i)ISTRiOLT ION: Europe

RCFeRENUES UBAUrIS, [969: CRIiI’s, 1989

Prokopicystis mergil CRIPES, 1989STRATIORAPFIICAI. OcCURRENCE: Midd[e Ordovician (Llandeilo)GeocavXPoic:rsL i)ISTRIBIJTIoN: Europe (Bohemia)Rvrvrsieces: CRípus, 1989

Proscotiaecystis me/chían (UBAGIIS, 1983); UBAGHS,

1994[=Co¡óus-uocvs/ismc/chioci L

tusc;rís, 1983]=Thoralicvsíis me/ch/oh (Ursscsís, 1983)1

STwsTscu-NursrcAs, OCCURRENcC: Lower Ordovician (ArenigíGEooRXuriiCXL i)IsTRiuonO~: Europe (France)REFETRENCES: UHAGES. 1983; LtB.NGIIs, 1994; CierrEN, 1991:GIL Cso ev al?, 1996

Protocystites menevensis HícKs, [872SrícsTrore’rtsrc-ss. oCCs’RRrINCF: Midéle Canbriao (Si.Davids)GEOGRAPrI5UAL Oi5TRI}31’TION: Europe (¡1K: Wa¡os)RFFERENUES: EiCKs hí HICaS & JONES__[172 JEFFERíES el al,,[987

Reticulocarpos hanusí JEEFERIES & PROI<or, 1972STRATIGRAFHíCAL OCCL’RRETNOS: Middlo Ordovician (Lían-ximiGEOGRAPOICAL DISTRIBIJTION: Europe (BoSomia)

RFsizRrzNCnS: JEPIFRIES & Przoieos’, [972; JEFEERIES, 1986:CRIFPS. [989

[Revialoeas-pos pissoteuisís CriAssvrsí. & NioN, 1977]—. Dovnfs-ontio pissovensts (CriAr551)1 & NION, 1977)REFERENCES: CHAP/FU & NIoN, 1977

Scoliaecystis co/lapsa CRIPES, [988STRAT5GRAPHICAL OCCusRRENCET: Ltpper Ordovician Asisgilí)GEOGRAPHIcAL i)ssTRIBI:TIoN: Europe (UK: Norihern Iroland)REFERENCES: Curpus, 1988

Scotuaecystis curvata (HAFHER. [913); CAsrílR &UBAGHS, [967

(=Govhuvnocvstis cus-tota Bmriizts, [913]STRAT5GRAPHICAL OcCURRENCE: Upper Ordovician (AsisgiS)

2>1>1223:33-6852

Patricio Donr/ssguez Alonso & el al, An aísrnotated check-iist of genera mrd species of caspoids

GEOGRAPHICAL i)ISTRIultzrIoN: Europa (UK: Scotland)REFERENCES: BATOER, [913; CASTER & UBAGHS in UBAGH5,[969; Jnrraieíns, 1967, 1968,1986; CRírus, 1988

I=Scoíiaecystis griffei UBAGHS, 19691=TIros-alicvstis gríffei (UBAGI iS, 1969); CHxusvEí., 197lREFERENCES: UBAGHS, 1969; CrsAs’vEs., 1971

[Scotiaecysíisgailloui LEFEBNRIS & VíZCAiNo, 1999]=Bo/re,n/aes:vstis gailíoai (LeversvaE & VIzCAÍNO, 1999),new combination Serein —>

REFERENI:ES: LEFETIWRE & VIZcAÍNO, 1999

[=Scotiaeczsrisjcfferiesi (GIL CID, DoMÍNGusiz, SILvAN & Escier-BANO, [996>;Leaenvae & Vizc:XiNo. 19991

Bohem/aecystis jefferiesi Grí Cío. DOMÍNGUEZ, SílváN &EscRIBANo, [996REFERENCES: GIL Cii) el al., [996; LEFEi)NRE & VIZCAÍNO,1999

[=T/ros-alicvs/isboacelci (CASrER, 1967), LEFEBvRE & VIZCAiNo,1999]

Boherniaecss/is boaceki CASTER, l967 —>

REPERENCES: CASTER la UBAISFis, 1 967c; GIL Cii) ev al., 1996;LEFEBNRE & VIZcAÍNo, [999

ThoTauicystis gnffei (UBAGHS, 1969); CHAUVEL,

97][=Scotiaesvstis gciffei UBA;írs, 1969]STRATICRAPHICAL OCCURRENCE: Lower Ordovician (Arenig)

GEeGRAuííícsL i)ISTRIBL’TION: Europe (France)

REFERENCLS: UBAGHS, [969; CnXr’vEI., 197l

[Thos-alicvstismelc/rios-i (UBAcurs, [983)][=Col/run,ocys/ismelc/riori Une« lIs, 1983]=Proscoviaecvs/is ,nelchiori (UBAOHS, [983);UBAGES, 1994REFERENCES: UBAUSHS, 983, [994: CRIPPS, 1991; GiL Cío eval?, 1996

Thonauicystis ubaghsi LEFEBVRE &VIZCAINO, ]999STRATIGRAPHICAL OCCURRENCE: Lower Ordovician (Areoig)

GEOGRAiHIcAL DISTRIBIsTION: Europe (France)REFERENCES: LEFEBNRE & VIzcAÍNo, 1999

Thonauicystis zagonaensis CIJALIvEL, 1971STrvsviískxvHiCxL OCCttRRENCE: Lower Ordovician (Arenig)GEtx;Rxuaic:s¿ DisTIIiBUTioN: Norrisero Africa (Merocce)

REETERENCCS: CHAUvEL, 1971

Tnigonocarpus singuhanis UHAGHS, 1994STRATIGRAuHICAI, OCCISRRENCE: Lower Ordovician (Aronig)

GEOGRAuHICÁL i)IsTRIBIsTIc)N: Europe (France)REFERENCES: UBAGHS, 1994

Mitrata JAEKEL, ]901

Adoketocarpus acherontkus RUTA & JELL, 1999SrrtATIC,RsxPHICAL OCCURRENCE: Upper Silurian (PIidoli)GEoGRAPIiICAL OiSTRiBUTION: Australia (Victoria)

REFERENCES: RUTA & JELE, 1999a

Adoketocarpusjaneae RuTA & JELL, 1999STRsTIorráríiicsL OCCURRENCE: Lower Devonian (Locisko-

vian)GEOGRAPHICAL DISTRIBUTION: AustraliaREFERENCES: RUTA & JETLL, 1999a

Atlanicytidium flemingí CASTER & GILL, 1967STRATIGRAPHICAL oCCIsRRPNCE: Lower Devonian (Emsian)GEOGRAPHICAL i)iSTRIBLrDoN: New Zealand (South Island)REFERENCES: CASTER & GILL in UBAGHS, 1967c; FLEMINO,l970; RUTA & JELL, 1999a

Anatzfopsis acuta BARRANDE, ]872STRATIGRAPHICAL OCCURRENCE: Upper Ordovician (Caradoc)

GEOGRAPHICAL DiSTRiBUTION: Europe (Bohemia)

REPERENCES: BARRANDE, 1872

Anatzfopsis ancona DOMINGUEZ & GUTIÉRREZ-MAR-

ce, 1990STRATíGRAPHICAL OCCURRENCE: Middlo Ordovician (Llandei-[o)GEOGRAPHICAI. i)isTRIBI:TIoN: Europe (Spain: Castilla-laMancSa)REPERENCES: DOMÍNGUEZ & GUTIÉRRr-z-MXRCo, 1990

Anat¡fopsis armoricana (CHAUVEL, 1941); DoMÍN-GUEZ & GUTIÉRREZ-MARCO, 1990

[—4naíiferocystisarmoricana CHAUNEL, 1941]STRATIRAPIIicAL OCCURRENCE: Middle Ordovician (Llandeilo)GEOGRAIHICAL DISTRIBUTION: Europe (France)

REFERENCES: CHAUvEL, 1941; DOMÍNGUEZ & GUTIÉRREZ-MARCO, 1990

Anatzfopsis balclatchiensis REED, [940[4 Anatífopsis barrandei (CHAUNEL, 1941); JEFFERIES &DALEY, 1996]STRATIGRAPHICAL OCCURRENCE: Upper Ordovician (Asisgilí)GEoGRxuHícxL DISTRiBUTION: Europe (UK: Scotland)REFERENCES: REED, 1940; JEr-PERIES & DALEY, 1996

Analifopsis barnandel (CEAUVEL, 1941); DoMÍN-GUEZ & GUTIÉRREZ-MARCO, 1990

[=Analiferocystisbarrandei CHAIsvEL, 19413STRATIGRAPHICAL tXCURRENCE: Middle Ordovician (Lían-viro)GEOCRAIHICAL DISTRIBUTION: Europe (Bohemia)REFERENCES: CHAUVEL, 1941; DOMÍNGUEZ & GUTIÉRREZ-MARCO, 1990


Patricio Domí,sgucz Alonso & el al, An annotated check-Iist of genera aud <pecies of carpoids

Anatifopsis bohemica BARRANDE, 1872STsvíxrrowxruieXL OC> URI*ETNCET. Upper Ordovician (Caradoc-

AslsrgiilY)GeoGrvXr¡rrCsL 0151 Ritís 5 l0N~ Europe (Bobomia)

Rí ETERIN> es: BXRRANDE. 1872

Anatifopsis cocaban KOBAYASHI, [960STRXrss;RXartseXs OCCIJRsZI:NCE: Lowcr OrdovicianGETR;RAFHICAL i)sSrRiBs,’rsON: KoreaRssFERIZNES: KorssxASHI. [960 DoMÍseistEz & GUTIÉRREz-

MARCO, 1990

(A ,sovifispsis cactus II ADi)INe;, ¡913]=YSoíe,ísscasis7 comer (Hsouíso, 1913)Nora: originally described as cirripeds trinavifops/s was Ilionconsidered crastacearr rernains) buí probably are isolalod pia-les of a polyplacopSoran irsolluse, Seo also Anaíífopsis e/em-galas (II ‘xi)i) INII. 1 913)> >) and A .rcstifopsi.s romes- 1 Hsssoisc,¡9l3>( >)Ssaár1s3aXrrdse:NI. 0CCrJrss<tzNCE: Uppor Ordtwieiao(ieooaXlritcXL DISTRIBUriON: Europe (Swedeo)

Rn>’aucscuss: Hsrsoisc, 1913, 1915; DoMÍNGUEz & GL’rrÉ-RRI)-MARtV5. 199>)

[Anatifopss?scloogaías HADDINUS, 1913]=Soíeoocaris elongatas HAi)i)INc. ¡913; Ssno & [lOARE, 1987NOTE: originally describod as Analrjhp.sis, Solenocaris clon-gatas consisís of isolated piales of a septeroehitonid (Mollus-ca Polyplacophoca). Scc also Aoatifopsis castas (HArDING,¡913) ( aod Anatifcspsis vesrvres- (Hái)DINc;, 1913) f—>)Sri<srsoRss’me.xr. OCCURRUNCe: Upper OrdovicianGeOcsRAeoIc:Xi, i)ISTRiBL:TIs)N: Europe (Swoden)REFERENCES: HADDINO, 1913, ¡915; SMITH & FInARE, 1987;DoMrss;sez & lis ~ ~~>rs1000

Anatífopsis escandei escandei TíioRAus, 1935

=Balttnoevst/ve.s escandti (TísoRAs., 1935): ÑIZCACNo &Leaersv~re, [998]STRArrcRexuí rUAr, OCCsSRRETNcET: Lower Ordovician

GEOGRAPHICAI. orsrarrs~~roa: Europe (France)REs’IzRIsNCsss: TuoRsí., 1935a; CNAs;ser,, 1941; JEFFERiES,[987; DoMÍNcrsEz & GLTIÉRRETZ-MARCO, [990; VIzCMNo &LtzrtvBssE, [998

Anatifopsis escandei? thorali (CHAUVEL, ]966);DOMÍNGUEZ & GLsríEnivez-MARdo, ¡990

(=Analiferescyslis esca,ídeiY 1/ros-ah CHileNa,, 194STRXriGRAPHICse (XiCURRENCE: Lower Ordosiciao

(‘sraocsRssuorcss. tsISTISIBUTION: Norlisoro AÍrica (Morocco)

Reruen=ices:CtixísveL. 941, 1966b; DOMÍNGUeZ & GuSTÉ-RRETZ-MARIKS, l990

Anahfopsis guichenensis (CHAUVEL, ]941); DoMÍN-(SUEZ & CUTIERREZ-MARCO. 1990

[=Anavifes-oevsv/sgaichenensis CHAusvEr., l 9413STrasTIcRsuurcsr. ocCIsRRFsNcF: Middle Ordovician (Llandeilo)

GEOGRAPHICAL DiSTRIBUTION: Europe (Franco)REFERENCES: CHAUvETL, 1941; DOMÍNGUeZ & GUTIÉRReZ-MARCO, ¡990

Anatífopsis longa BARRANDE, ]872STRATIGRAFHICAL OCCURRtiNCL: Upper Ordovician (Cara-

doc)GEOGRAPHICAL DiSTRSBUTION: Europe (Bohemia)REFERENCES: BARRANDET, 1872

,4natifopsis minuta (CHAusvEl., ¡94!); LEFEBVRE,

2000f=Asrat~/bpsis bacrandei minuto (CríXrsvar., 1941): Dor

5rÍN-GLEZ & GUTIÉRREZ-MARCO ¡990][=Ana/mjitrocvstisbarcandei mina/a CHAUVEL, 1941]STRATIGRAPHICAL occsSRRENc?E: Middle Ordoviciarsr (Llandei-¡o)Gt¿oGRAPHiCXu i)ISTRiBUTIoN: Europe (France)RFFFRFNCF CHsísvEs., 1 941 - [982; DoMÍNGIsEz & GIsTÉ-RRFZ-MARcO, ¡990; LEETEN’BRE, 2000a

Anatifopsis papi//ata (BASSLER, [943)l=Kirkocystis papi/lota IBAsSLER, 1943)][=Enoplous-aYpapilíava BASSLER, [9433STRXiIGRAPHIcAL OCCURRiLNCE: tlpper OrdovicianGEOURAI’HICAL DISTRiBUTION: NorIS America (USA)RaPERaNCeS: BASSLER, [943, [950; PARSLEY, 1991

Anatsrfopsis punctata (BASSLER, ] 950)Enoploara punclata BAsSLEII. 1950]

[=Ki,kocysvispunclala Bxssrnrs, [950]STRATIcRAuHrcssí. OCCURFsENCE: Middlo OrdovicianGancraAuIlICsm DISTRIBUTION: NoríS America (USA: Tennes--——5.SLLJRETFETReNCES: BA5SLETR, 1950; Psrísr.ísv, 1981

[Anattfopsisps-/ma BAR/ANDE. 1872][=Anatrfops/sprimas BAR/ANDE, 1872 (incorrecí gender)][=Sagillaeystis prima BARRANDE, [872:PARSLEY, PROKoP, &DERSTLER, 2000]=Balanoc-ystives pr/mus BARRANDE, 1887REFERENCES: BARRANDE, 1872, l887; JaFaErírEs, [986;PARS-LEY, PROKOP, & DERSTLER, 2000

Anatifopsis spinosa UBAGHS, [979STRATIORAPHICAL OCCURRENCE: Upper Ordovician (Caradee)

GeoGrs,uuIexL DISTRIBOTION: Europe (Bohemia)REFERENCES: 1JBAGIIS, [979;PETR, ¡984

Ánatifopsis tnapezijiormis THORAL, [935[=A triapezi]formis THORAL, [935 o DOMíNGUez & Gutie-rrez. 1990, p. 135 (incorrecí spol[ingflSTRATIGRAPHICAL OCCURRENCE: Lower Ordovician

GEOGRAPHICAL DiSTRiBUTiON: Europe (France)REFETRENCES: TFIoRAI., 1935a; DoMÍNCs’Pz & GIjireRRez-MARCO, [990

54(?csloquios de Pe¿/contsslesg/o2002, 53: 33-68

Patricio Domínguez Alonso & el al, Aa asrmnotated clieck-list of gonera and species of carpoids

Anatifopsis traveusotensis (CHAIJVEL, 194]);DOMÍNGUEZ & GuTIÉRREZ-MARCo, 1990

[=Anatífes-ocysíisirar’eusolensis CIIAUNEL, 1941]STRAOGRAPHICAL OCCUBRENCE: Middlo Ordevician (Llanviro)GEoORAFHIcAL oISTRIBUTION: Europe (France)REFERENCES: CHAUvEL, 1941; DOMíNGUEZ & GUTIÉRREZ-MARCO, [990

Ánatifopsis truncata KoBAYÁshíí, 1960[=Anat~fopsistrancature KorsrArtuHI, 1960, incorrecí gender]STRsrIcRAuíIIcsrL OCCURRENCE: Lower Ordevician

GEOGRAIHICAL i)iSTRiBUTiON: Asia (Korea)

REFERENCES: Konsyssrii, [960; DOMÍNGUEZ & GUTIÉRREZ-MARCO, 1990

LAnatifopsis comer (Hsi)DING, 1913)]=Solenocaris? vomer Hsi)i)rNo, [913; DOMíNGUEZ & GUTIÉ-RREZ-MARCO Marco, 1990Noru: originally described as Aoatifopsis tisese are iselatedplatos of a polyplacophorao mollusc, Seo alse Anaíifopsiscanas (HADDING, 1913) (—>) and Anatífopsis elongatas (Riso-OrNO, [913) ( QSTRATIGRAPHICAL OCCURRENCE: Upper OrdovicianGEooíur’HIcAL DISTRIBUTION: Europe (Sweden)RutRENcrsS: HADOINO, 1913, l915; DOMÍNGUEZ & GUTIÉ-RREZ-MARCO, [990

Anomalocystites cornutus (MILLER 1889; Subse-quení designation) HALL, 1859

STRATIGRAPHICAL OCCURRENCE: Lewer Devonian (Locisko-viso)GEOGRAPH]CAL i)ISTRIRUTIoN: NorIS America (USA: New York)REFERENCES: HALL, 1859; CASTER lo Uniscus, [967c

Anomalocystites disparilis His¡.í., 1859[=Bassles-ocysíisdisparilis (HALl, [859)]STRAOGRAPHICAL OCCURRENCE: Lower Devenian

GEoGItAPHICAI, DISTRIBUTiON: Nortis America (USA)REFERENCES: HALL, 1859; SCHI5CHERT, l904; KIRK, 1911;CASTER, 1952

[Anomocyslisensífer(BsRRXNi)E,1887); JAEKEL, 1918]—Spermacys/is enstfer UBAGHS 1967[=Anomaiocyslilesensjfer BARRANDE, 1887]STRATIGRAPHICAL OCCURRENCE: Ordovician

GnoGRAuHrcisL DISTRIBUTION: Europe (Bohemia)REFERENCES: BARRANDE, 1887 JAEKEI,, [900, 1918; CHAU-vEL, 1941: UBAOHS, 1967c

[=Anonaalocysiitesincipiens BARRANDE, 1887]Miíroeystella incipiens (BARRANDE, 1887); JAEREL, l901REPERENCES: BARRANDE, 1887; JAEKEL, 1901

[Anoenalocystlles pyramidalis BARRANDE, [887]=Lagynocysíispyramnidalis (BARRANOE, [887); JAEKEL, 19l8 —>

REFERENCES: BARRANDE, 1887; JAFNEL, l918

Aspidocarpus bohemicus UBAGHS, 1979STRATIGRAPHICAL OCCURRENCE: Upper Ordovician (Caradoc)

GEOGRAIHICAL DISTRIBUTION: Europe (Bohemia)REFERENCES: UBAC,HS, 1979

Aspidocarpus riadanensis (CHAUVEL, 1937);LEFEBVRE, 2000

hMitrocystites riadanensis CHAUX’EL, 1937]STRATIGRAPHICAL OCCURRENCE: Upper Ordovician (Caradoc)GEOGRAPHICAL DiSTRIBUTION: Europe (France)REPERENCES: CHAUVEI., 1937, l94[; LEFE5JBRE, 2000b;LEFENRRE, 2000a

[Aspidocas-pusdiscoidalis (CREuS, 1990); LEFEBVRE 2000]=Chaar’elia discoidalis CRIPFS, [990 —~

REFERENCES: CRrí’us, 1990; LEFEvERE, 2000b

Ateleocystites huxleyi BtLLrNGS, 1858NOTE: This is the firsí carpoid described,Srr-ATicRAPHiCAL OCcURRENCE: Middle OrdovicianGEOGRAIHICAL i)iSTRiBUTION: North America (Canada)

REFERENCES: BILLINOS, 1858; CsxSTER in KOLATA & JOLLIE,

1982

Ate/eocystites guttenbergensis KOLATA & JOLLIE,1982

STRATIGRAPHICAL OCCURRENCE: Upper Ordovician (Caradoc)

GEOGRAIHICAL i)ISTRIOUTION: North Amorica (USA: Wiscon-sin)

REFERENCES: KOLATA & JOLLIE, 1982

Austnatocystis ¡angel CASTER, 1954 [1956]STRATIGRAPHICA!. OCCURRENCE: Lower Devenian (Emsian)

GEOORAI’HICAL DISTRIBUTION: South America (Brasil: Paraná)

REFERENCES: CASTER, 1954a, 1954b

[Ralanocyslitesescandei (THORAL, 1935); VIZCMNo & LEFEBV-RE, 1998]

=Anat~fopsis escandel escanclei THORAL, 1935REFERENCES: THORAL, l935a; CHAUVEL, 1941; JEFEERIES,

1987; DOMíNGUEZ & GUTIÉRREZ-MARCO, 1990; VIZCAÍNO &LEEEvBRE, 1998

Bahanocystites ¡agenula BARRANDE, [887STRATIGRAI’HICAL OCCURRENCE: Middle Ordovician (Lían-viro)

GEoCRXPHIcAL i)ISTRiOUTioN: Europe (Bohemia)

REFERENCES: BARRANDE, l887 CHAP/EL, 1941

Balanocystitesprimnus (BARRANDE, 1872); JEEFERJES,

1986[=AnatrfopsisprOna BARRANDE, 18721[=Anaílfopsisprimas BARRANDE, 1872 (incorrect gender)][=Sagitlacystisprima BARRANDE, 1872; PARSLEY, PROKDP, &DERSTLER, 2000]

Coloquios de Paltonlologia2002,53:33-68

55

[‘anidoDcsmíngaez Alonso & es al, Art annosared check-liss of gesrera and species of carpnids

Strsxríc;Rss±tCXi.OCCIJRRFNCE: Middle Ordovician (Lían-virn)GazoGaoxuhIiCXL i)ISTRIBsJTIt)N: Europe (Bohemia)

RETI-EstrzNCas: BARRANDE, 1872; JISFFERIBS, [986: PARSLETY,PaoEctv, & Dr[RSTSrza, 201)0

[Bcroleross.sti.sdisparilis (lIsta, 1859)]=Arscsorolovseiees dis

1,nc/lis Hssr., 1859RÚi’erseNCres: HA>>, 1859; SUISUCIIIIas, [904: KIRE. 1911;CAvFIz, [952

Ilarrandeocarpusjaekeli UBAGI rs 1979Sstsrsí [RAu>ItUAS, oCCI!RRENCE: Upper Ordotician (Caradoc)

Gessort~s’rs (‘NS It [Sta ¡ IIL’]soN: Europe (Bohonria)Rs ir so vr s, tíissorrS, ¡979: Rr;s.x, 1997d

Barrandeocarpus norvegicus CíQexsau & JLFFERIIúS[989

STÑXrronsr’rasAL 0CCURt<et’CL: Uppor Ordovician (AsligilOGs1oonsr’uscsr I)sSTRIBs’TstSN: hisope (Norwzsy)RerETIIETNUES (RASEL & Jrit’Fr,RIES, 1989

I3okkeve/dia ooslhuizeni RUTA & ?HERON, [997SissÁrtuasFillc¼r.s)CCIsRIENCE: Devooian (Erosian)(i>-ooR’wsilC Nt, 1)55 tRiBUlION: Southem Africa (Seulis Africa)Rt:i i Rl N(l’< Rs’ SN & T>íeÑoN. [997

Chau ve/ja discoidalis CrRíi’ps, 1991)VAsísidsssaspu.s di.sses/elalie >Crcrn’sÍ. LFUI IsN’RU 2000]SsRAIs(iRAt’StsCAU 05CL’kRtNCt:: Midd[c Ordovician (Llandei-lo>GeooRXs’HsCXL IYssTRIOL’rloN: Norxhern Africa (Morocco)Rnr:eRvxcrtS: (‘szrpr’s. 1991); l±rrivtRe,20>)t)b

Chinianocarpos thonaui UtsAC;ris. 1961.STRATsCRXuissC:xs, OCCsJRRETN(ET: Lower (ilrdovician (Trema-doc>Geousitxs’mcsí. tssSsRIETL!Tlt)N: Europe (France>

RerozRrvtzs: UrrXs;vrs, [961, 1969: JwrETRrETs, 1986

Da/e,~ocyslis casteri PROKoP. [963SraATIc;tAs’orCAs 5t( Cl tRFMCF Middls- flevoniarí (Fifelian)Gí:oustAs’síIcNs. i)ISTRSISt ‘TsCsN: Europe (Bohenria)RI:rts*LNCi.S PRoEost [963,HADe. [98(1.[983

Diamphidiocysfis drepanon KoLAIA & GUENSBURO,1979

STn~NTscnsuíssetxi. oCcISRRENCE: Upper Ordovician (Asbgil 1)Gcocasr’tsícár. 0lS3RiRUT>ON: North Amnerica (LISA: Illinois>Rr:ri:ar:Nes:s: KnrxrX & GueNsissjao. ¡979: LeFevBÑe,2000a

Enop/oara ba/anoides WETIIILRHV, 1879 [Mí:r:a,18721

[zAssesotnletsssí/ts’s <A ¡elcscsobes’) balano/des MECE, l 872

Golsseíaios ele Palessnoslesg[es 5620>[3 53 536>1

STRATIGRAPHICAL OCCURRENCE: Upper Ordovician

GeooRXr’tsrcsL i)ISsRiBUTIoN: NoríS Arnerica (USA: Ohio)REFERENCES: METER, 1872: ‘,Vtr¡reísíy [879; WOoowAÑo,[880

Enop/oura crustacea (HAECKEL. [896>; CASTER,[952

[=PlacocvstisCrasloUea IteCací., 18961PErsoplossra beilemoides WI;rHIiRBy, [879(Mssr:R, 1 872)SrrcXTsGRsFrrIoNs. OCCI;RRENCiB: Upper Ordovician

GLOGRAPHICAL DIsTRIBUTION: North Anierica (USA: lísdiasra)Ri:simtzes:es: HAETUNEL, [896:CXsreí< 1952

Enop/oura mee/ej CASíLR. 1952STRÁTsGRsFiiIcAL OCCURRENCE: Upper Ordovician

GeOorssrEssCxr, Disí Rs» isroN: Nssrth Anscrica (USA: Ohio)REETERENCES: tiAsríR. [952

[=Enoplouis-nY papi/lara [3sso FR, 1 9~31—Aoarifispsis papillata (BAssLeis, [94fo[=K/s-kescvstispapillaS’; B,xssr s~ 1 950]Rs LRuNCI.S, Bssss í~í, 1943. [950;PXasrs;v, 991

Enop/oura popei CASruiR. [952STRATII; R XIISICNi (SIN IRRCN( U tI pper (Irdovie i an (Caradoc)GI-OGRAS’¡tt(\l DISTRInIsTIoN: Noríh America (USA: Ohio)It-sr RS-NCFS CSSTvte, 1952a. 1952b: RUTA & JEte [9995

[Lrsoploosa pare tato BNsss.r:a, ¡9501—. Anaítlopsss pune eau, (B~NSSr LE. 1 950)

Ktckoe x svrs puntenea 8A5SLLR, 19501RFFFRENCFS: [3SSS’ uit >950: PAÑSsrze. [98l

E»ophura wpthprhvi CÁsrI;ra 15>52STRMieRsss’ítsUXL OC(s’RRENCe: Upper lllrdovician

GEcscRXrsírcXL i)ISTRIBusrroN: Nortín s\mcrica (USA: Indiana)REETLRENCES: CASTeR, [952

Eumitrocysle/la sari/li Biíísswr-eriusízíz, [994STRATIcR-xurssCAL OCCURRETNCET: Miádle Ordovician (Llandei-lo)GEIN;RAFr-sss, L s)isTRsBIJTION: Northern Africa (Morocco>Rrzn RENCFN, OeisswETNoEre, 1994

Guichenocarpos nioni CíIAUvEL, ¡981SrsevvscRAFuIcXs OCCtRRFNCE: Middle Ordosician (Llandeilís)GIIOGRAI’HICAL 0iSlRiIsLSTION: Europe (France)

REí ERENCCS: Ci> NLNETL, 1982

.Jaekelocarpus okhahomensis KOLATA, FREST &MArES, 1991

STRsrisas. IHICAI. OCCURI<CN(L Upper Carboísriferous (Mos-

covian)GFooaXvHiem DrSirOHUiioN: NorIS Arnerica (USA: OklaSo-rna)

RsCFvuiesCr;s: KoI.XTA e/al,, 1991: DONIiNGIIFz ev al. - 2002

Patricio Domínguez Alonso & el al, Arr arsnotated clieck-list of gorrera aríd species of carpo¡ds

Kierocysfis inserta PARSLEY, 1991STRXTIGRAPHICAL oCCURRENc:E: Upper Ordovician (Caradoc)

GEoGRAPIIICAL i)iSTRiISUTION: NorIS America (USA: Ken-tuclcy)REFERENCES: PARSLEY, ¡991; Ruis. [999a

Kopficystis kirkfie/di PARSLUY, [991STRATIGRAPHICAL OCCURRENCL: Upper Ordovician (Caradoc)GEOGRAPHII:AL i)ISTRIBUTION: Nortis Amorica (Canada: Onta-rio)

REFERENtES: PARSLEY, [991: RI;TX, 1999a

L.agynocystis pyramida/is (BARRANDE. [887); JAEREL,

[918r=Aoo,nalosvsvlles pyra,nidalis BARRANIJE, 1887]STRXTrcrsxuHrcAí. t)CCLRRENCE: Middlo Ordovician (Lían-virn)

GEOGRAPHICAL DISTRIBUTION: Europe (Bohemia, Franco,

Spain)

REFERENCES: BARRANDE, l887; JAEKEL, 1918; CHAUNEL,

1941; JEFFERIES, l973; UBX:írS, 1991: PARSLEY, 2000;LEFEN’BRET. 2000a

Mi/onicysfis kerfornei CHAUNEL, [986 [Cornuta]->NOTE: origirsrally describod as a mitrate buí [atorwas interpre-ted as a cornuto: Seo Milonitvsei.s k-er-ftsnmei CHAUvEL, 1986under Cornuta

REFERENCES: CHAUVEI,, [986 CRIvus & DALEY, 1994

MitrocystelIa barrandel JAEKEL, [901[=Pro,oies-ocys/itesbarrandel (JAIzKFI.); LEFEBVRE, 2000]SnaATrcstxpHrcsr. OCCIJRRENCE: Middle Ordevician (Llanviro)GEoGRxuarcAL i)ISTRIBIsTII)N: Europe (Bohemia)

REFERENCCS: JAEKEL, [918; (HAIsvEL, 1941; LETEEVBRE,

200115

Mitrocystetla incipiens (BARRANDE, [887); JAEKEI.,

190][=Anomalocvstitesinc/picos BARRANDE, 1887]STRXTIcass’HIC’I. OCCURRENCE: Middle Ordevician (Llandeilo)GetxsRsrr’HicssI. DISTRIBUTION: Europe (Bohemia)

REFERENc?ES: BARRANDE, 1887 JAEREI,, IQOl, 1918; UBAGIIS,

1967c; JEprERIíss, 1967, [968,986; PARsí.EY, ¡994; LEPEN-RaE, 200>ía

Mitrocyste//a incipiens miloní (BARRANDE, 1887;CHAUVEL. 1941)

STRATIGRAPHICAL OcCURRENCE: Middle Ordevician (Llandeilo)GEocsrswírIc’i, DISTRIBUTION: Europe (France, Spain)

REFERENCES: CHAIsvEL, 1941, 1982: GUrIÉRREZ MARCO &MELÉNDEZ, [987

Mitrocystites mitra BARRANDE, 1887[=Mitrocvstismi/ra BARRANDE, [887,CI;A15NEL, 1941, neo-rrect spelling]

STRATIGRAI’HICAL OCCURRENCE: Midd[e Ordovician (L¡an-vira)

GFcXRAIHICAL DISTRIBUTION: Europe (Bohemia)

REFERENCES: BARRANDE, [887; JAEKEL, 1918; CHAUvEL,1941; UIIAGHS, 1967c;JnusmRIEs, l967, 1968,1986; PARSLEY,1994

[Mitrocvstilesriadanensis CHAUNEL, [941]=Aspidcse:arpas riadanensis (CHAUNEL, 1941); LFFEBN’RE,2000->REFERENCES: CHAUNEL, 1941; LEFEVBRE, 200Gb

«Mitrocysfites» styloideus DEHM, 1934NoTe: «Mitrocysiites» s/eloideas DEHM, 1934 from tSe l-luns-riick síato (Germaoy) is a species related witS stem-grouptunicates such as Anatlfopsis or Balanocystites. It is nol asíero group craniato sucis as Miírocystella.STresrrtiresuurCAL OCCURRENCE: Lower Devorjian (Emsian)

Gnocr>AuuícuxL i)ISTRIISUTION: Europe (Geranany)

REFERENCES: DEHM, 1934; KUTSCHER, 1975; BARTELS ev al?,[998

Mongolocarpos minzhini ROZHNOV, 1990STRATiGRAPHIcAL OCCURRENCE: Silurian (Ludlow)

GEOGRAPHICAL DISTRIBUTION: Asia (Mongolia)REFERENCES: ROzONON, l990; RUTA, 1999a

Notocarpos garratti PH¡LIP, 1981STRATIGRAPHICAL OCCURRENCE: Upper Silurian (Ludlow)

GEOGRAIHICAL i)ISTRIuLTION: Australia (Victoria)

REFETRENCES: PHILIP, 198l; JEETraRIES, 1982; RUTA & JELL,

l999c

Notocarpos praecox (CHAPMAN, 1904); JELL, 1999[=Divhyrocarispraccox CHAIMAN, 1904]NOTE: Originally described as aCoastacea PhyllocaridaSTRATIGRAPHICAL OCCURRENCE: Upper Silurian

GEOGRAIHICAL i)iSTRiBUTION: Australia (Victoria)

REFERENCES: CHAIMAN, 1904; ROLFE, 1969; JELL, 1999

Occu/tocysfis koeneni HAISDE, [995STRATIGRAI’HICAL OCCURRENCE: Lower Devonian (Emsian)

GEOGRAPOICAL i)ISTRiBUTION: South Amorica (Argentina: San

Juan)REFERENCES: HAIsDE, [995; RUTA & JELL, 19995; RUTA,1999a

Ovocarpus? circularis UBAGHS, [994SrrexTíc,rssuHícsL OCCURRENCE: Lower Ordovician (Arenig)

GEOORAPHICAL DISTRIBUTION: Europe (France)

REFERENCES: IJBAGHS, l994

Ovocarpus moncereti UBAGHS, 1994STRAT]GRAPHICAL OCCURRENCE: Lewer Ordovician (Arenig)

GEoGRAPIIICAL i)isTRmurrIoN: Europe (France)

REFERENCES: UBAGHS, 1994


57

Pcstrisio Dcsnsíngrtez A/es,rso & 05 al. Aa annrotased check-¡ist of gorrera mrd species of carpoids

Paranacysfis petrii CASTER, [954STRATIGRAPHICAI. OccísRnzNcr: Lower Devenían

GEOGRAFIOCAL i)SSTRIOUTII)N: South Amorica (Brasil: Paraná)

REFERENc:ETS: CASTER. 1 954a, 19Mb: CASTFR & FATON, 1956;UDACIIS. i967c: Ríns, [997a

Paranacystis simoneae Ríínx, 1997STRXTir;RsuIICAL OCCURRENCE: Lower Dovonian (Pragian)

Geoc;RXuhricsL i)15rRiBLTi(sN: Seutisero Africa (Republic of

South Africa)

REFERENCES: RusTA, ¡997a

Pe/tocystis cornata THORAL, [935SsÑXIIoRAFHJeXt, sXTURReNCtL: Lewer Ordoviciarí (Trenadoc)

GeoGrsXr’s 1 ¡CAL DISIRIisítIION: Europe (France)

REFEReNCES: TísoroAr, 1935b; Uwwsís. [969: JEFFERIES,

¡986; PARSEEN & PaoKou, ¡999

P/acocystel/a africana (REED, 1925); DuiííM, 19321 =Plas:ocvsíis nfrlsana RFF1), 1 925[=Plasosv.stella s ape’rs/s RFNMF, 1936]STaX]souXFISsCXs, ocCIsRRENt:E: Lower Devorrian Eíwian)

GETx;uXFIsmcXL DISTRIBUTION: Southern Africa (Republic of

South Africa)RvFvur;NcES: DEHM, [932; RUTA & IHERON. ¡997; RISTsN,1999a

rtlacoc}síella Capcnsi.s RENNIE, ¡936]= Placocvslello aflisana (Rrzni), 1925); Dursx4, [932STRsTrc,RXuííwXL oCcIJRRETNCE: Lower Devonian (Ensian>

GEoaÑAPHICsi. i)sSTRIBL’TII5N: Southern Africa (Republie of

South Africa)

Reí’uiarsNc:tes: RusENte, 1936; Rs:rs & Tr-reieoN, 1997

[Placoeyseisho/remitas (BARRANDF, [887)1j=Anomnalo¿:vsl/tes lsohenric:as BARRANDE. [887REsERENcEs: BARRANDE, 1887: CRsssvEs, 1941

[Píacocvs//.scraslacea FIAeCKES,, 1896][=Lnoplous-acnístercs’a (HAFURFI , 1896); C,XSTER. 1952]Rrsrínsr-NcFs: HAPCKI-:L, ¡896; CXssEíe, 1952

Placocystilesforbesianus ErE KoNINcK, [869SrroXTIcRXurrrc:XL t5CCsJRRENC1): Silurian (Wonlock)

GuiooRXi’r-rieXL DiSTRIBIsTION: Europe (UK: England)Reí’earzNces: CASTER, [952:CF5AP/EL, 1971; JEFETERIES &Lcwis, 1978: ifETER/reS & Lewrs, 1978: JrsuvERIr¿s, [984;SANAZzI ee al,. [982;Koreers & Jorire. ¡982; Rois, 1998;Rtjís & BArvíeLs, 1998

IPs-o;nitrocysviees barrandel (JAERFs., [901);LEFETWVRE, 2000J=Mieroevseclleí bar,aodc/ JAEREL, l90l-~RríFarNcrs: J’~rr<ri,, ¡918; LEETENBRIS, 201>1>5

Protocylidium e/liottae RUTA & JEEL, 1999SraXncRAFriíeXL OCCURRENCF: Upper Ordovician (Ashgill)GvouRsurncxI i)ISTRIEI;TiON: Australia

RisIriRUNCr-;s: Rí rs & Jvr .s , 19995

Co/oqs¿iess de Paleesnrolos>/e,

Pseudovictoriaeysfis problema¡¡ca RUTA & JELL,1999

SsroArrorsXr’nícXr. OCCItRFtETNCET: Lower Devonian (Lochko-

vEro)

GETOGRAPHICAL i)ISTRIBI;TioN: Australia

RFFETRFNCES: RUTA & JFLL, 1999d

khenocysfis ¡afipedunculata DEHM, 1932STRATIGRAPHICAL OCCsJRRENCI>: Lower Devenian (Erosian)

GsscscroNussrICst, i)ISTRIi)t;TION: Europe (Gennany)

REFERENCES: Derrra, 1932, 1934; KUTSCHER, [975; ¡lAUDE,

[983:KorATA & J(5J.IIET, ¡982; RISTA & BARTELS, l99& BAR-

TELS el al., 1998: SLsTCI rvrr el al?, 2000

Spermacystis ensifer (BARRANDE, 1887)[=Aoonsocysíisenstjér (BARRANDE, 1887)][=A,romalocysílees ensifer BARRANDE, 1887]STRATIORAPFICAL OCCURRENCE: Middle OrdovicianGEOGRAPHICAL i)ISTRII5IsTION: Europe (Bohemia)

REETERENCES: BAR/ANDE, [887; JAFKEL, [900; JAEREL, ¡918;Crrss:vnt. 1941; UnXc,us, 1967c

Tasmanicyfidium burretil CASTER, 1983STRATIGRAPIIicAL (sCCusRRENCE: Lower Silurian (L[andoveiy)

GEOGRsur¿Icsr isrsrRinUrroN: Australia (lasmania)ReecREroCes: CASTER, [983; RUTA & JELI., 1999c

Victoriacystis holmensorum RUTA & JELL, 1999STRAT5CRAPHICAL (sCCusRRFNCF: Lower Devonian (Lochko-vían)

GIsoURAPHICAi, DISTRIBUTION: Australia (Victoria)

REFERENTES: RUTA & Jnusr, 1999d

Victoriacysfis wilkinsi GILL & CASTER, 1960Noii: seo note for uSe solute Ratroclypeas cictos-iae GILL &CASTER, 1960STRATICRAPIIICAL oc:CURRETNCET: Silurian (Ludlew)

GETOCRAIHICAL i)ISTRiBUTION: Australia (Victoria)

RETFETRFNCF5: GILL & CASrere, 1960; RusiA, ¡997c

Vizcainocarpus denfiger RnA, 1997STÑATIcsRss’HSCXL t)CCURRENc:E: Lower Ordovician (Arenig)

GEoCRwííIesi, i)lSTRiIILSTION: Europe (Africa)

REPERENCES: RUTA, 1997b; LEFENuRE. 2000b

Vizcainocarpus rutai LEFEBVRE, 2000STRXTICRsurIIc:AI. OCCURRENCE: Lowor Ordovician (Trema-

doc)GEOc,RAPHICAL DISTRI}IISTION: Europe (France)REFERENCETS: LEFENBRE, 20005

Willtnanocysfis denfieu/ata KousATsx & JOLLEE, [982STRTroRXuíírcsL OCCISRRETNCET: Upper Ordovician (Caradee)

GCocRÁurIICst, i)ISTRIBUTION: NorIS America (USA)RnrvreENcns: KOLATA & JOIlIE, [982

21102, 53: 33-6858

Patricio Domínguez Alonso & el al, An arrnrolated check-liat of genera mrd species of carpoida

Yachalicysfis triangularis HAUDE, 1995STRATIORAI’HICAL OCCURPENCE: Lower Devonian (Emsian)

GEOCRAIHICAL DISTRiIIUTiON: South America (Argentina: SanJuan)

REFERENCES: HAUDE, [995; RUTA, 1 999a; LEFEN’BRS, 2000a

III: INDEX OF \‘ALID SPECIFIC NAMES

acheronticus: Adoketocarpusacrofera: Sucocystisacuta: Anaíifopsisafricana: Placocystellaamericana: Nevadaecystisancora: Anatifopsisarchaeocyathoides: Jugoszovia

armoncana: Anatífopsisaustra/is: Drepanocarposbadulesiensls: Gyrocystisbalanoides: Enoploarabaiclatchiensis: Anal ifopsis

barrandel: Dendrocystites, Ellipíocinctus, Mitrocys-tel/a, Anatífopsis

batheri: Girvanicystisbífidus: Archaeocot/turnusblayaci: Phyllocyslisbohentica/us: Anatifopsis,

chocysti/es, Aspidocarpusbouceki: Bohemiaecystisbretoni: Sucocys/isburretil: Tasnumnicytidiumcasterí: Girvanicystis, Da/ejocystiscircularis: Ot’ocarpus?cocaban: Anat~fopsiscollapsa: Scotiaecystiscolodon: Cfenocystiscornutajus: Peltocystis, Anomalocystitescourtesso/ei: Coíhurnocysíis, Lyricocarpuscrassimarginata: Playllocystiscross,naní: Myeinocystitescrustacea: Enopleuracruzae: Gyrocystiscurvata: Scotiaecystisdelficus: Acuticarpusdenficulata: Wil/manocysíisdenfiger: Vizcainocarpusdilabida: Babinocvstisdiscolda/is: Chauveliadisjuncta: Progyrocystisdisparilis: Anonealocystites

dolambii: Nanocarpus

Etoctenocystis, Tro-

drepanon: Dia¡nphidiocystiselizae: Cothurnocystiselliottae: Protocytidiumensifer: Spermacystiserecta: Gyrocystisescandei: Analrfopsisestonlea: Maenniliafellinensis: Cothurnocystisfiemingi: Allanicytidiutnforbesianus: P/acocystitesgarrati: Notocarposglobulus: Dehmicystisgoshutensis: Archaeocothurnusgnffei: Thoralicystis, Amygdalothecaguichenensis: Anatifopsisgui/buí: Bohemiaecystisguttenbergensis: Aíeleocystheshav/iceki: Asturicystishanusi: Reticulocarposhispanlea: Gyrocystisholmensoruin: Victoriacystishuxleyi: Ateleocystitesincipiens: Mitrocystellainserta: Kierocystisjaeketi: Asturicystis, Barrandeocarpusjagol: Cíenocystisjaneae: Adoketocarpusjeiferiesí: Bohemiaecystisjunorí: Rutroclypeus/eerfornei: Milonicystiskirkfieldi: Kopficystiskoeneni: Occultocystiskuckersiana: Heckericystisbagenula: Balanocystitesbandeyranensis: Atupelocarpus/angel: Australocystislafipeduncubala: Rhenocystislignieresí: Galliaecystis/onga: Anatifopsismaroccana: Sucocysíis?mee/el: Enoplourameichiorí: Proscotiaecystis, Sucocystismenevensis: Protocystitesmergil: Pro/eopicystismIloní: Mitrocystella

ml/nerorum: Juliaecarpusminuta:Anatr¿fopsisminzhini: Mongolocarpos

miranda: Berylliamitro: Mitrocystites

moncerefi: Ovocarpus


Pssesieio /Js»rssrigmsez .Aissrsso & es al. Arr arísrosared checklism of gesrera ssmm species of carpoids

natus: Mveiííocvsíiíc<s

nioni: Ca/ch enocal-pos

narr’egicas: f3arrandeoearpusobtusa: fiantísíít

occitana: A¡-cta ricv.stisoklabomensis: laekelocwpu.s

oosthreizeni: Be4ke ucidia

owensi: P,ocotlitunocvslispapillote: Anaítfapsí’

paradaxicas Svs-rnqac 1-inrisparvas: ha> bac x statdcspembro/eensís Dar íclocinc:iu,s

perneri: Cc la/ocx suspetrii: Pc>ranca xstts

písseuiensis: Damjrantiaplanas: Trac/roevsíoides

plale.ssa: CvroCNwlis

papel: Eoaploaaapraecox: Naloca ¡posprilepensis: linaasic>priínac’va: A¡atíricvsrís

primas: Ralcrnacvsíiíesproblc’matico: Psc>udos’,c:Ia s-¡ac:vst,spanclata: Anatifopsis

pyramida/is: f.ctgx’nacv.síis

quadricornata: Sacar vslisrepablicensis: Acuíicarpus?riadonensis: A spídocarpcts

rossícus.~‘c Deníhacvseiícss’rushtoni: [lOIé’lliUv.Stisratal: i’i~ctutacfí rpussalairica: Phxllocvstissagillaria: Jara w Vslis

sar/eensis: 1-lanasia

savi//i: Eumierocvsrella

scaficas: Dendracx’síiíessedgsvicki: Dendrocysrilc’ssemispinosa: Prochaavclicvsíissunonene: Paranacvsiissincíairi: Svringacrinnssinga/arls: Trigonar amassmilhi: É’tenacxsrspinoso: C/taas’c’hc s si>s Anat¡fápsissprink/ei: Calcitas-pussírimplel: Scalencsc y sítírsty/oideus: «Mili-ocx ‘Uve s»testudiformis: Gvrcícv sí> s

¡heranensis: Sucacvslis(horail: Ch brictuacarpos, Anattilapsis

trapezilformis: AnW/fcq’sisIraveusotensis: Aantij4osi.s

triangularis: Yac /talicysíis

trancatumius: Aaatrjfrpsis. Lcídwigiciactusabaghsi: Cima velicv.seis, Praga iíieczsei.s, T/toralicvs-

lis Solocinclasuní/ala: Sucocs’se/satahensis: Cteaocvstts

vail: Caslceicvstisvictorice: Ra ls-oc!zpcas

vida/i: Aiinerusaecvsvís

vizcainol: (/taauselicvslis, E/liptc>tincti~s, Ceratocvs—

lis, Laboí~a rpta.swetherbyi: /)clcmnacysíiíes, Enaploaraivilkinsi: Víctoriacvsevs

wilbersi: Rutrac/”pcas

zagoraensis: 7haraiicvs¡is

60Ceslese¡aios de Pss/s,s,sses/oy’/es

2<»>?. 5? 33-6>1

Patricio Dom/ngucz Alonso & el al.

IV. TABLES OF STRATICRAPHICAL OCURRENCE

Aa aranotated check-list of genero mrd apecies of carpoidí

Otn,bdío O,doeklnn SIsarían bevonlan Caetonlt.rous

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Coloquios de Paleontología2002, 53: 33-6[

61

Pcíííis/o Dssntíísgsacz Alonso & es al, An annotamed ctseck-Iist of genero and species of carpoids

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Coloquios eíc Palcesntoiogs/a

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Patricio Domínguez Alonso & cl al. An anorolated chcck-[ist of gencra and species of carpoids

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ACKNOWLEDGEMENTS

We tSank referee E. MoRENo, an anenymousreferee and P. SEVILLA, Directora of Coloquios dePaleontología, for useful suggestions to improve Ihetexí.

R DOMÍNGUEZ was supperted by [he SpanishGovernment (Ministerio de Educación Cultura yDeportes) through tSe Programa Nacional de For-mación de Personal Investigador, Becas en elExtranjero y, excepcionalmente, en España,

Secretaría de Estado de Edacación y Universida-des.

63

Recibido el día 15 de marzo de 2002Aceptado cl día 9 de septiembre de 2002

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Rec/rerc/res paléontologiques. Supp. Vol. 1. Trilobites, Crusta-ces divers et Poisins, 647 págs. Praga-Leipzig.1887. Sysítroe 8/luden du centre de la Bo/rhne. Vol. VIL Classe

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Patcis‘lis Dsss=sírss4sac;Alo>i=s=& es al, An arírotared check-iist of gcncra mrd speciesi of carpoid.s

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