Wildlife objectives: mixed-conifer dry forest · Home range area as indicator of habitat...

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Wildlife objectives: mixed-conifer dry forest John Lehmkuhl Research Wildlife Biologist Wenatchee Forestry Sciences Lab

Transcript of Wildlife objectives: mixed-conifer dry forest · Home range area as indicator of habitat...

Page 1: Wildlife objectives: mixed-conifer dry forest · Home range area as indicator of habitat quality….. 4.6 ha. 2.7 ha. 2.3 ha. Open Pine. Young. Mature > = Flyers in PIPO foraged .

Wildlife objectives: mixed-conifer dry forest

John LehmkuhlResearch Wildlife BiologistWenatchee Forestry Sciences Lab

Page 2: Wildlife objectives: mixed-conifer dry forest · Home range area as indicator of habitat quality….. 4.6 ha. 2.7 ha. 2.3 ha. Open Pine. Young. Mature > = Flyers in PIPO foraged .

Flying squirrel densityLincoln-Peterson estimator

Year2000199919981997

Mea

n de

nsity

3.0

2.5

2.0

1.5

1.0

0.5

Open pine

Young

Mature

Open pine forest consistently is poorer habitat.

Page 3: Wildlife objectives: mixed-conifer dry forest · Home range area as indicator of habitat quality….. 4.6 ha. 2.7 ha. 2.3 ha. Open Pine. Young. Mature > = Flyers in PIPO foraged .

Home range area as indicator of habitat quality…..

4.6 ha2.7 ha 2.3 ha

Open Pine Young Mature

> =

Flyers in PIPO foraged in mixed conifer stands

Page 4: Wildlife objectives: mixed-conifer dry forest · Home range area as indicator of habitat quality….. 4.6 ha. 2.7 ha. 2.3 ha. Open Pine. Young. Mature > = Flyers in PIPO foraged .

Low canopy closure = fewer flying squirrels

Canopy cover (%)9080706050403020

GLS

A d

ensi

ty

3.0

2.5

2.0

1.5

1.0

0.5

Mature

Young

Open pine

Page 5: Wildlife objectives: mixed-conifer dry forest · Home range area as indicator of habitat quality….. 4.6 ha. 2.7 ha. 2.3 ha. Open Pine. Young. Mature > = Flyers in PIPO foraged .

Flyer fitness increases with richness of plant understories…

Understory species richness

60504030

Sur

viva

l or R

ecru

itmen

t rat

e0.6

0.5

0.4

0.3

0.2

survival rate

recruitment rate

r = 0.98

r = 0.98

Page 6: Wildlife objectives: mixed-conifer dry forest · Home range area as indicator of habitat quality….. 4.6 ha. 2.7 ha. 2.3 ha. Open Pine. Young. Mature > = Flyers in PIPO foraged .

Truffle biomass important for survival & recruitment….

survival

recruitment

0.1

0.2

0.3

0.4

0.5

0.6

0.7

0 1 2 3 4 5 6 7

Truffle biomass (kg/ha)

Prob

abili

ty

Page 7: Wildlife objectives: mixed-conifer dry forest · Home range area as indicator of habitat quality….. 4.6 ha. 2.7 ha. 2.3 ha. Open Pine. Young. Mature > = Flyers in PIPO foraged .

Truffle biomass & richness is higher in cool-moist environments

aspect gradient index4.03.02.01.0

Bio

mas

s / R

ichn

ess

14

12

10

8

6

4

2

0

biomassr = 0.7

richnessr = 0.8

Page 8: Wildlife objectives: mixed-conifer dry forest · Home range area as indicator of habitat quality….. 4.6 ha. 2.7 ha. 2.3 ha. Open Pine. Young. Mature > = Flyers in PIPO foraged .

Canopy closure (%)9080706050403020

Bio

mas

s (k

g/ha

)

7

6

5

4

3

2

1

0

Biomass

r = 0.75

Truffle food increases with canopy closure

Page 9: Wildlife objectives: mixed-conifer dry forest · Home range area as indicator of habitat quality….. 4.6 ha. 2.7 ha. 2.3 ha. Open Pine. Young. Mature > = Flyers in PIPO foraged .

Coarse woody debris cover (%)403020100

Bio

mas

s / R

ichn

ess

14

12

10

8

6

4

2

0

Biomassr = 0.80

Richnessr = 0.85

Truffle richness & biomass increases with woody debris cover…

Page 10: Wildlife objectives: mixed-conifer dry forest · Home range area as indicator of habitat quality….. 4.6 ha. 2.7 ha. 2.3 ha. Open Pine. Young. Mature > = Flyers in PIPO foraged .

Flyer survival rate increases with forage lichen biomass……

0.40

0.45

0.50

0.55

0.60

0 2 4 6 8 10

Lichen biomass (kg/ha)

Prob

abili

ty

survival

Page 11: Wildlife objectives: mixed-conifer dry forest · Home range area as indicator of habitat quality….. 4.6 ha. 2.7 ha. 2.3 ha. Open Pine. Young. Mature > = Flyers in PIPO foraged .

Lichen biomass highest in old moist stands

0

1

2

3

4

5

6

7

Bryoria spp. All Species

Bio

mas

s (k

g/ha

)

OpenModerateMature

foragelichens

Page 12: Wildlife objectives: mixed-conifer dry forest · Home range area as indicator of habitat quality….. 4.6 ha. 2.7 ha. 2.3 ha. Open Pine. Young. Mature > = Flyers in PIPO foraged .

Bushy-tailed woodrat density

Density groups

0.13

0.28

0.54

0.93

0.00.10.20.30.40.50.60.70.80.91.0

Low Mod.Low

Mod.High

High

Den

sity

(n/h

a)

n = 3 n = 2 n = 4 n = 3

Page 13: Wildlife objectives: mixed-conifer dry forest · Home range area as indicator of habitat quality….. 4.6 ha. 2.7 ha. 2.3 ha. Open Pine. Young. Mature > = Flyers in PIPO foraged .

Woodrat density groupslow high

Large snag density <20/ha

Mistletoe index <5.0

Soft log cover <1.3%

0.13 0.28 0.54 0.93

Group mean density(woodrats/ha)

Woodrats most abundant

where more snags,

mistletoe, & logs

BT woodrats density highest where dead wood & mistletoe…

Page 14: Wildlife objectives: mixed-conifer dry forest · Home range area as indicator of habitat quality….. 4.6 ha. 2.7 ha. 2.3 ha. Open Pine. Young. Mature > = Flyers in PIPO foraged .

Deer mouse

no yes-1.00

-0.25

0.50

1.25

2.00

Log(

dens

ity)

unburnedburned

Thinned

Yellow-pine chipmunk

-2.00

-0.75

0.50

1.75

3.00

no yes

Treatments increase dominant small mammals

Thinned

thinned thinnedburned

Page 15: Wildlife objectives: mixed-conifer dry forest · Home range area as indicator of habitat quality….. 4.6 ha. 2.7 ha. 2.3 ha. Open Pine. Young. Mature > = Flyers in PIPO foraged .

Retain mesic patches of stands & landscapes…

Mesic stands Dry stands "Mesic-site" spp. southern red-backed vole 1 long-tailed vole 1 montane shrew 1 1 1 Townsend's chipmunk 1 1 creeping vole 1 1 1 1 1 1American shrew-mole 1 1 1deer mouse 3 3 3 3 2 2 1 1 1 1 2 1Trowbridge's shrew 1 2 2 1 1 1 2northern pocket gopher 1 1 1 1 1 "Dry site" spp. Great Basin pocket mouse 1 1 1 1 1vagrant shrew 1 1 1 1 1yellow-pine chipmunk 1 1 1 1 1 1 1 1 2 2 2 1

Page 16: Wildlife objectives: mixed-conifer dry forest · Home range area as indicator of habitat quality….. 4.6 ha. 2.7 ha. 2.3 ha. Open Pine. Young. Mature > = Flyers in PIPO foraged .

Treatment objectives to test…

Flying squirrelsMaintain 50% closure overall, but patchy…Leave patches large trees (lichens, truffles).Create open patches (understory richness).Large down wood (truffles).Large old snags, large mistletoe brooms (dens).

Bushy-tailed woodratsLarge old snags & replacements.Large mistletoe brooms.Large down wood.

Other small mammalsPatchy dense & open stands maintain diversity.