A Guide to the Adult Damselflies

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    A Guide to the Adult Damselflies & Dragonflies of the OttawaDistrict

    List of Species and their Preferred Habitats

    There are many words used in English to describe bodies of water. The meanings of many overlapconsiderably in common usage. I use the following definitions in this guide, which concentrate on thefeatures important to dragonflies:

    water volume mostly free of vegetation:

    o ditch: narrow (less than 5 meters wide), shallow (less than a meter deep), long, no visible current (includes man-

    made, and beaver/muskrat trails)

    o stream: a ditch with perceptible to moderate current, but no white water

    o river: a stream more than 5 m wide, much more than a meter deep, some current

    o rapids: white water (at least during spring runoff)

    o outlet: where a stream empties into a lake

    o puddle: shallow (much less than a meter), small ( a few meters or less across), no current

    o pond: moderate depth (1-5 m), moderate size (1 ha or less), no current

    o pool: a deep portion of a stream, with some eddy current

    o lake: large (greater than 1 ha), at least some deep portions (more than 5 m), no current

    o large: (lake or river) large enough to have breaking waves and wind-driven current

    o calcareous: limestone region, pH >7.5

    water heavily vegetated:

    o bog: most water volume filled with plant material such as sphagnum, pH less than 6

    o marsh: heavily-vegetated (typically cattails) pond or lake edge, pH 6 or greater

    Species noted as southern(S) can be expected only in the warmer portions of the district, northern(N) onlyin cold bogs. Species that are exceptionally local(L) in distribution or rare(R) here are noted as well.Excepting these species, habitats mentioned are those in which the species is most likely to be found here,and are not to be treated as the only places to look. Species in brackets are known only from close to thedistrict, not from within it.

    Zygoptera

    Calopteryx maculata small forest streams

    Calopteryx aequabile larger wooded streamsL Lestes eurinus small lakes, ponds

    Lestes congener temporary/permanent ponds

    S Lestes inaequalis marshy ponds, slow streams

    Lestes unguiculatus temporary ponds, overwinter as eggs

    Lestes vigilax marsh/bog-margined lakes

    S Lestes rectangularis partly shaded ponds

    Lestes dryas temporary/permanent ponds

    Lestes forcipatus temporary/permanent ponds, weedy streams

    Lestes disjunctus ponds, weedy streams

    L Amphiagrion saucium small, spring-fed bogs at hill bases

    Chromagrion conditum shady spring-fed brooks, pools

    Nehalinnia irene still marshes, bogs

    L Nehalinnia gracilis sphagnum bogs

    Argia moesta rocky shores or streams, lakes

    Argia fumipennis small shallow rocky streams

    Ischnura verticalis any still water

    Ischnura posita shaded spring-fed streams, ponds

    Coenagrion resolutum still shaded water

    (RL Coenagrion interrogatum quaking bogs)

    Enallagma ebrium limestone marshy ponds, streams

    N Enallagma boreale cold bogs

    Enallagma hageni acid marshy/boggy ponds

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    N Enallagma cyathigerum very cold bogs

    Enallagma carunculatum lakes, rivers

    S Enallagma civile ponds, slow streams

    S Enallagma geminatum small lakes, slow streams

    Enallagma exsulans streams, lake shores

    R Enallagma antennatum clayey quiet streams, small lakes

    S Enallagma vesperum marsh-bordered lakes, slow streams

    Enallagma signatum slow streams, quiet bays

    Anisoptera:Anax junius migratory, still water with emergent vegetation, esp.sand pits

    Basiaeschna janata small woodland streams with gentle current and their outlets

    R Nasiaeschna pentacantha slow woodland streams, small lakes

    S Epiaeschna heros shady ponds at wood edges

    Boyeria grafiana fly until well after dusk, rock- margined lakes and streams

    Boyeria vinosa fly until well after dusk, shady streams with gentle rapids

    Aeshna umbrosa shady wooded ditches and streams

    Aeshna constricta open marshes and ponds with emergent vegetation

    (N Aeshna sitchensis puddles in almost filled cold bogs)

    Aeshna interrupta shallow marshy or boggy lakes

    S Aeshna clepsydra shallow marsh-bordered lakes

    Aeshna eremita marsh-bordered lakes, sand pit ponds(N Aeshna subarctica puddles in cold sphagnum bogs)

    (N Aeshna juncea peaty puddles and ditches, shallow lakes)

    Aeshna tuberculifera ponds, especially in bogs

    Aeshna canadensis still marshes on lake edges, slow streams

    LS Aeshna verticalis spring ponds, marsh-bordered lakes

    Hagenius brevistylus slow rocky forest streams and their outlets

    N Stylogomphus albistylus small rapid gravel streams

    Dromogomphus spinosus rapid streams, rivers, lakes

    N Ophiogomphus colubrinus rapid gravel streams

    L Ophiogomphus anomalus wide rivers, rapid streams

    Ophiogomphus rupinsulensis streams and rivers

    Ophiogomphus aspersus cold clear rapid streams

    RN Ophiogomphus mainensis clear running streams

    R Gomphus lividus sheltered inlets on wavy lakes, gentle streams

    Gomphus exilis quiet soft-bottomed streams and lake borders

    Gomphus spicatus still marshy waters

    S Arigomphus furcifer marshy ponds and lakes

    SL Arigomphus cornutus small marshy lakes

    L Gomphus descriptus rapid streams with pools

    R Gomphus borealis lakes, slow streams, sand pit ponds

    Gomphus fraternus turbulent water, calcareous

    R Gomphus ventricosus rivers

    Gomphus vastus large rivers, lakesGomphus adelphus clear running streams, exposed lake shores

    R Stylurus amnicola rapid streams, wide rivers

    N Stylurus scudderi cold gravelly forest streams and rivers with rapids

    Stylurus notatus wide rivers

    L Stylurus spiniceps wide rivers

    L Cordulegaster obliqua tiny shaded rapid streams in deciduous woods

    Cordulegaster diastatops streams with rapids in spring, pools in clearings

    Cordulegaster maculata rapid streams in forests

    Didymops transversa slow forest streams, wavy lakes

    Macromia illinoiensis fast streams, exposed wavy lakes and wide rivers

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    L Nannothemis bella puddles and ponds in floating sphagnum bogs

    (S Pachydiplax longipennis lakes, ponds, slow streams)

    RS Pantala hymenea small standing water esp.sand pits

    S Pantala flavescens small standing water esp.sand pits, migrant

    Erythemis simplicicollis clear ponds, lake borders

    S Celithemis eponina ponds, lakes and slow streams

    LS Celithemis elisa marshy lakes and ponds

    Libellula luctuosa ponds, small lakes and marshes

    Libellula julia bog ponds to swampy lakes

    Libellula lydia puddles to ponds

    Libellula pulchella sunny ponds, lake borders, calcareous

    Libellula quadrimaculata still water

    Libellula incesta ponds, slow streams

    Leucorrhinia glacialis bog lakes and marshes

    (N Leucorrhinia patricia puddles in peat bogs)

    Leucorrhinia hudsonica puddles in cold marshes and bogs

    N Leucorrhinia frigida floating bog lakes, marshy ponds

    Leucorrhinia proxima still marshes

    Leucorrhinia intacta slow streams, marshy bays, farm ponds

    Sympetrum internum slow shady streams, ponds

    R Sympetrum rubicundulum still water

    Sympetrum obtrusum tiny, even temporary, ponds in wet meadows and gravel pits

    R Sympetrum semicinctum spring-fed ponds, marshes

    Sympetrum danae peaty marshes, bog ponds

    Sympetrum vicinum slow streams, permanent ponds

    Sympetrum costiferum sand or gravel pit ponds

    Neurocordulia yamaskanensis wave-washed rocky shores of rivers and lakes, rapids, flies only briefly after dusk

    RN Williamsonia fletcheri puddles in sphagnum bogs

    Epicordulia princeps quiet small lakes, rivers

    Helocordulia uhleri edges of rapid streams and their outletsDorocordulia libera bog ponds, sluggish marshy streams

    Cordulia shurtleffi quiet marshes, boggy ponds

    Tetragoneuria spinigera marshy non-stagnant borders of lakes and streams

    Tetragoneuria canis acid ponds and streams

    Tetragoneuria cynosura marsh-bordered lakes and slow streams

    Somatochlora tenebrosa tiny shaded deciduous forest streams

    (N Somatochlora cingulata lakes)

    N Somatochlora albicincta boggy places with slight water movement

    N Somatochlora franklini puddles in spring-fed bogs

    Somatochlora minor clear slow forest streams

    Somatochlora walshi small ditches in bogs

    Somatochlora forcipata tiny boggy spring-fed streams, puddles in bogs

    (N Somatochlora elongata forest streams with rapids and their outlets)

    Somatochlora williamsoni quiet shady forest streams and their outlets

    N Somatochlora kennedyi cold bog ponds and puddles

    (N Somatochlora incurvata sphagnum ponds and puddles)

    Identification Drawings

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    Side view(Note that no single dragonfly has all the features as shown. They were selected for

    clarity from several genera.)

    Front wing (Libelluloidea)

    Hind wing

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    A Guide to the Adult Damselflies & Dragonflies of the OttawaDistrict

    Key to Species

    Most of this key is reduced from published works that include many more species than occur in the OttawaDistrict, primarily from E.M.Walker, 'The Odonata of Canada and Alaska'. I also consulted in detail

    J.G.Needham & M.J.Westfall, 'A Manual of the Dragonflies of North America'; A.Robert, 'Les libellules du

    Qubec'; and R.Hutchinson & A.Larochelle, 'Manuel d'identification des Libellules du Qubec'. In thereduction, I favoured terms that are visible on a living insect. Each term of the key was verified using allapplicable specimens in the Canadian National Collection of Insects (Ontario & Quebec, excluding the farnorth). The field-mark oriented sections marked by symbols are my own, are based solely upon livinginsects within the Ottawa District, and appear only in cases where I am certain that they are significantlymore reliable in the field here than the specimen-based key.1 fore and hind wings similar; eyes separated by more than their own width Zygoptera 2

    hind wing broader than forewing Anisoptera 11

    2 Zygoptera: damselfiles

    wings not stalked at base, many crossveins before nodus Calopteryx 3

    wings stalked at base, two crossveins before nodus 2a

    2a wings held spread open when resting; vein M3 rises from M1-2 closer to arculus than to nodus Lestes 4

    wings held together above back when resting; M3 rises behind nodus Coenagrionidae 5

    3 Calopteryx: unstalked broad wings held together above back when resting

    male: wing all clear, female: wing diffuse colour to beyond nodus, end clear; 3Jun-2Sep C.aequabile

    male: wing all dark, female: wing all smoky with white spot at stigma; 27May-10Sep C.maculata

    4 Lestes male

    lower appendages much less than half the length of upper; thorax dark metallic green, wings

    yellowish 27May-28Jun

    Lestes eurinus

    lower appendages half or a bit less the length of upper; thorax nearly black 9Aug-13Oct L.congener

    lower appendages longer than upper, ends curved in; metallic green 30May-8Aug L.inaequalis

    lower appendages S-curved with divergent ends; blue eyes yellow lower lip black thorax 8Jun-9Sep L.unguiculatus

    upper appendages meeting at acute angle; dark metallic green 16Jun-27Aug L.vigilax

    abdomen very long and slender; hairy green lower lip, 15Jun- 10Sep L.rectangularis

    thorax metallic green above yellow sides; stout, 30May-25Aug L.dryassegment 2 3/5 as long as 3, rear tooth of upper appendages blunt, much smaller than front one;

    blackish, greenish face 6Jun- 21SepL.forcipatus

    segment 2 1/2 as long as 3, rear tooth of upper appendages acute, as large as front one; blackish,

    greenish face 2Jul-21Sep

    L.disjunctus

    4a Lestes: female

    ovipositor 1/2 or less as long as segment 7 4b

    ovipositor as long as segment 7 4c

    wing yellowish L.eurinus

    thorax dark brown, narrow pale yellow stripe L.congener

    dark areas of abdomen bronzy colour L.disjunctus

    these areas greenish L.unguiculatus

    4b grey humeral stripe more than 1/2 as wide as dark brown stripe above; fewer than 14 postnodalcrossveins

    L.rectangularis

    reddish-brown humeral stripe 1/3 or more thickness of metallic stripe above L.vigilax

    this stripe yellowish, less than 1/4 thickness of stripe above L.inaequalis

    4c thorax metallic green, stripe clear yellow L.dryas

    thorax dull bronze-brown, stripe pale yellow L.forcipatus

    5 Coenagrionidae:

    leg hairs twice as long as the space between them Argia 6

    leg hairs only a bit longer than the space between them 5a

    5a two spots on top back of head next to eye 5b

    no such spots 5d

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    5b M2 arises near the 4th crossvein after the nodus in forewing, between 2nd & 3rd vein in hindwing Ischnura 7

    M2 arises near the 5th crossvein after the nodus in forewing, near the 4th vein in hindwing 5c

    5c male: lower appendages with notch when viewed from side, female: segment 8 without spine on

    bottom

    Coenagrion 8

    male: lower appendages triangular when viewed from side, female: segment 8 with spine on bottom

    rear

    Enallagma 9

    5d body only red and black; small 1Jun-8Aug Amphiagrion saucium

    body blue & black with bright sulphur yellow stripe on thorax rear; 7Jun-4Aug Chromagrion conditum

    thorax metallic green top blue bottom, abdomen blue (male) blue-green (female) with black areas

    all the way to segment 10; 1Jun-28Aug

    Nehalinnia irene

    as N.irene but no black on segments 9 or 10; 30May-12Aug N.gracilis

    6 Argia: argos = bright (coloured)

    upper parts of segments 3-6 black-brown; 7Jun-12Sep Argia moesta

    violet-blue; 19Jun-19Aug A.fumipennis

    7 Ischnura: ischnos = thin, oura = tail

    two narrow long shoulder stripes; 5May-15Oct Ischnura verticalis

    two exclamation marks on shoulder; 4Jun-9Sep I.posita

    8 Coenagrion: agrios = field

    two long shoulder stripes; 12Jun-29Jul Coenagrion resolutum

    two wide exclamation marks on shoulder; (Muskoka 3Jun-25Jul) C.interrogatum

    illustrations for 1-89 Enallagma male: enallex = crosswise (striped)

    black of segments 3-4 less than 2/3 length of segment; pale blue 9a

    this black more than 4/5 length of segment 9c

    9a upper appendages longer than lower, a pale bump on upper end 9b

    upper appendages clearly split in two; 30May-24Aug Enallagma ebrium

    upper appendages short and rounded; 16May-24Jul E.boreale

    upper appendages slender with points curved up; 30May-21Aug E.hageni

    upper appendages with prominent tooth/ridge; 29May-27Jun E.cyathigerum

    9b upper appendages symmetrical viewed from side; 18May-20Sep E.carunculatum

    upper appendages asymmetrical; 6Jun-12Sep E.civile

    9c upper appendages tapered to slender upturned points; 15Jun-3Sep E.geminatum

    upper appendages split in profile 9dotherwise 9e

    9d upper branch of upper appendages shorter than lower branch; 7Jun-19Sep E.exsulans

    both these branches about equal; 9Jun-14Aug E.antennatum

    9e upper appendages' height greater than 1/2 their length; 18Jun-29Aug E.vesperum

    otherwise; 3Jun-9Sep E.signatum

    10 Enallagma female (these are reliably separable only by microscopic examination of thorax plate)

    segment 8 with pair of large blue spots E.geminatum

    segments 8-10 entirely black above E.hageni or E.ebrium

    segments 7,8,9 with similar marking E.civile or E.carunculatum

    segment 8 entirely black, 9 or 10 not; eye spots elongated to or almost to occiput 10b

    segment 8 black only on rear half or all pale 10a

    10a fine line between spots of segment 8 E.boreale

    these spots separated by a line at front, a wider gap at rear, side spots reach top of segment 10 E.cyathigerum

    10b black humeral stripe twice or more width of pale stripe above 10c

    black stripe about same width or narrower 10d

    10c black stripe divided by pale line E.exsulans

    not so divided E.antennatum

    10d pale stripe with gap in middle; M2 begins between crossveins 4- 5 E.vesperum

    no such gap; M2 begins between crossveins 5-6 E.signatum

    11 Anisoptera:

    eyes separated by about half their width Gomphidae 17

    eyes touching on top of head or separated by only a tiny fraction of their width 11a

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    11a triangles about equally distant from arculus in both wings, both oriented lengthwise on wing Aeshnoidea 12

    triangle at least twice as far from arculus in forewing as in hindwing, forewing triangle orientedacross wing, hindwing lengthwise

    Libelluloidea 22

    illustrations for 9-11

    12 Aeshnoidea:

    eyes barely touch at top or slightly separated Cordulegaster 21

    eyes meet in seam at least height of occiput Aeshnidae 13

    13 Aeshnidae:

    medial space with more than 1 crossvein; 2 rounded pale spots on thorax sides, yellow stigma Boyeria 14

    with one crossvein or none 13a

    13a sectors of arculus arising from its upper end, thorax uniform green; 21Apr-15Oct Anax junius

    sectors of arculus arising near its middle, thorax brown 13b

    13b Rs vein not forked; 3 or more cubito-anal cross veins, blue body with bright yellow strips on thorax

    sides 8Jun-17Jul

    Basiaeschna janata

    stalk of Rs vein with symmetrical fork 13c

    stalk of Rs vein up-arched and fork askew forward Aeshna 15

    13c radial planate subtends 1 row of cells; very prominent blue frons, face yellowish margined with

    brown 20Jun-16Jul

    Nasiaeschna pentacantha

    subtends more than 1 row at least basal half; largest Odonata here, brown & green, 7Jun-9Sep Epiaeschna heros

    14 Boyeria: fly until well after dusk

    wing base completely clear; side spots of thorax grey-bluish, segments 9 & 10 similar in colour,body grey-brown 27Jun- 7Oct Boyeria grafiana

    wings with brown spots at base (usually small); 2 clear yellowish thorax side spots, segment 10 more

    yellow than 9, body brown, 5Jul-22Sep

    B.vinosa

    15 Aeshna, male:

    anal triangle with 3 cells; 2 yellow to green stripes on thorax 15a

    with 2 cells 15b

    15a rear of head marked with yellow or brown, row of pale spots each side of segments 4-6; some green

    on segments 5-10, 12Jun-21Oct

    Aeshna umbrosa

    rear of head black, no such spots; 19Jul-11Oct A.constricta

    15b central thorax stripe 1 mm or less wide and bent in sharp Z; eye seam little if any longer than

    occiput, greenish face, T-spot with crescent base (Tremblant 21Jun-14Sep)

    A.sitchensis

    central stripe not bent sharply, more than 1 mm wide; eye seam at least 2x length of occiput 15c

    15c central and lower thorax stripes interrupted, reduced to two elongate spots or absent; yellow head,

    dark body 2Jul-1Oct

    A.interrupta

    these stripes wider, generally full length 15d

    15d face with fine but clear black-grey stripe on face suture 15e

    stripe not clear, more like a shadow than a marking; upper side of upper appendages hairy 15f

    15e large pale spots between green side stripes of thorax, sometimes confluent with central stripe; 5

    paranal cells, slender 19Jul-12Sep

    A.clepsydra

    spotted with blue; 4-5 paranal cells 19Jun-15Sep A.eremita

    central thorax stripe clearly concave on front edge; 3-4 paranal cells 12Jul-10Sep A.subarctica

    central thorax stripe convex on leading edge (may be only slightly); 3-4 paranal cells 14Jul-31Aug A.juncea

    15f segment 10 all black; large thorax stripes 27Jul-22Sep A.tuberculifera

    pale spots on dorsum of segment 10 15g

    15g central thorax stripe mostly or all blue; 21Jun-27Sep A.canadensis

    this stripe all green; 14Jul-11Aug A.verticalis

    16 female:

    central thorax stripe 1 mm or less wide and bent in sharp Z; eye seam little if any longer than

    occiput, greenish face, T-spot with crescent base

    A.sitchensis

    central stripe not bent sharply, more than 1 mm wide; eye seam at least 2x length of occiput 16a

    16a palp of ovipositor (see 4a) as long as segment 10 16b

    palp much shorter than segment 10 16c

    16b front edge of central thorax stripe nearly straight; thin pencil of hairs at end of vulva A.tuberculifera

    this edge distinctly sinuate; no such pencil of hairs A.constricta

    16c face with black line on face suture 16d

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    any such line indistinct 16f

    16d central and rear thorax stripes interrupted, reduced to two elongate spots or absent A.interrupta

    these stripes well developed 16e

    16e front thorax suture with large triangular pale spot below A.clepsydra

    4-5 paranal cells; spotted with green 19Jun-15Sep A.eremita

    central thorax stripe concave on front edge; 3-4 paranal cells A.subarctica

    central thorax stripe convex on leading edge; 3-4 paranal cells A.juncea

    16f central thorax stripe straight, margined with blackish A.umbrosa

    central stripe distinctly sinuate 16g16g central thorax stripe mostly or all blue A.canadensis

    this stripe all green A.verticalis

    illustrations for 12-16

    17 Gomphidae:

    triangles with a cross vein, each with a supplementary longitudinal vein springing from its outeredge; very large, small head, black and greenish yellow, eats other dragonflies 21Jun-21Aug

    Hagenius brevistylus

    triangles clear, no such vein, rarely 65 mm long 17a

    17a male: hind wings with anal loop, usually 3 cells, lower appendage ends separated slightly, female:

    vulva nearly as long as the rear of segment 9

    Ophiogomphus 18

    male: without anal loop, lower appendage ends widely separated, generally divergent, female: vulva

    usually less than half as long as the rear of segment 9

    17b

    17b small, 25-40 mm long, stigma less than 4x as long as wide and more than 2x as wide as the spacebehind its middle; greenish with lots of black 21Jun-14Aug Stylogomphus albistylus

    42-65 mm long, stigma rarely less than 4x as long as wide, less than 2x as wide as this space 17c

    17c long hind legs reaching the base of segment 3 with 4-7 long spines below; dark brown, distinctive

    pale green markings on thorax 5Jun-31Aug

    Dromogomphus spinosus

    hind legs not reaching beyond the middle of segment 2, only short spines Gomphus 20

    18 Ophiogomphus: male

    face striped across with black 18a

    face uniformly pale 18b

    18a upper appendages (top view) bowed, separated by a U-shaped space, ends blunt, lower appendagesnearly as long as upper; greenish with black- brown markings 20Jun-5Sep

    Ophiogomphus colubrinus

    upper appendages triangular with straight edges, separated by a V-shaped space, ends pointed, lower

    appendages shorter than upper; greenish with clear black markings 20Jun-2Aug

    O.anomalus

    18b black legs 18c

    legs with extensor surfaces yellow; greenish with brown markings 20Jun-28Aug O.rupinsulensis

    18c lower appendages much shorter than upper and not projecting out to sides beyond them; greenish,

    brown markings on thorax 15Jun-11Aug

    O.aspersus

    lower appendages little if any shorter than upper and projecting sideways beyond them; yellowishgreen with brown markings 4Jun-7Jul

    O.mainensis

    19 female

    face striped across with black 19a

    face uniformly pale 19b

    19a thorax without a dark central stripe; occiput without spines but with a definite fringe of hairs, horns

    behind occiput stout

    O.colubrinus

    thorax with dark central stripe; occiput with a pair of black spines, hair not in fringe, horns behind

    occiput very small

    O.anomalus

    19b horns behind occiput stout and conspicuous O.rupinsulensis

    horns absent or small 19c

    19c segments 8-9 without yellow spots; with a pair of high contiguous occipital spines O.mainensis

    segments 8 and 9 each with a yellow spot; occipital spines if present small and well separated O.aspersus

    20 Gomphus:

    face entirely yellow-green or olive 20a

    face partly or entirely dark 20g

    20a legs yellow on outside 20b

    legs entirely black or brown 20c

    20b male: upper appendages without spine under, female: occiput with slightly convex margin fringed Gomphus lividus

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    sparsely with hair, vulva less than 1/6th as long as segment 9 with lobes widely separated; dull

    brown and green 14Jun-10Jul

    male: upper appendages with obtuse protrusion under, female: occiput slightly bilobed; dull yellow

    with pale brown thorax stripes 8Jun-12Aug

    G.exilis

    male: upper appendages with spine under, female: occiput with a central prominence and slight

    emargination on each side; dull greenish brown 24May-14Jul

    G.spicatus

    illustrations for 15-20

    20c occiput half height of eye, shape as G.lividus, fringed with long black hair, male: upper appendages'

    ends convergent, slender and not meeting in middle, female: vulva about 1/3 as long as segment 9,

    triangular with a short narrow cleft, no horns on vertex; greenish with black markings 1Jun-17Aug

    Arigomphus furcifer

    occiput height of eye, notched in middle, no hair protruding, male: these ends meeting in middle,

    female: vulva as G.furcifer; yellowish green with brown and black markings 6Jun-27Jul

    A.cornutus

    male: upper and lower appendages' ends parallel, directed rearward, female: vulva less than 1/4 as

    long as segment 9, angle between the lobes obtuse

    20d

    20d segments 7-9 only slightly enlarged, the greatest width less than twice the least width, 8 and 9 with

    scarcely any rim, edge of 8 straight in profile, 10 half as long as 9

    20e

    segments 7-9 greatly enlarged, the greatest width more than twice the least width, 8 and 9 with wide

    rim, edge of 8 convex in profile, 10 1/3 as long as 9 or less

    20f

    20e male: front hamules almost as long as rear, ending in a long slender hook, female: vulva less than 1/2

    as long as segment 9, about as wide as long, not tapering to end; clear brown green and yellow 4Jun-

    30Jul

    Gomphus descriptus

    male: front hamules much shorter than rear, rounded at end, female: vulva more than 1/2 as long assegment 9, much narrower than long, tapering to end; brown green and yellow, very hairy 26May-

    22Jul

    G.borealis

    20f front diameter of segments 8 and 9 about equal, 8 with a single triangular yellow spot on top, occiput

    of female with a small central hump; pale, hairy, wasp waist 1Jun-23Jul

    G.fraternus

    diameter of segment 8 much greater than 9, 8 with two spots on top, occiput of female without

    hump; green with bright brown markings, abdomen blackish, wasp waist 15Jun-14Jul

    G.ventricosus

    20g upper pale thorax stripes flowing together with collar stripe 20h

    these stripes isolated 20i

    20h segments 7-9 as wide as thorax, 8 & 9 with wide curved side rims, 8 more curved than 9, 10 about a

    third as long as 9; thorax greenish yellow with bright brown markings, abdomen blackish with

    yellow at front and back 1May-31Jul

    G.vastus

    segments 7-9 much narrower than thorax, rim of 9 more curved than 8, 10 more than half as long as

    9; yellowish green with strong black markings 7Jun-21Jul

    G.adelphus

    20i abdomen not more than 1/7th longer than hind wing, male: rear hamules leaning somewhat backward 20j

    abdomen 1/5 to 1/3 longer than hind wing, male: rear hamules leaning somewhat forward 20k

    20j small, 32 mm abdomen 28 mm hindwing, pale yellow upper thorax stripes near middle line only

    slightly divergent; darkish, late June

    Stylurus amnicola

    large and robust, 40 mm abdomen 34 mm hindwing, these stripes not near middle line and very

    divergent; blackish with yellow bands on abdomen 25Jun-9Sep

    S.scudderi

    20k segments 8 & 9 about equal in length, male: rear hamules tilted only slightly forward, front edges

    nearly straight; waspwaist, black and greenish yellow 7Jul-12Oct

    S.notatus

    segment 9 about 1/4 longer than 8, male: rear hamules tilted far forward, front edges concave; wings

    shorter than abdomen, black green and yellow 30Jun-16Sep

    S.spiniceps

    21 Cordulegaster:

    single row of yellow abdominal spots on top; July Cordulegaster obliquapaired abdominal spots large, triangular and widely separated and eyes separated by a small space on

    males, usually on females; female ovipositor barely longer than segments 9+10, 28May-14Jul

    C.diastatops

    paired spots small, narrowly separated or confluent, rounded behind on males usually on females,

    eyes meet at point; female ovipositor 2x length of segments 9+10, 29May-14Jul

    C.maculata

    illustrations for 20-21

    22 Libelluloidea:

    hind wing triangle about half as far from arculus as in forewing, anal loop about as long as wide, its

    included cells not in 2 long rows, undeveloped radial and medial planates

    Macromiidae 23

    hind wing triangle at or very close to arculus, anal loop somewhat foot shaped, divided lengthwise

    by a midrib into 2 rows of cells

    22a

    22a forewing triangle 1 times as high as wide, anal loop with little if any development of toe, males: Corduliidae 29

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    well developed anal triangle

    forewing tringle 2x as high as wide, anal loop with well developed toe (except Nannothemis rearwing), male: no anal triangle

    Libellulidae 24

    23 Macromiidae:

    occiput and vertex pale; nodus of forewing about equally distant from stigma and wing base 23May-

    30Jul

    Didymops transversa

    occiput and vertex black; nodus of forewing clearly closer to stigma than to wing base 15Jun-28Aug Macromia illinoiensis

    24 Libellulidae:

    forewing triangle with front side broken making a quadrangle of it; anal loop of hind wing open

    rearward, smallest dragonfly here (wing 20 mm) 20Jun-9Aug

    Nannothemis bella

    triple-length vacant space under stigma; (Lanark 11Jun-30Sep) Pachydiplax longipennis

    vein M2 distinctly waved 24a

    vein M2 smoothly curved 24b

    24a wings with several bridge crossveins Libellula 26

    single bridge crossvein, large brown spot on rear base of hind wing; 13Jul-10Sep Pantala hymenea

    single bridge crossvein, no such spot; 11Jun-27Aug P.flavescens

    24b brilliant three-colour wings: clear, yellow, brown Celithemis 25

    wings almost clear 24c

    24c vein Cu1 in hind wing arising from hind end of triangle Sympetrum 28

    arising from outer side, sometimes only by vein width 24d

    24d forewing triangle inner side less than twice as long as front side, stigma short and thick, face bone

    white (if not check Sympetrum 28)

    Leucorrhinia 27

    inner side more than twice as long as front, stigma about 3 times as long as wide, face greenish with

    brown hairs; radial planate subtends 1 row of cells (if 2 rows, it's Pantala flavescens 24a), mostly

    greenish to grey-blue, brown spots in segments 3-9 19Jun-5Sep

    Erythemis simplicicollis

    25 Celithemis: brilliant yellow and brown spots covering half or more of wings

    4 spots per wing, end one beyond stigma; 18Jun-23Aug C.elisa

    5 spots per wing, end one inside stigma 15Jun-16Aug C.eponina

    26 Libellula: stout body shorter than wing span, medium size, hunch-backed appearance both at rest and

    in flight

    wing clear (may be smoky at tip); forewing triangle 3-celled, eye seam longer than occiput, longer

    and slimmer than L.julia; 21Jun-14Aug

    L.incesta

    wings clear except for small dark basal spots; forewing triangle 2-celled, eye seam shorter than

    occiput; 7Jun-30Jul[I am not certain that the wing basal spot can be used to separate julia from incesta - a few females in

    the national collection have basal spots, eye seam shorter than occiput, but 3-celled triangles.]

    L.julia

    basal third of both wings covered full width by blackish band, tips clear; 26Jun-30Aug L.luctuosa

    nodus wing band at rear wider than 1/5 hindwing length, no spot at stigma, 28May-25Aug male L.lydia

    large clear separate basal, nodal and wing-end spots, forewing basal colour does not extend into

    topmost cell of triangle, female

    female L.lydia

    illustrations for 22-26

    as female L.lydia, but forewing basal colour extends into topmost cell of triangle making a notch in

    its rear; normally longer wingspan than lydia, males have prunescent spots on wings with age; 1Jun-

    28Sep

    L.pulchella

    hind wing with triangular yellow to brown spot at base with white crossveins and dark rear border,

    small clear brown spot at nodus; usually yellow costal streak, sometimes swarms 7May-28Aug

    L.quadrimaculata

    27 Leucorrhinia: white face on black head; small, not stout, wings mostly clear

    all blackish except for prunescence; trigonal interspace of forewing with two rows of cells inner

    third (other L. have 3 rows); 30May-21Aug

    L.frigida

    'blood'-red flush on thorax (only when alive) especially around wing roots, usually on sides of

    segments 1-2 as well, otherwise black abdomen; prunescence can form a row of spots on top of

    abdomen of pattern similar to the yellow spots of L.hudsonica; 31May-19Aug[Many publications use a 2-cell width of the radial planate to key this species. It is inapplicable here;

    several local populations have 1-cell planates.]

    L.glacialis

    with line of broad oval yellow spots on top of segments 1-6 (reddish on some males), usually

    yellow is visible at the bottom of segments 1-2 even through prunescence; a bit stouter than other L.,

    thorax may show a bit of red; 9Jun-30Aug

    L.hudsonica

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    clear yellow or reddish spot on top of segment 7 much larger and brighter than anything on other

    segments; 19May-26Aug

    L.intacta

    white ring, or at least a lot of white, at junction of thorax and abdomen; a bit larger than other L.,

    thorax with no red, the white can be largely obscured by prunescence; 1Jun-10Aug

    L.proxima

    segments 1-3 with line of narrow linear yellow streaks on top (may be faint); check of appendage

    shapes required to be sure (Renfrew 20Jun-7Aug)

    L.patricia

    28 Sympetrum: hybridise often, so expect intermediate forms

    male: upper abdominal appendages with prominent tooth on bottom; female: subgenital plate deeply

    cleft

    28a

    male: these appendages only finely toothed; female: plate with tiny tooth or none 28c28a wing veins reddish, overall brownish red; male: hamuli forked for about 1/4 visible length, rear

    branch 4 times as stout as inner, face cherry red with clear black line bottom quarter of lower lip;21Jun-18Sep

    Sympetrum internum

    wing veins not reddish, little or no black on lower lip or lower edge of black indistinct 28b

    28b reddish throughout, pale stripes on thorax; male: hamuli forked for about 1/3 visible length, rear

    branch barely twice as stout as inner, female: subgenital plate tips upturned against rear of segment

    9, face yellowish red 13Jun-24Sep

    S.rubicundulum

    abdomen with black side triangles, usually linked into a line with triangle-jagged top; initially

    yellow-brown turning red with age, male: hamuli forked for less than 1/4 its visible length, rear

    branch 2-3 times as stout as inner, frons white upper lip yellow-white, female: face greenish white20Jun- 29Oct

    S.obtrusum

    28c wings tinged with dark yellow over basal half, larger area and darker on hindwings; brown, smallpale yellow spots behind head 1Jul-1Oct S.semicinctum

    wings clear, yellow only on costal strip or at extreme base 28d

    28d legs black; only Sympetrum with no red anywhere on male, only black and yellow 3Jul-26Sep S.danae

    legs more or less yellow 18e

    28e legs entirely yellow; yellow when young all red when mature, slender, thin legs 24Jul-3Nov S.vicinum

    legs striped with black on sides; brownish red, wings with yellow veins at tips 14Jul-14Sep S.costiferum

    29 Corduliidae:

    M4 and Cu1 in forewing parallel to divergent 29a

    these veins convergent at wing margin 29b

    29a forewing triangle 3 cells, wings >30 mm; brownish, no visible spots or stripes, flies only briefly after

    dusk 7Jun-30Jul

    Neurocordulia

    yamaskanensis

    triangles and subtriangles each of a single cell, tiny (24 mm); black with reddish lower lip and paleyellow spot on thorax 19May-11June Williamsonia fletcheri

    29b wings with blotchy smoky brown spots at base, nodus and end; hindwing triangle 2-celled, large,brown hairy thorax 7Jun- 20Aug

    Epicordulia (Epitheca)princeps

    no such spots 29c

    29c hindwings with opaque brown spots at base; brownish, face yellowish 25May-7Jul Helocordulia uhleri

    illustrations for 26-29

    hind wing with 2 cubito-anal crossveins Somatochlora 31

    with 1 such crossvein 29d

    29d black abdomen 2x(male) to 3x(female) as wide at end as at waist; male with white abdominal ring

    segment 2 and sometimes 3, green eyes; female yellow-brown segments 1 and 2, brown eyes;

    18May-21Jul

    Dorocordulia libera

    metallic green with green eyes, abdomen only twice as wide at widest point as at narrowest 24May-

    8Aug

    Cordulia shurtleffi

    brown

    30 Epitheca (Tetragoneuria):

    T-spot on frons clearly defined; male: upper appendages each with small sharp spine projectingdownward at about 2/5th of its length 8May-10Jul

    E.spinigera

    T-spot indistinct or incomplete (usually base only) 30a

    30a male: upper appendages with outer tip bent downward, with a bump on upper side where slope

    begins, no crossbar on frons T-spot, female: vulva ends cylindrical extending to margin of segment

    9; stocky 6May-18Jul

    E.canis

    large triangular basal spot rear wings only, row of yellow spots each side of abdomen; male: without

    such a bump, female: ends conical extending well beyond margin of segment 9; hairy thorax 5Jun-

    E.cynosura

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    18Jul

    31 Somatochlora:

    upper lip yellowish, often with black band across front border, face below suture mostly yellow;

    brilliant bronze and green thorax, male: scythe-shaped appendages 1Jul-9Sep

    Somatochlora tenebrosa

    upper lip black, if partly yellow with no such black band 31a

    31a middle segments with clear rings of white about 1/10th their length; one pale stripe/spot on side of

    metallic green thorax

    31b

    no such white rings 31c

    31b thorax stripe barely distinguishable; male: lower appendage squared off, nearly parallel to its widely

    forking tip, female: appendages viewed from side straightish below, much more convex on uppermargin (Tremblant 24Jun-4Sep)

    S.cingulata

    yellow elongate spot each side of thorax; male: lower appendage elongate-triangular, stronglytapering to single upturned tip, female: appendages about equally convex on upper and lower

    margins, 26Jun-1Sep

    S.albicincta

    31c two side stripes on thorax 31d

    only one such stripe; face blackish green with 3 yellow spots, thorax metallic green with brown-

    violet markings 23May- 29Aug

    S.franklini

    31d each thorax stripe represented by round spot; face blackish blue 6Jun-30Aug S.minor

    these spots elongated 31e

    31e first thorax stripe narrower and much longer than 2nd; face brownish yellow with black markings on

    frons, lower lip black 18Jun-1Sep

    S.walshi

    this stripe shorter, about 1/2 thorax height, usually narrower than 2nd 31f31f both thorax stripes bright yellow and sharply defined 31g

    these stripes dull yellow and ill-defined 31h

    31g middle segments with pale spots at front bases; brown with metallic sheen 27May-27Aug S.forcipata

    these segments with no pale spots; yellowish face, barely visible yellow bars on metallic green

    thorax with light brown hairs (Tremblant 8Jun-30Aug)

    S.elongata

    31h male: upper appendages very hairy, tips recurved, female: subgenital plate triangular in profile, taper

    pointed and erect; dark with patches of dark yellow, hairy thorax (Tremblant 21Jun-15Sep)

    S.williamsoni

    male: upper appendages not hairy, not recurved at tips, lower curved upwards, female: subgenital

    plate broadly triangular, not erect; blackish face with some pale yellow, hairy thorax 5Jun-11Jul

    S.kennedyi

    male: upper appendages not hairy, not recurved at tips, lower not curved upwards, female: subgenital

    plate cylindrical; segment 2 with single small pale spot each side, little thorax hair (19Jul-30Aug)

    S.incurvata

    illustrations for 30-31main page

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