Theories on the ORIGIN OF LIFE
Basic enigma of life: HOW DID LIFE ORIGINATE?
Nothing is directly known about the origin of life, the answer
to this question are exceedingly complex and at best only
tentative.
Theories of the origin of life: Still a
matter of speculation
Several intelligent explanations
account for the origin of life on earth
Religion
The origin of life must be attributed to an
agency outside nature called a creator.
Mythology Philosophy
1. THEORY OF SPECIAL CREATION
support or acceptance mostly due to faith rather than experimental or scientific evidences.
Supporters recently created a new discipline called “creation science”.
Proposed that universe started from a primeval fireball and had
been expanding and cooling since its inception 10-20 billion years
ago (bya).
Life originated from outer planets in the form of a resistant spore
(cosmozoa) propelled by radiation pressure, reached earth and
started the first form of life.
Idea was proposed by Richter in
1865. The theory did not gain any
support. Needs evidence for the
existence of ET life.
2. Cosmozoic or Interplanetary
The cosmozoic theory speculates that life
arrived on Earth as bacterial spores,
perhaps enclosed in a comet.
3. ABIOGENESIS (SPONTANEOUS GENERATION)
600 BC up to 2nd half of 19th century – believed that life could arise
spontaneously from nonliving substances.
Living organisms originated in sea lime under the influence of
factors in the environment s.a. heat, air, sun.
Thales (624-548 BC) ---“oceanic
water was the mother from which all
living forms originated”.
The origin of life without apparent cause
3. ABIOGENESIS (SPONTANEOUS GENERATION)
Aristotle‟s Hypothesis
Aristotle (384-322 BC)--- proposed
that living forms are animated forms of
non-living matter.
-Vital forces operate constantly and
improve the living world
Empedocles (540-433 BC) ---“life
originated by itself from non living matter
and imperfect forms were replaced by
perfect forms”
Biogenesis. Every living thing on earth is the offspring of other living things.
“Life comes from life” is referred to as the law of biogenesis, which asserts that
modern organisms do not spontaneously arise in nature from non-life.
Louis Pasteur
Working hypothesis: life arose from pre-existing life
Anton van Leeuwenhoek
Believed that microorganisms arose
from pre-existing organisms.
4. Biogenesis Theory
Francisco Redi
opposed abiogenesis
proposed that life could arise only
from pre-existing living things (thru
experiment) .
Louis Pasteur was the first to
be able to prove this theory
proposed that the organisms
that are not visible to the
naked eye are present in air.
Broth is boiled. Broth is free of
microorganisms
for a year.
Curved neck
is removed.
Broth is
teeming with
microorganisms
.
Pasteur’s Experiment
demonstrated that fermentation is caused by the growth
of micro-organisms, and that the emergent growth of bacteria in
nutrient broths is not due to spontaneous generation
OBSERVATIONS: Flies land on meat that is left uncovered. Later, maggots appear on the meat.
HYPOTHESIS: Flies produce maggots.
PROCEDURE
Controlled Variables: jars, type of meat, location, temperature, time
Manipulated Variables: gauze covering that keeps flies away from meat
Responding Variable: whether maggots appear
CONCLUSION: Maggots form only when flies come in contact with meat.
Spontaneous generation of maggots did not occur.
Redi’s Experiment
5. Natural or Marine (Primeval Soup)
Proposed that life did not originate in the surface of the earth but deep beneath the sea in or around hydrothermal vents.
In 1929 by J.B.S. Haldane; suggested that life was the result of UV radiation converting methane, ammonia and water into the first organic compounds in the early earth oceans.
6. Physico-chemical or coacervate droplet
theory (Oparin and Haldane) Chemical evolution:
1. Formation of simple organic compounds
the primitive inorganic molecules of earth interacted and combined
with one another to form simple organic compounds. These were in
the form of simple sugars, fatty acids, glycerol, amino acids and
nitrogen bases.
2. Formation of complex organic compounds
-Simple sugars combined, form complex polysaccharides (starch,
cellulose). Fatty acids and glycerol molecules combined to form
lipids. Amino acids combined forming polypeptides and proteins.
-Purines and pyrimidines combined with simple sugars and phosphates
to form nucleotides, which then formed nucleic acids.
Harold C. Urey and Stanley L. Miller
(1953) – conducted an experiment
simulating the primitive condition of the
Earth.
− discovered that a variety of amino
acids and organic acids were formed
Urey-Miller hypothesis
Proposed that amino acids can
be synthesized outside living
systems.
They conducted experiments
in which a gas mixture
containing hydrogen, ammonia,
methane and water vapor was
subjected to electric spark.
It yielded aldehydes, amino
acids and carboxylic acids.
3. Formation of molecular aggregates (Coacervates):
•Oparin and Fox proposed that the complex organic molecules synthesized
abiotically on the primitive earth formed large spherical aggregates as
cluster of complex organic molecules bound by fatty acids and divide.
•They remained suspended as droplets in sea water
•The coacervates had all the basic properties of living cells like,
metabolism, growth etc.
•However they lacked the complexity of the living cells like organelles.
•Thus these particles with proteins as enzymes and ATP as source of
energy were the first structures at the margin of non-living and living.
4. Formation of first primitive living cell:
Theories on the ORIGIN OF LIFE
Basic enigma of life:
Nothing is directly known about the origin of life, the answer
to this question are exceedingly complex and at best only
tentative.
Theories of the origin of life: Still a
matter of speculation
Several intelligent explanations
account for the origin of life on earth
HOW DID LIFE ORIGINATE?
Fossils as evidence of change
Features of Earliest Organisms
Extinction
• the earth is about 4.5 billion years old,
• the earliest known cells are found in 3.5 billion year old
rocks
• the earliest known eukaryotic cells date to 1.5 billion years
• the earliest multicellular animals date to 650 million years
• the earliest land animals date to about 450 million years
• the earliest mammals date to about 230 million years
• 65 m.y. ago there was a mass extinction of many living things
• the human family tree diverged from the other apes about
4.5 million years ago
Fossils as evidence of change
Remains or traces of prehistoric life
preserved remains of animals, plants or their parts
Can be of entire organisms or a part which got
buried, a mould or cast, foot prints or imprints on a
stone.
• Physical evidence of organisms from the past
• Provides visible evidence that takes us back in time and shows organisms
have changed.
• The fossil record provides incomplete information about the history of life. • Paleontologists are scientists who collect and study fossils.
• Over 99% of all species that have lived on Earth have become extinct.
• Where are all these fossils??
Fossils as evidence of change
Careful study of fossils
Opens a window into the lives of organisms that existed long ago and
provides information about the evolution of life over billions of years
The strata of one location can often be correlated with strata at
another location by the presence of index fossils
Index fossils (also known as guide fossils, indicator fossils or zone
fossils) are fossils used to define and identify geologic periods (or
faunal stages).
Example of Index fossils: Shelled animals called brachiopods were extremely abundant in ancient seas. Their fossils are useful indicators of the relative ages of rock strata in different locations
Methods of fossilization
Petrifaction- Turn into
stones.
This is due to formation
of sedimentary rocks
under water.
soft parts: disappear;
hard parts: preserved
due to mineralization.
muscles and other
soft organs: get
mineralized and form
rocky fossils.
Preservation of foot prints
prints, if left undisturbed: hardened and form rocky
fossils.
such imprints can provide clues regarding the
body form and characteristics of the extinct
animal.
A dinosaur footprint
Preservation in ice
Entire animals can get
frozen and may be
preserved
Body parts remain
intact without change.
E.g. woolly mammoth
from Siberia
Fossils: evidence of past life Types of fossils
• Carbonization – organic
matter becomes a thin
residue of carbon
• Preservation in amber
– hardened resin of
ancient trees surrounds an
organism
• Impression – replica of
the fossil's surface
preserved in fine-grained
sediment
Impressions: Moulds and cast
• Mold – a part/structure is buried and then dissolved by underground water • Fossilized moulds are found in volcanic ashes.
• Cast – hollow space of a mold that is filled with mineral matter
Natural casts of shelled
invertebrates
Natural mould of a trilobite
Types of fossils
• Petrified– cavities and pores are filled with precipitated mineral matter
• Permineralization– the pores of the organism are filled with minerals but some tissues of the plant or animal are still intact and are not replaced with minerals.
• Formed by replacement - cell material is removed and replaced with mineral matter
Fossils: evidence of past life
Trace – aka ichnofossils.
May be impressions made
on the substrate by an
organism. For ex: burrows,
footprints and feeding
marks, and root cavities.
Coprolites – fossil dung
and stomach contents
Gastroliths – stomach
stones used to grind food
by some extinct reptiles
Indirect fossil evidence includes:
The History of Life
Relative dating is a method used to
determine the age of rocks by
comparing them with those in other
layers.
Rock layers form in order by age—
the oldest on the bottom, with more
recent layers on top.
Dating Fossils
Radioactive dating (Absolute Dating)
• Uses the decay of radioactive isotopes to measure the age of a rock. the use of half-lives to determine the age of a sample. A half-life is the length of time required for half of the radioactive atoms (C, N) in a sample to decay.
Dating Fossils
Early members of the animal fossil record
◦ Include the Ediacaran fauna
Important fossils
(features of earliest organisms)
These animals were the precursors of organisms with
skeletons.
Ichthyostega - interconnecting link
between fishes and amphibians.
Important fossils
(features of earliest organisms)
Seymouria - Interconnecting link
between Amphibians and Reptiles.
Important fossils
(features of earliest organisms)
Dinosaurs - Extinct group of reptiles.
Important fossils
(features of earliest organisms)
Archaeopteryx - Ancestral form of birds
Important fossils
(features of earliest organisms)
Hyracotherium - Early ancestor of
horses.
Important fossils
(features of earliest organisms)
Theropods
Allosaurus
Sinornis
Velociraptor
Archaeopteryx
Robin
Light bones
3-toed foot;
wishbone Down
feathers
Feathers with
shaft, veins,
and barbs
Flight feathers;
arms as long
as legs
Important fossils
(features of earliest organisms)
Rodhocetus, an
ancient whale, lived
about 47 million
years ago.
Rodhocetus ankle bone
(left), a modern
artiodactyl, pronghorn
antelope ankle bone
(right).
Its distinctive ankle
bones point to a
close evolutionary
connection to
artiodactyls.
Important fossils
(features of earliest organisms)
Dorudonatrox, an ancient whale that lived about
37 million years ago.
Important fossils
(features of earliest organisms)
Extinction
The termination of a lineage without issue
or abrupt disappearance of specific groups
of organisms without leaving descendents.
The death of a species or group of taxa
Extinction Types: True Extinction and Pseudoextinction
1. True Extinctions -particular lineage totally disappears
without any progeny or evolutionary descendents.
E.g. extinction of dinosaurs as a group ; trilobites.
2. Pseudoextinction (phyletic extinction or phyletic
transformation) - a group may disappear leaving descendents
with evolutionary modifications.
Example:
Horse evolution while the earliest ancestor became extinct its
descendant survived to produce the modern „Equus‟.
Pattern of extinction
Major groups of herbivorous vertebrates are more susceptible than the carnivorous vertebrates
Larger organisms easily became extinct.
VanValen (1973) recorded a constancy in the rate of extinction in a number of groups.
Explained using „Mac Arthor‟s law‟ - “every new
adaptation encourages the survival of a possessor it also decreases a fitness of other species of that area”.
Causes of extinction
(1) A mass extinction: due to drastic changes in the environmental conditions.
(2) Any adaptive advance in one species decreases the fitness of all other species.
Red Queen‟s hypothesis : you have to keep running pretty fast, just in order to stay in the same place.
(3) Over specialization to a specific situation may cause extinction
(ex. Antlers)
(4) The spread of an epidemic disease without any control can cause extinction.
(5) An increase in the population strength of herbivorous animals cause rapid food shortage and cause extinction for several inter-related groups (Predation, competition)
Causes of extinction
(6) A sudden cosmic radiation can
cause the death of large organisms.
(7) A dust storm formed due to falling
of a meteorite is commonly
mentioned as a cause for the
disappearance of dinosaurs.
(8) Habitat degradation: The
degradation of a species' habitat may
alter the fitness landscape or such an
extent that the species is no longer
able to survive and becomes extinct.
Causes of extinction
(9) Coextinction: the loss of a species due to the extinction of another; for example, the extinction of parasitic insects following the loss of their hosts
(10) Genetic Pollution: uncontrolled hybridization, introgression genetic swamping which leads to homogenization or replacement of local genotypes as a result of a numerical and/or fitness advantage of the introduced plant or animal
Causes of extinction
In the recorded history of earth,
extinctions of major groups of organisms
were due to natural causes.
By end of Permian period of the Paleozoic
Era, nearly 60% of the varieties then
existed, became extinct.
Similar large scale extinctions have been
observed by the end of Mesozoic era
Causes of extinction
2006 IUCN Red List data.
2006 IUCN Red List data.
At present … mostly due to human interference.
The realization of such a cause lead to starting of several international voluntary agencies to monitor and control extinctions.
“The red-data book brought out regularly by W. W. F (World Wide Fund for nature, IUCN - international union for the conservation of nature and natural resources) provides a list of animals and plants that are endangered or have become extinct.
Causes of extinction