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Island Specialists: Shared Flora of the Alta and Baja California Pacific Islands Authors: Ratay, Sarah E., Vanderplank, Sula E., and Wilder, Benjamin T. Source: Monographs of the Western North American Naturalist, 7(1) : 161-220 Published By: Monte L. Bean Life Science Museum, Brigham Young University URL: https://doi.org/10.3398/042.007.0116 BioOne Complete (complete.BioOne.org) is a full-text database of 200 subscribed and open-access titles in the biological, ecological, and environmental sciences published by nonprofit societies, associations, museums, institutions, and presses. Your use of this PDF, the BioOne Complete website, and all posted and associated content indicates your acceptance of BioOne’s Terms of Use, available at www.bioone.org/terms-of-use. Usage of BioOne Complete content is strictly limited to personal, educational, and non - commercial use. Commercial inquiries or rights and permissions requests should be directed to the individual publisher as copyright holder. BioOne sees sustainable scholarly publishing as an inherently collaborative enterprise connecting authors, nonprofit publishers, academic institutions, research libraries, and research funders in the common goal of maximizing access to critical research. Downloaded From: https://bioone.org/journals/Monographs-of-the-Western-North-American-Naturalist on 24 Jun 2020 Terms of Use: https://bioone.org/terms-of-use

Transcript of ,V OD Q G 6 S H F LD OLV WV 6 K D UH G ) OR UD R I ......San Martín—Junak and Philbrick 1994b...

Page 1: ,V OD Q G 6 S H F LD OLV WV 6 K D UH G ) OR UD R I ......San Martín—Junak and Philbrick 1994b Guadalupe—Moran 1996, Rebman 2006, Rebman et al. 2007 San Benito—Junak and Philbrick

Island Specialists: Shared Flora of the Alta and BajaCalifornia Pacific Islands

Authors: Ratay, Sarah E., Vanderplank, Sula E., and Wilder, BenjaminT.

Source: Monographs of the Western North American Naturalist, 7(1) :161-220

Published By: Monte L. Bean Life Science Museum, Brigham YoungUniversity

URL: https://doi.org/10.3398/042.007.0116

BioOne Complete (complete.BioOne.org) is a full-text database of 200 subscribed and open-access titlesin the biological, ecological, and environmental sciences published by nonprofit societies, associations,museums, institutions, and presses.

Your use of this PDF, the BioOne Complete website, and all posted and associated content indicates youracceptance of BioOne’s Terms of Use, available at www.bioone.org/terms-of-use.

Usage of BioOne Complete content is strictly limited to personal, educational, and non - commercial use.Commercial inquiries or rights and permissions requests should be directed to the individual publisher ascopyright holder.

BioOne sees sustainable scholarly publishing as an inherently collaborative enterprise connecting authors, nonprofitpublishers, academic institutions, research libraries, and research funders in the common goal of maximizing access tocritical research.

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The islands of the California Floristic Prov -ince (CFP) are an important component of oneof the world’s great biodiversity hotspots(Myers et al. 2000). The CFP has commonlybeen referred to as an environmental islanddue to its regionally unique mediterranean-typeclimate, isolation (via oceanic and orographicboundaries), and corresponding globally sig-nificant levels of endemism. The plant com-munity has had ample time to evolve sincethe emergence of wet winters and dry sum-mers at least 5–10 Mya (Axelrod 1973) andperhaps earlier (Ackerly 2009, Keeley et al.2011). The remarkable diversity and endemismof the CFP has been attributed to relative cli-matic stability during glacial-interglacial tran-sitions of the Pleistocene (Lancaster and Kay2012, Sniderman et al. 2013). The antiquityand distinctiveness of the California flora are

well-recognized attributes that are accentuatedon the islands of the Californias.

The degree of floristic similarity of 16 off-shore islands, and their apparent role as refugiafor species once present on the mainland, hasbeen a persistent biogeographical puzzle. TheCFP includes 16 islands from the adjacentcoast of Alta and Baja California and extendssouth to the 28th parallel, including the CFPvegetation found on Cedros and GuadalupeIslands (Raven and Axelrod 1978, Moran 1996,Oberbauer 2002a). The CFP has an increasinggradient of aridity from north to south, butvegetation distributions are heavily influencedby fog presence; this fog provides additionalmoisture and reduces radiant loadings, signifi-cantly changing conditions for plants (Dawson1998, Fischer et al. 2008, Vanderplank 2013).Physical data suggest that ocean currents have

Monographs of the Western North American Naturalist 7, © 2014, pp. 161–220

ISLAND SPECIALISTS: SHARED FLORA OF THE ALTA AND BAJA CALIFORNIA PACIFIC ISLANDS

Sarah E. Ratay1, Sula E. Vanderplank2, and Benjamin T. Wilder3

ABSTRACT.—The floristic connection between the mediterranean region of Baja California and the Pacific islands ofAlta and Baja California provides insight into the history and origin of the California Floristic Province. We presentupdated species lists for all California Floristic Province islands and demonstrate the disjunct distributions of 26 taxabetween the Baja California and the California Channel Islands. These 26 plant taxa are found among the 16 Pacificislands without occurring on the intervening mainland of Alta California. Separate species lists for each island group (8California Channel Islands and 8 Baja California Islands) were compiled. These lists were compared to the mainland Cali-fornia flora to identify species that occur on the California Islands and either the Baja California Pacific Islands or themediterranean region of the Baja California Peninsula, but not the mainland of Alta California. This first compilation ofthe flora of the Baja California Islands and nomenclatural updates for the Channel Islands provide a platform for futureresearch and conservation planning.

RESUMEN.—El grado de conexión florística entre la región mediterránea de Baja California y las Islas del Pacífico deAlta y Baja California proporciona un entendimiento de la historia y el origen de la Provincia Florística de California.Presentamos listados actualizados de especies de todas las islas de la Provincia Florística de California y mostramos ladistribución aislada de veintiséis taxones entre la Baja California y las Islas del Canal de California. Identificamos aquel-las especies de plantas que se pueden encontrar dentro de las dieciséis islas del Pacífico, pero no se encuentran en lapenínsula de Alta California. Con este objetivo, recopilamos listados separados de las especies que habitan en cadagrupo de islas (las ocho Islas del Canal de California y las ocho islas de Baja California). Estos listados se compararoncon la flora existente en la península de California con el fin de identificar aquellas especies que se encuentran en lasislas de California, y tanto en las Islas del Pacífico de Baja California como en la región mediterránea de la Península deBaja California, pero no en la península de Alta California. La primera recopilación de flora de las Islas de Baja Califor-nia y actualizaciones nomenclaturales en las Islas del Canal, proporcionan una plataforma para futuras investigaciones yplanes de conservación.

1University of California, Los Angeles, Department of Ecology and Evolutionary Biology, 612 Charles E. Young Drive East, Box 957246, Los Angeles, CA90095. E-mail: [email protected]

2Botanical Research Institute of Texas, 1700 University Dr., Fort Worth, TX 76107.3University of California, Riverside, Department of Botany and Plant Sciences, 900 University Avenue, Riverside, CA 92507.

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been stable for millions of years (Jacobs et al.2004), which indicates the presence of fogduring the development of the CFP.

Within the large number of plant specieson the CFP islands and the mainland of Altaand Baja California is the occurrence of dis-junct Baja California taxa on the CaliforniaChannel Islands. These species are seen todayonly on the CFP islands and a distinct portionof the California mainland termed the DieganScrub, the San Diego region southward intocoastal northwestern Baja California (Fig. 1;Axelrod 1978). However, the number and iden-tity of this group of disjunct species has re -mained unknown.

To identify the species that define this pat-tern, we present (1) the first compiled check-list of the 8 CFP Baja California Islands, (2)

nomenclatural updates to the flora of the 8California Channel Islands, and (3) the plantsthat occur on the California Channel Islandsand in the Diegan Scrub but not the rest ofmainland California, hereafter referred to as“exemplar taxa.”

METHODS

Development of Checklists

We assembled all existing species checklistsfor the 2 island groups (Alta California andBaja California) into 2 lists, then comparedthose lists with the flora of the CaliforniaFloristic Province for both mainland Alta andBaja California to generate the list of exemplartaxa. The nomenclatural updates presented hereare based on our best understanding of the

162 MONOGRAPHS OF THE WESTERN NORTH AMERICAN NATURALIST [Volume 7

Fig. 1. California Channel and Baja California Pacific Islands with Diegan Scrub as defined by Axelrod (1978). Mapby B.T. Wilder, adapted from Google Earth™ imagery.

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island floras at the date of publication and fol-low the models of the sources indicated belowfor the 2 lists. Species now thought to be extir-pated from the islands are denoted by anampersand (&). Unvouchered taxa (includingreports, photo vouchers, and personal commu-nications) are indicated by a question mark (?).Nonnative species are indicated by an asterisk(*). When a subspecies was not recorded forall the islands and some ambiguity remains,the taxon is recorded at the species level only.Exotic species that were planted or are notnaturalized (e.g., Nerium oleander) were deletedfrom checklists.

Baja California Islands Checklist

The most recent species lists for each ofthe 8 Baja California Islands (Appendix 1;Coronados, Todos Santos, San Martín, Jeron-imo, Guadalupe, San Benito, Cedros, andNatividad) were compiled from the followingdata sources:

Coronados—Oberbauer 2002bTodos Santos—Junak and Philbrick 1994aSan Martín—Junak and Philbrick 1994bGuadalupe—Moran 1996, Rebman 2006, Rebman et

al. 2007San Benito—Junak and Philbrick 2002b, Rebman 2007aCedros—Oberbauer 1987, Rebman 2007bNatividad—Junak and Philbrick 2002a

The list of Jeronimo island flora, consisting of8 taxa, was provided via personal communica-tion from Steve Junak. We verified mainlandBaja California flora distributions with theunpublished data of Jon Rebman and BartO’Brien (personal communication 2013). Sev-eral taxa were added to the checklists basedon recent collections, herbarium specimensrecently encountered, and recent reports bySula Vanderplank, Jon Rebman, and SteveJunak; these taxa are indicated by footnotes.The taxonomy for this checklist follows theChecklist of Baja California Plants in prepara-tion by Jon Rebman.

California Channel Islands Checklist

The updated flora of the 8 California Chan-nel Islands (Appendix 2; San Clemente, SanNicolas, Santa Catalina, Santa Barbara, SantaRosa, Santa Cruz, Anacapa, and San Miguel) isbased on the master checklist complied byGary Wallace (1985). We included publishedupdates to individual island floras found in the

National Park Service checklist from Junak etal. (1997) for San Miguel, Santa Rosa, SantaCruz, Anacapa, and Santa Barbara islands. ForSan Clemente Island we included updatesfrom Ross et al. (1996) and for San NicholasIsland updates from Junak (2008). Additionaltaxa and island records are included based onrecent collections on Santa Catalina Island bySarah Ratay, observations and collections onSan Clemente Island by Emily Howe (SoilEcology and Restoration Group, SDSU), andherbarium specimens found in the Consor-tium of California Herbaria (CCH 2014), whichare listed in footnotes. The nomenclatural re -vision of this checklist is consistent with tax-onomy of the new Jepson Manual (Baldwin etal. 2012) through the use of the dynamic con-cordance tool provided on the Jepson e-florawebsite (Jepson Flora Project 2013). Thischecklist was developed to identify distribu-tion patterns in the flora, and though thenomenclature has been updated, we have notconfirmed identifications of questionable her -barium specimens, nor have we diligently pur-sued the lowest taxonomic rank beyond thepublished sources. We are also aware of addi-tional updates to the flora of many of theChannel Islands in process by Steve Junak,including publication of the flora of CatalinaIsland; Junak’s updates are not included here.

Exemplar Taxa

The identification of exemplar taxa wasgenerated through a comparison of the 2 newlygenerated island checklists (Appendixes 1, 2)with the Jepson Manual (Baldwin et al. 2012)and the inventory of rare and endemic plantsof CFP Baja California (O’Brien et al. 2014). Alarge number of species from CFP Baja Cali-fornia extend just slightly into southern Califor -nia and are considered near-endemic to BajaCalifornia (O’Brien et al. 2014). As such, weinclude the near-endemic species of CFPBaja California as ‘absent’ in the mainland ofAlta California (with the exception of Crosso-soma californicum, which occurs on the PalosVerde Peninsula, and Euphorbia misera, whichoccurs in the South Coast region of Californiaand beyond; Baldwin et al. 2012). These fewspecies are consistent with the concept of Die-gan Scrub as proposed by Axelrod (1978).Diegan Scrub is named for its occurrence insouthern San Diego County and south intoBaja California. It is a coastal scrub, rich in

2014] CALIFORNIA ISLAND PLANT DISTRIBUTION PATTERNS 163

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succulents, that includes many endemic andnear-endemic species from CFP Baja; it alsoincludes the coastal succulent scrub and coastalsage scrub ecoregions (González-Abraham etal. 2010).

RESULTS

The California Pacific Islands in total contain1239 unique taxa. The California Islandsseparately contain 976 taxa and the Baja Califor -nia Islands 535 taxa. Twenty-six of these taxa havea disjunct occurrence between the Baja Cali-fornia Pacific Islands and the California Chan-nel Islands, absent from Alta California out-side the Diegan Scrub (Table 1). These exem-plar taxa represent 16 families and 26 uniquegenera. No one growth form predominates; ofthe 26 taxa, there are 12 shrubs, 10 herba-ceous annuals, 2 trees, 1 vine, and 1 xerophyticsucculent (Table 1). Fabaceae is the largestfamily with 3 species on the exemplar taxa list.

These 26 taxa fall into 6 biogeographic sub-patterns:

(1) multiple California Channel Islands andGuadalupe Island, 13 taxa

(2) Guadalupe Island–San Clemente Island only,4 taxa (Hazardia cana, Leptosiphon pyg-maeus ssp. pygmaeus, Lupinus guadalupen-sis, Phacelia floribunda)

(3) Diegan Scrub and the Channel Islandsonly, 2 taxa (Prunus ilicifolia ssp. lyonii andSalvia brandegeei)

(4) widespread, 1 taxon (Eschscholzia ramosa)occurs on 14 of 16 islands

(5) certain Channel and Baja California Islandsand Diegan Scrub, 4 taxa (Bergerocactusemoryi, Crossosoma californicum, Euphor-bia misera, Ribes viburnifolium)

(6) certain Channel and Baja California Islands,2 taxa (Calystegia macrostegia ssp. macro -stegia and Senecio lyonii)

DISCUSSION

The complex biogeography of the Califor-nia Floristic Province can be explained in part

164 MONOGRAPHS OF THE WESTERN NORTH AMERICAN NATURALIST [Volume 7

TABLE 1. The 26 exemplar taxa as they occur on 8 Baja California Pacific Islands and the 8 Channel California Islands.aCRPR column provides the California Rare Plant List ranking, and the BC column provides the State of Baja California

Family Exemplar taxa Habit NAT CED BEN GUA JER MAR

Apiaceae Lomatium insulare Herb GUAAsteraceae Hazardia cana Shrub GUAAsteraceae Senecio lyonii Shrub BEN MARBoraginaceae Phacelia floribunda Herb GUACactaceae Bergerocactus emoryi Succulent MARConvolvulaceae Calystegia macrostegia Vine GUA MAR

subsp. macrostegiaCrossosomataceae Crossosoma californicum Shrub GUAEuphorbiaceae Euphorbia misera Shrub NAT CED BEN GUA MARFabaceae Acmispon argophyllus Shrub GUA

subsp. argenteusFabaceae Lupinus guadalupensis Herb GUAFabaceae Trifolium palmeri Herb GUAFagaceae Quercus tomentella Tree GUAGrossulariaceae Ribes viburnifolium Shrub CEDLamiaceae Salvia brandegeei ShrubPapaveraceae Eschscholzia ramosa Herb NAT CED BEN GUA MARPhmyraceae Mimulus latifolius Herb GUAPlantaginaceae Gambelia speciosa Shrub GUAPoaceae Dissanthelium californicum Herb GUAPolemoniaceae Gilia nevinii Herb GUAPolemoniaceae Leptosiphon pygmaeus Herb GUA

subsp. pygmaeusRhamnaceae Ceanothus arboreus Shrub GUARhamnaceae Rhamnus pirifolia Shrub GUARosaceae Prunus ilicifolia subsp. lyonii TreeSaxifragaceae Jepsonia malvifolia Herb GUAScrophulariaceae Scrophularia villosa Shrub GUASolanaceae Solanum wallacei Shrub GUANUMBER OF SPECIES 2 3 3 21 0 5aNatividad [NAT], Cedros [CED], San Benito [BEN], Guadalupe [GUA], Jeronimo [JER], San Martín [MAR], Todos Santos [TOS], Coronados [COR], San Clemente

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by analyzing unusual floristic patterns seenon the California Pacific Islands. The 26 taxashared by the Baja California and ChannelIslands indicate a clear connection betweenthese isolated and disjunct regions. What pre-cludes these taxa from occurring on the AltaCalifornia mainland, and what does this sug-gest for the past extent of the CFP? Did thesetaxa evolve lo cally and then migrate, or doesthis pattern instead reflect a distant legacy?

Three current hypotheses attempt to ex -plain the presence of these disjunct popula-tions on the California Pacific Islands: (1)these species have the ability to disperse longdistances yet for some reason have failed toestablish on the California mainland; (2) thesespecies were his torically more widespread,connected by former geologic linkages, andhave been unable to persist on the Alta Cali-fornia mainland, remaining in refugia on theislands and in Baja California; or (3) the Chan-nel Islands originally occurred farther southnear San Diego (Atwater 1998) and as they

moved north, fog-moderated climates retainedspecies native to that region.

A number of the exemplar taxa have distri-butions on the Channel Islands and farthersouth in Baja California, suggestive of therefugium hypothesis (i.e., these species wereonce more widespread). Epling and Lewis(1942) proposed that the diverse Diegan Scrubhas many taxa derived from Miocene vegeta-tion that came from the north Mexican plateau.Axelrod (1978) states that Diegan Scrub wasmore restricted in the Pleistocene and ex -panded in the Holocene as arid-adapted taxamoved northward into the southern ChannelIslands. Many CFP species are found in dis-junct occurrences, scattered across Arizonaand Mexico, not unlike the presence of these26 species on the islands (Valiente-Banuet etal. 1998, Bhaskar et al. 2007). Small-scaletopo graphic niches and facilitation by otherspecies often support these occurrences. Axel-rod (1978) also points out that the most impor-tant conditions for Diegan Scrub vegetation

2014] CALIFORNIA ISLAND PLANT DISTRIBUTION PATTERNS 165

Islands are listed left to right by increasing latitude. The Mainland column indicates continental occurrences. The protected rankings.

TOS COR Mainland CLE NIC CAT BAR ROS CRU ANA MIG # ISL CRPR BC

No CLE NIC 3 1B.2 1bNo CLE 2 1B.2 1bBaja only CLE CAT 4 2aNo CLE 2 1B.2 1b

TOS COR Baja & CA CLE CAT 5 2.2 4No CAT ROS CRU ANA MIG 7 2a

CA only CLE CAT 3 1B.2 1bTOS COR Baja & CA CLE CAT CRU 10 2.2

No CLE NIC CAT 4 2a

No CLE 2 1B.2 1bNo CLE NIC CAT BAR ANA 5 4.2 2aNo CLE CAT ROS CRU ANA 6 4.2 2aBaja & CA CAT 2 1B.2 4Baja only ROS 1 1B.2 1b

TOS COR No CLE NIC CAT BAR ROS CRU 13 4.3 2aNo CAT CRU 3 1A 1bNo CLE CAT BAR 4 1B.2 1bNo CLE CAT 3 1B.2 1aNo CLE NIC CAT BAR ROS CRU ANA 8 4.3 2aNo CLE 2 1B.2 1b

No CAT ROS CRU 4 2aNo CLE CAT ROS CRU MIG 6 4.2 2aBaja only CLE CAT ROS CRU ANA 5No CLE NIC CAT ROS CRU 6 4.2 2aNo CLE CAT 3 1B.2 1bNo CAT ROS CRU 4 1B.1 1b

3 3 20 6 20 4 10 11 5 2 21 24

[CLE], San Nicolas [NIC], Santa Catalina [CAT], Santa Barbara [BAR], Santa Rosa [ROS], Santa Cruz [CRU], Anacapa [ANA], and San Miguel [MIG].

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appear to be the absence of frost and the pres-ence of a degree of summer moisture (in theform of fog). The cold California current anddense fog banks suggest that similarities in cli-mate may help to explain the similarities inthe floras (Fischer et al. 2008).

On the other hand, a number of the exem-plar taxa are perhaps better explained as hav-ing evolved on the islands. This is especiallytrue of many island Lotus species (Acmispon;McGlaughlin et al. 2011, McGlaughlin personalcommunication), Eschscholzia ramosa (Still andPotter 2013), and Crossosoma californicum(Wallace and Helenurm 2009), among others(Baldwin, 2007). For this pattern, we hypothe-size local evolution and subsequent dispersalto similar niches on the CFP Baja Islands.

Guadalupe Island poses a particularly in -teresting study system. As the most isolatedisland, located roughly 270 km offshore, Guada -lupe is a true oceanic island never connectedto the Baja California Peninsula. Understandingthe colonization of Guadalupe may contributeto understanding the dispersal patterns thatwe see today. Of 26 exemplar taxa, 21 of themoccur on Guadalupe. Floristic similarities tothe Channel Islands flora (at least 400 kmnorth) may be due to the California currentmoving species from the California islandssouthward. Guadalupe Island has been abovewater for millions of years, and its current floracould represent the accumulation of speciesfrom the possible sources over this time.

In addition to a better understanding of theorigin and evolution of the CFP, these specieslists allow for improved conservation planning.Twenty-one of the 26 species are listed by theCalifornia Rare Plant Ranking (CNPS 2013)yet currently have no protection with the Mexi -can government. Twenty-five of the 26 speciesare listed in the rare plant ranking for BajaCalifornia (O’Brien et al. in press). These rank-ings are utilized in conservation planning inCalifornia and would be useful for land plan-ning in Baja California. These combinedislands species lists provide important infor-mation about the level of invasive species inthe re gions. Of the 535 taxa on the Baja Cali-fornia Islands, roughly 14% (74) are intro-duced species (excluding planted species,which were not included in our analyses). Incomparison, of the 976 species on the Califor-nia Channel Islands, roughly 28% (278) areintroduced species. Presumably, the lower

proportion of invasive species on the BajaCalifornia Islands is due to less human useand fewer impacts to date.

The exemplar taxa identified by this processcan now serve as the focal point of research tobetter understand the region’s historical bio-geography. Such research can incorporatephylogenetic relationships and their importancein community assembly (Webb et al. 2008).Functional trait analyses would help deter-mine if these taxonomically diverse speciesshare inherent traits that explain their disjunctdistributions. Comparative phylogeographicstudies of these shared taxa would test thespecific hypothesis discussed above, amongothers, and offer a glimpse into the past. Addi-tionally, the knowledge of patterns and modesof dispersal would provide valuable naturalhistory information. Further genetic work toresolve the origin of these island specialistspecies would help to elucidate questions oftiming, and definitively separate neo- and paleo -endemics in the exemplar list.

Conservation activity in Baja California hasrecently expanded, yet knowledge of plant dis-tributions and ecology remains limited. Wehope that these updated checklists of the Cali-fornia Pacific Islands will facilitate speciesstatus and distribution updates on both sidesof the border, and foster well-informed conser-vation decisions that promote expanded scienceand management for the transborder ecoregion.

ACKNOWLEDGMENTS

This work is based on the legacy of bo -tanists who have explored and documentedthe floristic diversity of California and thePacific Islands. In particular, we thank the fol-lowing individuals who have directly aidedthis paper. Steve Junak’s expertise has beeninvaluable; we are most grateful for his per-sonal communications and guidance. His dedi -cation and role as an information clearing-house for the island floras is vital for island-focused plant work. We are extremely gratefulto Jon Rebman of the San Diego Natural His-tory Museum (SD) for his assistance withthe Baja California Island checklist, as wellas his additional data, nomenclatural referenceworks, and taxonomic assistance. Karen Richand Judy Gibson at SD also provided informa-tion. Bart O’Brien and coauthors kindly gaveus access to a great deal of informative data

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in their in-press manuscript. Two anonymousreviewers contributed greatly to the quality ofthis manuscript.

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Received 27 April 2013Accepted 27 May 2014

Early online 25 September 2014

168 MONOGRAPHS OF THE WESTERN NORTH AMERICAN NATURALIST [Volume 7

Appendix 1 on page 169.Appendix 2 on page 188.

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2014] CALIFORNIA ISLAND PLANT DISTRIBUTION PATTERNS 169A

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CO

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ativ

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]; Sa

n B

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n M

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d To

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170 MONOGRAPHS OF THE WESTERN NORTH AMERICAN NATURALIST [Volume 7A

PPE

ND

IX1.

Con

tinue

d

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ily/s

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reen

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ngel

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ichx

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ees

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pect

inac

eaN

AT

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uca

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onym

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.) G

ray

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2014] CALIFORNIA ISLAND PLANT DISTRIBUTION PATTERNS 171A

PPE

ND

IX1.

Con

tinue

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ily/s

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mel

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nym

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rin.

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kwor

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nym

: Nas

sella

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itchc

.) B

arkw

orth

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172 MONOGRAPHS OF THE WESTERN NORTH AMERICAN NATURALIST [Volume 7A

PPE

ND

IX1.

Con

tinue

d

Fam

ily/s

peci

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p./v

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ayne

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2014] CALIFORNIA ISLAND PLANT DISTRIBUTION PATTERNS 173A

PPE

ND

IX1.

Con

tinue

d

Fam

ily/s

peci

esss

p./v

ar. a

nd in

fran

ame

CO

RT

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MA

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emis

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char

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174 MONOGRAPHS OF THE WESTERN NORTH AMERICAN NATURALIST [Volume 7A

PPE

ND

IX1.

Con

tinue

d

Fam

ily/s

peci

esss

p./v

ar. a

nd in

fran

ame

CO

RT

OS

MA

RJE

RG

UA

BE

NC

ED

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zies

ii (H

ook.

& A

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var.

men

zies

iiT

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CE

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sian

a A

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yC

ED

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then

ia c

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aria

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then

ia g

raci

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ayia

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tygl

ossa

(Fis

ch. &

Mey

.) A

. Gra

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ne g

igan

tea

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logg

26G

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(Nut

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y27

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RT

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MA

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. Gra

y) H

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rica

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ene3

0 *G

UA

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lifor

nicu

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sch.

& C

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ey.

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a B

enth

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EN

Peri

tyle

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oryi

Tor

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RT

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MA

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UA

BE

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ED

NA

TPe

rity

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cana

A. G

ray

GU

APl

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a od

orat

a (L

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ass.

var.

odor

ata

CE

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roph

yllu

m g

raci

le B

enth

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lium

ben

eole

ns D

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TO

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2014] CALIFORNIA ISLAND PLANT DISTRIBUTION PATTERNS 175A

PPE

ND

IX1.

Con

tinue

d

Fam

ily/s

peci

esss

p./v

ar. a

nd in

fran

ame

CO

RT

OS

MA

RJE

RG

UA

BE

NC

ED

NA

T

Sene

cio

lyon

ii A

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30)

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176 MONOGRAPHS OF THE WESTERN NORTH AMERICAN NATURALIST [Volume 7A

PPE

ND

IX1.

Con

tinue

d

Fam

ily/s

peci

esss

p./v

ar. a

nd in

fran

ame

CO

RT

OS

MA

RJE

RG

UA

BE

NC

ED

NA

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cura

inia

pin

nata

ssp.

gla

bra

(Woo

t. &

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ndl.)

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l.C

OR

CE

DD

escu

rain

ia p

inna

ta (C

ocke

rell)

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ling

ssp.

hal

icto

rum

(Coc

kere

ll) D

etlin

gM

AR

CE

DD

escu

rain

ia p

inna

ta (C

ocke

rell)

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ling

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men

zies

ii (D

C.)

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ling

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UA

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ba c

unei

folia

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atso

nva

r. in

tegr

ifolia

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DE

ruca

ves

icar

ia (L

.) C

av.3

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p. s

ativ

aC

ED

Ery

sim

um m

oran

ii R

ollin

sG

UA

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nung

ia p

rocu

mbe

ns (L

.) H

ayek

38*

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AL

epid

ium

lasi

ocar

pum

Nut

t.va

r. la

tifol

ium

MA

R?

GU

AB

EN

?N

AT

Lep

idiu

m n

itidu

m T

orre

y &

A. G

ray

TO

SG

UA

Lep

idiu

m o

blon

gum

Sm

all

var.

insu

lare

C.L

.Hitc

hc.

TO

SM

AR

GU

AB

EN

CE

DN

AT

Rap

hanu

s sa

tivus

L.*

GU

ASi

bara

ang

elor

um (S

. Wat

son)

Gre

ene

CE

DSi

sym

briu

m ir

io L

.*C

OR

TO

SG

UA

CE

DN

AT

Sisy

mbr

ium

ori

enta

le L

.*T

OS

GU

AC

ED

Thy

sano

carp

us e

rect

us S

. Wat

son

GU

AC

ED

Thy

sano

carp

us la

cini

atus

Nut

tall

CE

D

35N

ew r

ecor

d; h

erba

rium

spe

cim

en @

RSA

, SD

36Sy

nony

m: G

uille

nia

lasi

ophy

lla(H

ook.

& A

rn.)

Gre

ene

37N

ew r

ecor

d; h

erba

rium

spe

cim

en @

SD

(Joh

n B

row

n, s

.n.,

Apr

il 4

1983

)38

Syno

nym

: Hut

chin

sia

proc

umbe

ns(L

.) D

esv.

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2014] CALIFORNIA ISLAND PLANT DISTRIBUTION PATTERNS 177A

PPE

ND

IX1.

Con

tinue

d

Fam

ily/s

peci

esss

p./v

ar. a

nd in

fran

ame

CO

RT

OS

MA

RJE

RG

UA

BE

NC

ED

NA

T

Cac

tace

aeB

erge

roca

ctus

em

oryi

(Eng

elm

.) B

r. &

R.

CO

RT

OS

MA

RC

oche

mie

a po

ndii

Wal

ton

CE

DN

AT

Cyl

indr

opun

tia a

lcah

es (F

.A.C

. Web

er) F

.M. K

nuth

var.

alca

hes

CE

DC

ylin

drop

untia

cho

lla (F

.A.C

. Web

er) F

.M. K

nuth

NA

TC

ylin

drop

untia

pro

lifer

a (E

ngel

m.)

F.M

.Knu

thC

OR

TO

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AR

? G

UA

CE

D?

Cyl

indr

opun

tia s

p. n

ova3

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EN

CE

DN

AT

?E

chin

ocer

eus

mar

itim

us (M

.E. J

ones

) K. S

chum

.va

r. m

ariti

mus

MA

RC

ED

?N

AT

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cact

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thus

(Orc

utt)

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tt. &

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EN

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ED

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cact

us fo

rdii

(Orc

utt)

Bri

tt. &

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r. gr

andi

floru

s G

.E. L

inds

.M

AR

NA

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opho

cere

us s

chot

tii (E

ngel

m.)

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& R

.va

r. sc

hott

iiM

AR

40B

EN

CE

DM

amm

illar

ia b

loss

feld

iana

Boe

deke

rva

r. sh

urlia

na G

ates

GU

AC

ED

Mam

mill

aria

dio

ica

M.K

. Bra

ndeg

eeC

OR

TO

SM

AR

Mam

mill

aria

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drid

gei S

chee

rva

r. r

ectis

pina

E.Y

. Daw

son

CE

DM

amm

illar

ia g

oodr

idge

i Sch

eer

var.

good

ridg

eiC

ED

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mill

aria

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chis

onia

na (G

ates

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d.C

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amm

illar

ia lo

uisa

e G

.E. L

inds

MA

R41

Mam

mill

aria

neo

palm

eri R

.T. C

raig

BE

N?

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cact

us c

ocha

l (O

rcut

t) B

ritt

. & R

ose

MA

RO

punt

ia fi

cus-

indi

ca (L

.) M

iller

TO

SM

AR

Opu

ntia

litt

oral

is (E

ngel

m.)

Coc

kere

llC

OR

TO

SM

AR

42 a

ff.O

punt

ia o

rico

la P

hilb

rick

CO

RT

OS

MA

R43

CE

D?

Pach

ycer

eus

prin

glei

(S. W

atso

n) B

ritt

on &

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eC

ED

?N

AT

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ocer

eus

gum

mos

us (E

ngel

m.)

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. M

AR

CE

DG

ibso

n &

K.E

. Hor

akC

ampa

nula

ceae

Gith

opsi

s di

ffusa

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ray

var.

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s (M

orin

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mer

sG

UA

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dani

s bi

flora

(Rui

z &

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.) G

reen

eG

UA

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naba

ceae

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tis la

evig

ata4

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r. re

ticul

ata

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enso

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appa

race

aePe

rito

ma

arbo

rea

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t.) I

ltis4

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gust

ata

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ish)

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sC

ED

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tom

a ar

bore

a (N

utt.)

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s46

var.

glob

osa

(Cov

ille)

Ilti

sC

OR

39N

ew r

ecor

d; p

ers.

com

m. J

on R

ebm

an, S

D (i

nclu

des

Cyl

indr

opun

tia c

edro

sens

isR

ebm

an)

40N

ew r

ecor

d; h

erba

rium

spe

cim

en @

RSA

, SD

41R

e-id

entif

icat

ion,

cha

nged

from

M. h

utch

inso

nia

42N

ew r

ecor

d; h

erba

rium

spe

cim

en @

RSA

43N

ew r

ecor

d; h

erba

rium

spe

cim

en @

RSA

44Sy

nony

m: C

eltis

dou

glas

iiPl

anch

.45

Syno

nym

: Iso

mer

is a

rbor

eaN

utt.

ssp.

ang

usta

taPa

rish

46N

ew r

ecor

d; h

erba

rium

spe

cim

en @

SD

Reb

man

623

0

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178 MONOGRAPHS OF THE WESTERN NORTH AMERICAN NATURALIST [Volume 7A

PPE

ND

IX1.

Con

tinue

d

Fam

ily/s

peci

esss

p./v

ar. a

nd in

fran

ame

CO

RT

OS

MA

RJE

RG

UA

BE

NC

ED

NA

T

Cap

rifo

liace

aeL

onic

era

hisp

idul

a D

ougl

.47

GU

AL

onic

era

subs

pica

ta H

ook.

& A

rn.

var.

denu

data

Reh

der

CE

DC

aryo

phyl

lace

aeA

chyr

onyc

hia

coop

eri T

orre

y &

A. G

ray

CE

DC

eras

tium

glo

mer

atum

Thu

ill.*

GU

AD

rym

aria

hol

oste

oide

s B

enth

.va

r. ho

lost

eoid

esC

ED

Her

niar

ia h

irsu

ta L

.va

r. ci

nere

a (D

C.)

Cou

tinho

GU

APo

lyca

rpon

dep

ress

um N

utt.

TO

SC

ED

Sile

ne a

ntir

rhin

a L

.T

OS

GU

ASi

lene

gal

lica

L.*

CO

RT

OS

GU

AC

ED

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lene

laci

niat

a C

av.4

8ss

p. la

cini

ata

CO

RSp

ergu

lari

a bo

ccon

ii (S

chee

le) M

erin

o*G

UA

Sper

gula

ria

mac

roth

eca

var.

leuc

anth

a (G

reen

e) B

.L. R

ob.

GU

A(H

orne

m. e

x C

ham

. & S

chltd

l.) H

eynh

.49

Sper

gula

ria

mac

roth

eca

var.

talin

um (G

reen

e) J

eps.

GU

A(H

orne

m. e

x C

ham

. & S

chltd

l.) H

eynh

.Sp

ergu

lari

a m

arin

a J.&

K. P

resl

MA

R50

GU

ASp

ergu

lari

a vi

llosa

(Per

s.) C

amb.

*C

OR

Stel

lari

a m

edia

(L.)

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*C

OR

Stel

lari

a pa

llida

(Dum

ort.)

Cré

p*

CO

RSt

ella

ria

nite

ns N

utt.

GU

AA

llenr

olfe

a oc

cide

ntal

is (S

. Wat

son)

Kun

tze

CE

DC

heno

podi

acea

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phan

ism

a bl

itoid

es N

utt.

CO

RT

OS

MA

RG

UA

BE

NC

ED

NA

TA

rthr

ocne

mum

sub

term

inal

e (P

aris

h) S

tand

l.51

MA

RA

trip

lex

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laya

na (B

enth

.) D

ietr.

GU

AB

EN

CE

D?

NA

TA

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lex

calif

orni

ca M

oq.

CO

RT

OS

GU

AC

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iple

x ca

nesc

ens

(Pur

sh) N

utt.

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can

esce

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ulte

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oq.)

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ietr.

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NC

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?N

AT

?A

trip

lex

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cea

S. W

atso

nT

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MA

RC

ED

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lex

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ophy

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n D

C.)

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ietr.

MA

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ED

NA

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trip

lex

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fica

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s.C

OR

MA

RC

ED

NA

TA

trip

lex

sem

ibac

cata

R. B

r.*C

OR

TO

SM

AR

GU

AC

ED

Atr

iple

x se

rena

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. Nel

son

ex A

bram

sva

r. da

vids

onii

(Sta

ndl.)

Mun

zC

OR

MA

R52

aff.

NA

T ×

aff.

47Sy

nony

m: L

onic

era

hisp

idul

a(L

indl

.) D

ougl

as e

x To

rr. &

A. G

ray

var.

vaci

llans

A. G

ray

48Sy

nony

m: S

ilene

laci

niat

aC

av. s

sp. m

ajor

C.L

. Hitc

h. &

Mag

uire

49N

ew r

ecor

d; h

erba

rium

spe

cim

en @

RSA

, SD

50N

ew r

ecor

d; h

erba

rium

spe

cim

en @

RSA

, SB

BG

51Sy

nony

m: S

alic

orni

a su

bter

min

alis

Pari

sh52

New

rec

ord;

her

bari

um s

peci

men

@ R

SA, p

oten

tially

new

spe

cies

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2014] CALIFORNIA ISLAND PLANT DISTRIBUTION PATTERNS 179A

PPE

ND

IX1.

Con

tinue

d

Fam

ily/s

peci

esss

p./v

ar. a

nd in

fran

ame

CO

RT

OS

MA

RJE

RG

UA

BE

NC

ED

NA

T

Atr

iple

x su

bere

cta

Verd

oorn

CO

R?5

3G

UA

Atr

iple

x w

atso

nii A

. Nel

son

MA

RC

heno

podi

um a

lbum

L.

CE

D?

Che

nopo

dium

cal

iforn

icum

(S. W

atso

n) S

. Wat

son

CO

RT

OS

Che

nopo

dium

flab

ellif

oliu

m S

tand

l.M

AR

Che

nopo

dium

mur

ale

L.*

CO

RT

OS

MA

RJE

RG

UA

BE

NC

ED

NA

TSa

licor

nia

paci

fica

Stan

dl.5

4M

AR

Sals

ola

aust

ralis

R.B

r.*C

OR

TO

SC

ED

Suae

da n

igra

(Raf

ines

que)

J.F.

Mac

brid

eB

EN

NA

TSu

aeda

taxi

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ndl.)

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ndl.

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AR

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55G

UA

BE

NC

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volv

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rost

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ene)

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mm

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mac

rost

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(Gre

ene)

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mm

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term

edia

CO

R56

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yste

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mac

rost

egia

(Gre

ene)

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mm

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p. m

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steg

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GU

AC

uscu

ta c

alifo

rnic

a H

ook.

& A

rn.

MA

RN

AT

Cus

cuta

cor

ymbo

sa R

uiz

& P

avon

GU

AC

uscu

ta s

alin

a E

ngel

m.5

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p. s

alin

aB

EN

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hond

ra o

ccid

enta

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ouse

CO

RT

OS

Cra

ssul

acea

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rass

ula

conn

ata

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z L

opez

& P

avon

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OR

TO

SM

AR

GU

AB

EN

?C

ED

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erge

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udle

ya a

cum

inat

a R

ose

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DD

udle

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idso

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oran

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. cul

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udle

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oran

CO

RT

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Dud

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.A. J

ohan

s.T

OS

Dud

leya

can

dida

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tton

CO

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udle

ya c

edro

sens

is M

oran

CE

DD

udle

ya c

ultr

ata

Ros

eM

AR

Dud

leya

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dalu

pens

is M

oran

GU

AD

udle

ya in

gens

Ros

eC

ED

? af

fD

udle

ya la

nceo

lata

(Nut

t.) B

ritt

on &

Ros

eC

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Dud

leya

line

aris

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ene)

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tt. &

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eB

EN

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udle

ya p

achy

phyt

um M

oran

& B

ened

ict

CE

D

53N

ew r

ecor

d; p

ers.

com

m. J

on R

ebm

an54

Syno

nym

: Sal

icor

nia

virg

inic

aL

.55

New

rec

ord;

her

bari

um s

peci

men

@ S

D56

New

rec

ord;

per

s. c

omm

. Jon

Reb

man

57N

ew r

ecor

d; h

erba

rium

spe

cim

en @

RSA

, SD

Mor

an 2

0315

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180 MONOGRAPHS OF THE WESTERN NORTH AMERICAN NATURALIST [Volume 7A

PPE

ND

IX1.

Con

tinue

d

Fam

ily/s

peci

esss

p./v

ar. a

nd in

fran

ame

CO

RT

OS

MA

RJE

RG

UA

BE

NC

ED

NA

T

Dud

leya

vir

ens

(Ros

e) M

oran

ssp.

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anG

UA

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leya

×se

mite

res

(Ros

e) M

oran

CO

RT

OS

Cro

ssos

omat

acea

eC

ross

osom

a ca

lifor

nicu

m N

utt.

GU

AC

ucur

bita

ceae

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inop

epon

min

imus

(Kel

l.) S

. Wat

son

var.

min

imus

CE

DN

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ah g

uada

lupe

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Gre

ene

GU

AM

arah

mac

roca

rpus

(Gre

ene)

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s sp

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(Mill

sp.)

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upho

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. & A

. Gra

y*C

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?E

upho

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bar

tolo

mae

i Gre

ene

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horb

ia c

renu

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elm

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OS

Eup

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iser

a B

enth

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el) B

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llet6

1C

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pon

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diflo

rus

(Ben

th.)

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uille

t62

var.

gran

diflo

rus

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cmis

pon

mar

itim

us (N

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olof

f63

ssp.

bre

vive

xillu

s (O

ttle

y) B

roui

llet

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NC

ED

NA

TA

cmis

pon

nuda

tus

(Gre

ene)

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uille

t64

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DA

cmis

pon

rigi

dus

(Ben

th.)

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uille

t65

CE

DA

cmis

pon

stri

gosu

s (N

utt.)

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uille

t66

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SM

AR

CE

DA

stra

galu

s af

f.ga

mbe

lianu

s E

. She

ld.

NA

TA

stra

galu

s fa

stid

ius

(Kel

l.) M

.E. J

ones

CE

DA

stra

galu

s in

sula

ris

Kel

l.va

r. in

sula

ris

CE

D

58Sy

nony

m: C

ham

aesy

ce p

ondi

i(M

illsp

.) M

illsp

.59

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nym

& p

er J

on R

ebm

an m

ay b

e re

cogn

ized

as:

Lot

us a

rgop

hyllu

s(A

. Gra

y) G

reen

e ss

p. o

rnith

opus

(Gre

ene)

P.H

. Rav

en60

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nym

: Lot

us c

edro

sens

isG

reen

e

61Sy

nony

m: L

otus

sco

pari

us(T

orr.

& A

. Gra

y) O

ttle

y62

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nym

: Lot

us g

rand

iflor

us(B

enth

.) G

reen

e63

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nym

: Lot

us s

alsu

gino

sus

Gre

ene

ssp.

bre

vive

xillu

sO

ttle

y64

Syno

nym

: Lot

us n

udat

us(G

reen

e) G

reen

e65

Syno

nym

: Lot

us r

igid

us(B

enth

.) G

reen

e66

Syno

nym

: Lot

us s

trig

osus

(Nut

t.) G

reen

e

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2014] CALIFORNIA ISLAND PLANT DISTRIBUTION PATTERNS 181A

PPE

ND

IX1.

Con

tinue

d

Fam

ily/s

peci

esss

p./v

ar. a

nd in

fran

ame

CO

RT

OS

MA

RJE

RG

UA

BE

NC

ED

NA

T

Ast

raga

lus

mag

dale

nae

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ene

var.

mag

dale

nae

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stra

galu

s nu

ttal

lianu

s D

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var.

cedr

osen

sis

M.E

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stra

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icho

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s A

. Gra

yva

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s (M

.E. J

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) Bar

neby

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ea m

ollis

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th.

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onfin

is (S

tand

.) N

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rand

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upin

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son

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ene)

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ithM

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upin

us s

uccu

lent

us D

ougl

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inus

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ook.

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edic

ago

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mor

pha

L.*

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ilotu

s in

dicu

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aseo

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th.

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nii (

Vase

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Ros

e) B

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68T

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epau

pera

tum

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v.

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e) M

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? 69

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ntum

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r. &

A. G

ray

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lium

mic

roce

phal

um P

ursh

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ifoliu

m p

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eri S

. Wat

son

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i Spr

enge

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sei S

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.) W

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keni

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ryac

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ithec

ello

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con

fine

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dl.

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otus

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sey

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ose)

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ene

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ew r

ecor

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com

m. J

on R

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rec

ord;

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bari

um s

peci

men

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D71

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rec

ord;

per

s. c

omm

. Jon

Reb

man

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r @

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; syn

onym

: Aca

cia

farn

esia

nava

r. m

inut

a72

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nym

: Cen

taur

ium

ven

ustu

m(A

. Gra

y) B

.L. R

ob.

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182 MONOGRAPHS OF THE WESTERN NORTH AMERICAN NATURALIST [Volume 7A

PPE

ND

IX1.

Con

tinue

d

Fam

ily/s

peci

esss

p./v

ar. a

nd in

fran

ame

CO

RT

OS

MA

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.) L’

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orda

ta K

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ene

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vace

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alifo

rnic

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alch

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ray)

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ene

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Mal

va a

ssur

gent

iflor

a (K

ello

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.F. R

ay74

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ssur

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iflor

a T

OS

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va li

ndsa

yi (M

oran

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. Ray

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va p

acifi

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eral

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an75

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ater

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2014] CALIFORNIA ISLAND PLANT DISTRIBUTION PATTERNS 183A

PPE

ND

IX1.

Con

tinue

d

Fam

ily/s

peci

esss

p./v

ar. a

nd in

fran

ame

CO

RT

OS

MA

RJE

RG

UA

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ED

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tiace

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ia c

iliat

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uiz

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ez &

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on) D

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ista

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: Cam

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aven

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184 MONOGRAPHS OF THE WESTERN NORTH AMERICAN NATURALIST [Volume 7A

PPE

ND

IX1.

Con

tinue

d

Fam

ily/s

peci

esss

p./v

ar. a

nd in

fran

ame

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RT

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nthu

s di

anth

iflor

us (B

enth

.) G

reen

eC

OR

86Sy

nony

m: S

tylo

mec

on h

eter

ophy

lla(B

enth

.) G

. Tay

lor

87Sy

nony

m: M

imul

us a

uran

tiacu

sC

urtis

var

. aur

antia

cus

88Sy

nony

m: M

imul

us la

tifol

ius

A. G

ray,

Dip

lacu

s la

tifol

ius

(A. G

ray)

G.L

. Nes

om (D

ougl

as e

x B

enth

.) Sp

ach

89Sy

nony

m: M

imul

us c

ardi

nalis

Dou

glas

ex

Ben

th.

90Sy

nony

m: G

alve

zia

junc

ea(B

enth

.) B

all

91Sy

nony

m: G

alve

zia

spec

iosa

(Nut

t.) A

. Gra

y

92Sy

nony

ms:

Lin

aria

texa

naSc

heel

e an

d Li

nari

a ca

nade

nsis

(L.)

Dum

. Cou

rs.

93Sy

nony

m: L

inan

thus

pyg

mae

us(B

rand

) J.T

. How

ell s

sp. p

ygm

aeus

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Page 26: ,V OD Q G 6 S H F LD OLV WV 6 K D UH G ) OR UD R I ......San Martín—Junak and Philbrick 1994b Guadalupe—Moran 1996, Rebman 2006, Rebman et al. 2007 San Benito—Junak and Philbrick

2014] CALIFORNIA ISLAND PLANT DISTRIBUTION PATTERNS 185A

PPE

ND

IX1.

Con

tinue

d

Fam

ily/s

peci

esss

p./v

ar. a

nd in

fran

ame

CO

RT

OS

MA

RJE

RG

UA

BE

NC

ED

NA

T

Lina

nthu

s un

cial

is (B

rand

egee

) Mor

anC

ED

?Li

nant

hus

veat

chii

(C. P

arry

) J.M

. C

ED

Port

er &

L.A

. Joh

nson

Mic

rost

eris

gra

cilis

(Hoo

k.) G

reen

eC

ED

?Po

lygo

nace

aeE

riog

onum

fasc

icul

atum

Ben

th.

CO

RT

OS

CE

DE

riog

onum

gra

nde

Gre

ene

var.

test

udin

um R

evea

lT

OS

Eri

ogon

um in

tric

atum

Ben

th.

CE

DE

riog

onum

mol

le G

reen

eC

ED

Eri

ogon

um p

ondi

i Gre

ene

Torr

ey e

x B

enth

.C

ED

NA

TE

riog

onum

zap

atoe

nse

Mor

anG

UA

Eri

ogon

um w

righ

tiiva

r. ta

xifo

lium

(Gre

ene)

Par

ish

CE

DH

arfo

rdia

mac

ropt

era

(Ben

th.)

var.

frut

icos

a (G

reen

e) R

evea

lC

ED

Gre

ene

& P

arry

Las

tarr

iaea

cor

iace

a (G

oodm

an) H

oove

r94

CE

D?

Pter

oste

gia

drym

ario

ides

F. &

M.

CO

RT

OS

MA

RG

UA

CE

DPo

rtul

acac

eae

Port

ulac

a ol

erac

ea L

.*N

AT

Prim

ulac

eae

Dod

ecat

heon

cle

vela

ndii

Gre

ene

ssp.

insu

lare

(H.J.

Tho

mps

.) R

evea

lG

UA

Ran

uncu

lace

aeC

lem

atis

pau

ciflo

ra N

utt.

CO

RT

OS

CE

D?

Del

phin

ium

car

dina

le H

ook.

CE

D?

Del

phin

ium

par

ryi A

. Gra

yC

OR

TO

SM

yosu

rus

min

imus

L.

GU

AR

anun

culu

s he

beca

rpus

H. &

A.

GU

AC

ED

?R

esed

acea

eO

ligom

eris

lini

folia

(Vah

l.) M

acbr

.C

OR

TO

SG

UA

BE

NC

ED

NA

TR

ham

nace

aeC

eano

thus

arb

oreu

s G

reen

eG

UA

Cea

noth

us c

rass

ifoliu

s To

rr.

GU

AC

eano

thus

cun

eatu

s (H

ook.

) Nut

t.G

UA

Cea

noth

uspe

rple

xans

Tre

l.95

GU

AC

eano

thus

ver

ruco

sus

Nut

t.C

ED

?R

ham

nus

insu

la K

ello

ggC

ED

?R

ham

nus

piri

folia

Gre

ene

GU

AZ

izip

hus

parr

yi T

orr.

ssp.

mic

roph

ylla

CE

D?

Ros

acea

eA

deno

stom

a fa

scic

ulat

um H

ook.

& A

rn.

var.

obt

usifo

lium

S. W

ats.

CE

D?

94Sy

nony

m: C

hori

zant

he c

oria

cea

Goo

dman

95N

ew r

ecor

d; h

erba

rium

spe

cim

en @

RSA

, SD

, SB

BG

; syn

onym

: Cea

noth

us g

regg

iiA

. Gra

y va

r. pe

rple

xans

(Tre

l.) J

eps.

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Page 27: ,V OD Q G 6 S H F LD OLV WV 6 K D UH G ) OR UD R I ......San Martín—Junak and Philbrick 1994b Guadalupe—Moran 1996, Rebman 2006, Rebman et al. 2007 San Benito—Junak and Philbrick

186 MONOGRAPHS OF THE WESTERN NORTH AMERICAN NATURALIST [Volume 7A

PPE

ND

IX1.

Con

tinue

d

Fam

ily/s

peci

esss

p./v

ar. a

nd in

fran

ame

CO

RT

OS

MA

RJE

RG

UA

BE

NC

ED

NA

T

Aph

anes

occ

iden

talis

Ryd

berg

GU

AH

eter

omel

es a

rbut

ifolia

(Lin

dl.)

M. R

oem

.C

OR

TO

SG

UA

CE

D?

Rub

iace

aeG

aliu

m a

ngul

osum

A. G

ray

GU

AG

aliu

m a

ngus

tifol

ium

Nut

t.ss

p. a

ngus

tifol

ium

CO

RG

aliu

m a

pari

ne L

.C

OR

96M

AR

GU

AC

ED

?G

aliu

m c

oron

adoe

nse

Dem

pste

rC

OR

Gal

ium

ste

llatu

m K

ello

ggva

r. er

emic

um H

ilend

& J.

How

ell

CE

DR

utac

eae

Rut

a ch

alep

ensi

s L

.G

UA

Sapi

ndac

eae

Aes

culu

s pa

rryi

A. G

ray

CO

RSa

xifr

agac

eae

Jeps

onia

mal

vifo

lia (G

reen

e) S

mal

lG

UA

Saxi

frag

acea

eJe

pson

ia p

arry

i (To

rr.)

Smal

l.C

OR

Scro

phul

aria

ceae

Myo

poru

m la

etum

For

st. f

. Nga

io.*

CO

RSc

roph

ular

ia v

illos

a Pe

nnel

lG

UA

Sim

mon

dsia

ceae

Sim

mon

dsia

chi

nens

is (L

ink)

C. S

chne

ider

GU

AC

ED

NA

TSo

lana

ceae

Dat

ura

disc

olor

Ber

nh.

NA

TD

atur

a w

righ

tii R

egel

MA

RB

EN

?L

yciu

m a

nder

soni

i A. G

ray

MA

RC

ED

?L

yciu

m b

revi

pes

Ben

th.

var.

brev

ipes

TO

SM

AR

JER

CE

D?

NA

TL

yciu

m c

alifo

rnic

um N

utt.

CO

RT

OS

MA

RG

UA

BE

N?

NA

TL

yciu

m e

xser

tum

A. G

ray

BE

NC

ED

?L

yciu

m fr

emon

tii A

. Gra

yG

UA

Lyc

oper

sico

n es

cule

ntum

L.*

CO

RT

OS

Nic

otia

na a

tten

uata

Tor

r.G

UA

Nic

otia

na c

leve

land

ii A

. Gra

yC

OR

TO

SM

AR

CE

DN

AT

Nic

otia

na g

lauc

a G

rah.

*G

UA

Phys

alis

cra

ssifo

lia B

enth

.va

r. cr

assi

folia

TO

SC

ED

Sola

num

am

eric

anum

Mill

er*

TO

SM

AR

GU

ASo

lanu

m d

ougl

asii

Dun

alG

UA

Sola

num

hin

dsia

num

Ben

th.

CE

DSo

lanu

m n

odifl

orum

Jac

q.*

CO

R

96N

ew r

ecor

d; h

erba

rium

spe

cim

en @

SD

.

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Page 28: ,V OD Q G 6 S H F LD OLV WV 6 K D UH G ) OR UD R I ......San Martín—Junak and Philbrick 1994b Guadalupe—Moran 1996, Rebman 2006, Rebman et al. 2007 San Benito—Junak and Philbrick

2014] CALIFORNIA ISLAND PLANT DISTRIBUTION PATTERNS 187A

PPE

ND

IX1.

Con

tinue

d

Fam

ily/s

peci

esss

p./v

ar. a

nd in

fran

ame

CO

RT

OS

MA

RJE

RG

UA

BE

NC

ED

NA

T

Sola

num

pal

mer

i Vas

ey &

Ros

eT

OS

MA

RSo

lanu

m w

alla

cei (

A. G

ray)

Par

ish

ssp.

clo

keyi

GU

ATa

mar

icac

eae

Tam

arix

ram

osis

sim

a L

edeb

.*C

ED

Urt

icac

eae

Hes

pero

cnid

e te

nella

Tor

rey

TO

SM

AR

GU

APa

riet

aria

hes

pera

B.D

. Hin

ton

CO

RT

OS

GU

AC

ED

?U

rtic

a ur

ens

L.*

CO

RVe

rben

acea

eG

land

ular

ia li

laci

na (G

reen

e) U

mbe

r.97

CE

DVi

scac

eae

Phor

aden

dron

den

sum

Tor

r. ex

Tre

l.G

UA

Zyg

ophy

llace

aeFa

goni

a la

evis

Sta

ndl.

CE

DN

AT

Vis

cain

oa g

enic

ulat

a (K

ello

gg) G

reen

eva

r. ge

nicu

lata

CE

DN

UM

BE

RO

FSP

EC

IES

129

145

107

823

257

266

79

97Sy

nony

m: V

erbe

na li

laci

naG

reen

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188 MONOGRAPHS OF THE WESTERN NORTH AMERICAN NATURALIST [Volume 7A

PPE

ND

IX2.

Pla

nt s

peci

es c

heck

list f

or th

e 8

Cal

iforn

ia C

hann

el I

slan

ds (A

naca

pa [A

NA

], Sa

n C

lem

ente

[CL

E],

San

Mig

uel [

MIG

], Sa

n N

icol

as [N

IC],

Sant

a B

arba

ra [B

AR

], Sa

nta

Cat

alin

a [C

AT

], Sa

nta

Cru

z [C

RU

], an

d Sa

nta

Ros

a [R

OS]

). A

n am

pers

and

(&) i

ndic

ates

an

extir

pate

d sp

ecie

s; a

que

stio

n m

ark

(?) i

ndic

ates

an

unvo

uche

red

taxa

; and

an

aste

risk

(*)

indi

cate

s a

nonn

ativ

e sp

ecie

s.

Fam

ily/s

peci

esss

p./v

ar. a

nd in

fran

ame

CL

EN

ICC

AT

BA

RR

OS

CR

UA

NA

MIG

FE

RN

S

Azo

llace

ae

Azo

lla fi

licul

oide

s L

am.

CA

TB

lech

nace

ae

Woo

dwar

dia

fimbr

iata

Sm

.C

LE

? 1C

AT

2R

OS

CR

UD

enns

taed

tiace

ae

Pter

idiu

m a

quili

num

(L.)

Kuh

nva

r. pu

besc

ens

Und

erw

.C

AT

RO

SC

RU

Dry

opte

rida

ceae

D

ryop

teri

s ar

guta

(Kau

lf.) M

axon

CL

EC

AT

RO

SC

RU

AN

AM

IGPo

lyst

ichu

m m

unitu

m (K

aulf.

) C. P

resl

RO

SC

RU

Equ

iset

acea

e E

quis

etum

hye

mal

e L

.ss

p. a

ffine

(Eng

elm

.) C

alde

r R

OS

CR

U&

Roy

L. T

aylo

rE

quis

etum

laev

igat

um A

. Bra

unC

AT

RO

SC

RU

Equ

iset

um te

lmat

eia

Ehr

h.ss

p. b

raun

ii (J

. Mild

e) H

auke

CA

TC

RU

Poly

podi

acea

e Po

lypo

dium

cal

iforn

icum

Kau

lf.C

LE

NIC

CA

TB

AR

RO

SC

RU

AN

APo

lypo

dium

sco

uler

i Hoo

k. &

Gre

v.C

RU

Pter

idac

eae

Adi

antu

m a

leut

icum

(Rup

r.) C

.A. P

aris

CR

UA

dian

tum

cap

illus

-ven

eris

L.

CA

TR

OS

CR

UA

NA

Adi

antu

m jo

rdan

ii M

üll.

Hal

.C

LE

NIC

CA

TR

OS

CR

UA

NA

MIG

Asp

idot

is c

alifo

rnic

a (H

ook.

) Cop

el.

CA

TC

RU

Che

ilant

hes

clev

elan

dii D

.C. E

aton

RO

SC

RU

Che

ilant

hes

coop

erae

D.C

. Eat

onC

RU

Che

ilant

hes

new

berr

yi(D

.C. E

aton

) Dom

inC

LE

Pella

ea a

ndro

med

ifolia

(Kau

lf.) F

éeC

LE

NIC

CA

TR

OS

CR

UA

NA

MIG

Pella

ea m

ucro

nata

(D.C

. Eat

on)

var.

muc

rona

taC

LE

? 3C

AT

RO

SC

RU

D.C

. Eat

onPe

ntag

ram

ma

tria

ngul

aris

(Kau

lf.) Y

atsk

., ss

p. tr

iang

ular

isC

LE

NIC

CA

TR

OS

CR

UA

NA

MIG

Win

dham

, & E

. Wol

lenw

.Pe

ntag

ram

ma

tria

ngul

aris

(Kau

lf.) Y

atsk

., ss

p. v

isco

sa (D

.C. E

aton

) Yat

sk.e

t al.

CL

EC

AT

RO

SC

RU

Win

dham

, & E

. Wol

lenw

.Se

lagi

nella

ceae

Se

lagi

nella

big

elov

ii U

nder

w.

CL

EC

AT

RO

SC

RU

AN

A

1 Per

s. c

om. E

. How

e2 S

. Rat

ay c

olle

ctio

n 3 P

ers.

com

. E. H

owe

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2014] CALIFORNIA ISLAND PLANT DISTRIBUTION PATTERNS 189A

PPE

ND

IX2.

Con

tinue

d

Fam

ily/s

peci

esss

p./v

ar. a

nd in

fran

ame

CL

EN

ICC

AT

BA

RR

OS

CR

UA

NA

MIG

Woo

dsia

ceae

Ath

yriu

m fi

lix-f

emin

a (L

.) R

oth

var.

cycl

osor

um R

upr.

RO

SC

RU

Cys

topt

eris

frag

ilis

(L.)

Ber

nh.

CR

UG

YMN

OSP

ER

MS

Cup

ress

acea

e H

espe

rocy

pari

s m

acro

carp

a (H

artw

.) B

arte

l*C

AT

RO

SC

RU

AN

APi

nace

ae

Pinu

s m

uric

ata

D. D

onva

r. m

uric

ata

RO

SC

RU

Pinu

s pi

nea

L.*

CR

UPi

nus

radi

ata

D. D

on*

MIG

Pinu

s to

rrey

ana

Parr

y ex

Car

rièr

ess

p. in

sula

ris

J.R. H

alle

rR

OS

MO

NO

CO

TS

Aga

vace

ae

Aga

ve a

mer

ican

a L

.*C

RU

Aga

vace

ae

Chl

orog

alum

pom

erid

ianu

m (D

C.)

Kun

thC

AT

RO

SA

lliac

eae

Alli

um la

cuno

sum

S. W

atso

nva

r. la

cuno

sum

RO

SC

RU

Alli

um p

raec

ox B

rand

egee

CL

EC

AT

RO

SC

RU

MIG

Am

aryl

lidac

eae

Am

aryl

lis b

ella

donn

a L

.*A

NA

MIG

Nar

ciss

us ta

zett

a L

.*C

RU

Ara

ceae

L

emna

min

or L

.M

IGZ

ante

desc

hia

aeth

iopi

ca (L

.) Sp

reng

.*M

IGA

reca

ceae

W

ashi

ngto

nia

filife

ra (A

ndré

) de

Bar

yC

RU

MIG

Asp

arag

acea

e A

spar

agus

asp

arag

oide

s (L

.) D

ruce

*N

ICA

spar

agus

offi

cina

lis L

.*C

AT

Asp

hode

lace

ae

Asp

hode

lus

fistu

losu

s L

.*C

LE

4

Cyp

erac

eae

Bol

bosc

hoen

us m

ariti

mus

(L.)

Palla

ssp.

pal

udos

us (A

. Nel

son)

T. K

oyam

aC

RU

MIG

Car

ex b

arba

rae

Dew

eyC

RU

Car

ex g

lobo

sa B

oott

CA

TR

OS

CR

UC

arex

gra

cilio

r M

ack.

RO

SC

arex

har

ford

ii M

ack.

RO

SC

RU

4 NY:

T.S

. Ros

s, 6

137,

04

13 1

992

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190 MONOGRAPHS OF THE WESTERN NORTH AMERICAN NATURALIST [Volume 7A

PPE

ND

IX2.

Con

tinue

d

Fam

ily/s

peci

esss

p./v

ar. a

nd in

fran

ame

CL

EN

ICC

AT

BA

RR

OS

CR

UA

NA

MIG

Car

ex p

ansa

L.H

. Bai

ley

RO

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egra

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oott

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ex s

enta

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a B

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RO

SC

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erus

era

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tis L

am.

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CL

EC

AT

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erus

invo

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s m

acro

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tton

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EN

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ernu

a (V

ahl)

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m. &

Sch

ult.

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hoen

ople

ctus

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eric

anus

N

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AT

7R

OS

(Per

s.) S

chin

z &

R. K

elle

rSc

hoen

ople

ctus

cal

iforn

icus

(C.A

. Mey

.) So

ják

RO

SC

RU

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enop

lect

us p

unge

ns (V

ahl)

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var.

long

ispi

catu

s (B

ritt

on) S

.G. S

m.

RO

SC

RU

Irid

acea

e Si

syri

nchi

um b

ellu

m S

. Wat

son

CA

TR

OS

CR

UM

IGJu

ncac

eae

Junc

us a

cutu

s L

.ss

p. le

opol

dii (

Parl

.) Sn

oger

upC

AT

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us b

altic

us W

illd.

ssp.

ate

r (R

ydb.

) Sno

geru

pC

AT

RO

SM

IGJu

ncus

buf

oniu

s L

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LE

NIC

CA

TR

OS

CR

UA

NA

MIG

Junc

us e

ffusu

s L

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p. p

acifi

cus

(Fer

nald

& W

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nd)

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per

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eatt

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mex

ican

us W

illd.

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OS

CR

UM

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ncus

pat

ens

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ey.

CL

ER

OS

CR

UM

IGJu

ncus

pha

eoce

phal

us E

ngel

m.

var.

phae

ocep

halu

sR

OS

Junc

us te

xtili

s B

uche

nau

CA

TJu

ncus

xip

hioi

des

E. M

ey.

CA

TC

RU

Luz

ula

subs

essi

lis (S

. Wat

son)

Buc

hena

u8R

OS

CR

UL

iliac

eae

Cal

ocho

rtus

alb

us (B

enth

.) B

enth

.C

AT

RO

SC

RU

Cal

ocho

rtus

cat

alin

ae S

. Wat

son

CA

TR

OS

CR

UC

aloc

hort

us lu

teus

Lin

dl.

CR

UC

aloc

hort

us s

plen

dens

Ben

th.

CA

TLi

lium

hum

bold

tii D

uch.

ssp.

oce

llatu

m (K

ello

gg) T

horn

eR

OS

CR

UA

NA

5 RSA

: Mar

k H

oefs

, Ste

ven

A. J

unak

, Jan

et T

akar

a, M

. Gay

, 238

4, J

ul 1

4 19

956 S

BB

G: M

.L. H

oefs

, R.F

. Tho

rne,

245

8, J

ul 2

6 19

957 R

SA: M

ark

Hoe

fs, R

.F. T

horn

e, J

anet

Tak

ara

Jul 2

6 19

95 2

482

8 Syn

onym

: Luz

ula

com

osa

E. M

ey.;

taxo

nom

ic is

sues

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2014] CALIFORNIA ISLAND PLANT DISTRIBUTION PATTERNS 191A

PPE

ND

IX2.

Con

tinue

d

Fam

ily/s

peci

esss

p./v

ar. a

nd in

fran

ame

CL

EN

ICC

AT

BA

RR

OS

CR

UA

NA

MIG

Mel

anth

iace

ae

Toxi

cosc

ordi

on fr

emon

tii (T

orr.)

Ryd

b.R

OS

CR

UA

NA

MIG

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hida

ceae

E

pipa

ctis

gig

ante

a H

ook.

CR

UPi

peri

a co

oper

i (S.

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son)

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ria

elon

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peri

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(Gre

ene)

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RU

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eae

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cyl

indr

ica

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ostis

exa

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n.C

AT

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ostis

pal

lens

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n.C

LE

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OS

CR

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gros

tis s

tolo

nife

ra*

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CA

TA

NA

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mop

hila

are

nari

a (L

.) L

ink*

NIC

And

ropo

gon

glom

erat

us (W

alte

r) B

ritt

on e

t al.

var.

scab

rigl

umis

C.S

. Cam

pb.

CR

UA

rist

ida

adsc

ensi

onis

L.

CL

EC

AT

RO

SC

RU

Ari

stid

a te

rnip

es C

av.

var.

gent

ilis

(Hen

rard

) Allr

edC

RU

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ndo

dona

x L

.*N

ICR

OS

CR

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IGA

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bar

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k*C

LE

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CA

TB

AR

RO

SC

RU

AN

AM

IGA

vena

fatu

a L

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LE

NIC

CA

TB

AR

RO

SC

RU

AN

AM

IGA

vena

sat

iva

L.*

CL

EC

AT

RO

SC

RU

Bot

hrio

chlo

a ba

rbin

odis

(Lag

.) H

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rC

AT

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odiu

m d

ista

chyo

n (L

.) P.

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uv.*

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E? 1

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min

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mus

ari

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cus

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ar) S

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CL

EN

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AT

BA

RR

OS

CR

UA

NA

MIG

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mus

ber

tero

anus

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la12

CL

EN

ICC

AT

BA

RR

OS

CR

UA

NA

MIG

Bro

mus

car

inat

us H

ook.

& A

rn.

var.

cari

natu

sC

LE

?N

ICC

AT

RO

SC

RU

AN

AM

IGB

rom

us m

ariti

mus

(Pip

er) H

itchc

.R

OS

CR

UA

NA

MIG

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mus

cat

hart

icus

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l*C

LE

13N

ICC

RU

*Bro

mus

dia

ndru

s R

oth*

CL

EN

ICC

AT

BA

RR

OS

CR

UA

NA

MIG

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mus

hor

deac

eus

L.*

14C

LE

NIC

CA

TB

AR

RO

SC

RU

AN

AM

IGB

rom

us la

evip

es S

hear

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AT

RO

SC

RU

Bro

mus

mad

rite

nsis

L.*

CL

EN

ICC

AT

BA

RR

OS

CR

UA

NA

MIG

Bro

mus

vul

gari

s (H

ook.

) She

ar15

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AT

Cal

amag

rost

is r

ubes

cens

Buc

kley

CR

U

9 Per

s. c

om. E

. How

e10

SBB

G: S

.A. J

unak

Apr

30

1998

SC

a-56

811

Pers

. com

. E. H

owe

12Sy

nony

m: B

rom

us tr

inii

Des

v.; P

er J

epso

n In

terc

hang

e: m

olec

ular

evi

denc

e ne

eded

to r

esol

ve q

uest

ion

of n

ativ

ity

13SB

BG

: T.S

. Ros

s, E

. Kel

logg

Apr

11

1992

612

214

Syno

nym

: Bro

mus

mol

lisL

.15

UC

SB: D

an H

anco

ck 0

3/30

/196

1 14

6

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192 MONOGRAPHS OF THE WESTERN NORTH AMERICAN NATURALIST [Volume 7A

PPE

ND

IX2.

Con

tinue

d

Fam

ily/s

peci

esss

p./v

ar. a

nd in

fran

ame

CL

EN

ICC

AT

BA

RR

OS

CR

UA

NA

MIG

Chl

oris

vir

gata

Sw

.*C

LE

16

Cor

tade

ria

sello

ana

(Sch

ult.

& S

chul

t. f.)

C

LE

? 17

NIC

CA

TC

RU

MIG

Asc

h. &

Gra

ebn.

*C

ryps

is v

agin

iflor

a (F

orss

k.) O

piz*

CA

T18

Cry

psis

alo

pecu

roid

es (P

iller

& M

itter

p.) S

chra

d.*

CL

E? 1

9

Cyn

odon

dac

tylo

n (L

.) Pe

rs.*

CL

EN

ICC

AT

BA

RR

OS

CR

UA

NA

MIG

Dac

tylis

glo

mer

ata

L.*

CL

EC

AT

CR

UD

anth

onia

cal

iforn

ica

Bol

.R

OS

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cham

psia

dan

thon

ioid

es (T

rin.

) Mun

roC

LE

Des

maz

eria

rig

ida

(L.)

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AT

20

Dig

itari

a sa

ngui

nalis

(L.)

Scop

.*C

AT

21

Dis

sant

heliu

m c

alifo

rnic

um (N

utt.)

Ben

th.

CA

TD

istic

hlis

spi

cata

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ene

CL

EN

ICC

AT

RO

SC

RU

AN

AM

IGD

istic

hlis

litt

oral

is (E

ngel

m.)

H.L

. C

AT

RO

SB

ell &

Col

umbu

sE

chin

ochl

oa c

rus-

galli

(L.)

P. B

eauv

. *C

LE

CA

TE

hrha

rta

caly

cina

* Sm

.C

LE

CA

TE

hrha

rta

erec

ta L

am.*

CA

T22

CR

UE

lym

us g

lauc

us B

uckl

eyss

p. g

lauc

usC

AT

RO

SC

RU

Ely

mus

rep

ens

(L.)

Gou

ld*

MIG

Ely

mus

con

dens

atus

J. P

resl

CL

EN

ICC

AT

RO

SC

RU

AN

AM

IGE

lym

us p

acifi

cus

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ldN

ICR

OS

MIG

Ely

mus

triti

coid

es B

uckl

eyN

ICC

AT

RO

SC

RU

AN

AM

IGFe

stuc

a ar

undi

nace

a Sc

hreb

.*

CL

EN

ICC

AT

RO

SC

RU

MIG

Fest

uca

pere

nnis

(L.)

Col

umbu

s &

J.P.

Sm

.*C

LE

NIC

CA

TB

AR

RO

SC

RU

MIG

Fest

uca

tem

ulen

ta (L

.) C

olum

bus

& J.

P. S

m.*

CL

EC

AT

RO

SC

RU

Fest

uca

brom

oide

s L

.*C

LE

CA

TR

OS

CR

UA

NA

MIG

Fest

uca

mic

rost

achy

s N

utt.

CL

EC

AT

CR

UA

NA

MIG

Fest

uca

myu

ros

L.*

CL

EN

ICC

AT

BA

RR

OS

CR

UA

NA

MIG

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uca

octo

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ter

CL

EN

ICC

AT

BA

RR

OS

CR

UA

NA

MIG

Gas

trid

ium

phl

eoid

es (N

ees

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eyen

) C.E

. Hub

b.*

CL

EC

AT

RO

SC

RU

Hai

nard

ia c

ylin

dric

a (W

illd.

) Gre

uter

*C

LE

23C

AT

24R

OS

CR

UM

IG

16C

AS:

Ste

ven

A. J

unak

9 N

ov 1

990

SCI-

143

17Pe

rs. c

om. E

. How

e18

RSA

: Mar

k H

oefs

, R.F

. Tho

rne,

Jan

et T

akar

a, M

. Gay

Jul

27

1995

249

119

Pers

. com

. E. H

owe

20C

AS:

Pet

er H

. Rav

en 2

2 M

ay 1

962

1782

621

UC

D: B

eech

er C

ram

pton

08

10 1

968

8322

22SS

BG

: S.A

. Jun

ak J

un 1

3 19

98 S

Ca-

743

23SB

BG

: S.A

. Jun

ak M

ay 1

4 19

85 S

Cl-

7124

SBB

G: S

.A. J

unak

, M. L

. Hoe

fs J

un 4

199

7 SC

a-42

1

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2014] CALIFORNIA ISLAND PLANT DISTRIBUTION PATTERNS 193A

PPE

ND

IX2.

Con

tinue

d

Fam

ily/s

peci

esss

p./v

ar. a

nd in

fran

ame

CL

EN

ICC

AT

BA

RR

OS

CR

UA

NA

MIG

Hor

deum

bra

chya

nthe

rum

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ski

ssp.

cal

iforn

icum

(Cov

as &

Ste

bbin

s)

NIC

CA

T25

RO

SC

RU

AN

AB

othm

er e

t al.

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deum

dep

ress

um (S

crib

n. &

J.G

. Sm

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orde

um in

terc

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s N

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AR

RO

SC

RU

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AM

IGH

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um m

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urin

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RU

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um v

ulga

re L

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OS

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CR

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ica

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CR

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lenb

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icro

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AR

RO

SC

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AM

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raph

olis

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UA

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alum

dis

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m L

.C

AT

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nnis

etum

cla

ndes

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ov.*

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RO

SC

RU

AN

AM

IGPe

nnis

etum

set

aceu

m (F

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k.) C

hiov

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a L

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ICC

AT

CR

UA

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aris

can

arie

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AR

CR

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mm

onii

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aris

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etz.

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TB

AR

RO

SC

RU

AN

AM

IGPh

alar

is p

arad

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RR

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CR

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ecun

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AT

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OS

CR

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pogo

n vi

ridi

s (G

ouan

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istr.

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AT

BA

RR

OS

CR

UA

NA

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pogo

n in

terr

uptu

s K

unth

*C

LE

CA

TR

OS

CR

UM

IG

25SB

BG

: L.W

. Nut

tall

May

12

1920

190

26PO

M: P

.A. M

unz

Apr

8 1

923

6613

27SB

BG

: S.A

. Jun

ak M

ay 3

200

1 SC

a-14

6428

SBB

G: P

.H. R

aven

May

10

1962

177

3829

CA

S: E

.R. B

lakl

ey 7

Dec

. 196

3 63

2730

S. R

atay

col

lect

ion

31SB

BG

: S.A

. Jun

ak, M

.L. H

oefs

Jun

5 1

997

SCa-

427

32U

C: P

eter

H. R

aven

Apr

12

1962

173

3233

UC

: F.R

. Fos

berg

Mar

20

1931

S43

10

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194 MONOGRAPHS OF THE WESTERN NORTH AMERICAN NATURALIST [Volume 7A

PPE

ND

IX2.

Con

tinue

d

Fam

ily/s

peci

esss

p./v

ar. a

nd in

fran

ame

CL

EN

ICC

AT

BA

RR

OS

CR

UA

NA

MIG

Poly

pogo

n m

onsp

elie

nsis

(L.)

Des

f.*C

LE

NIC

CA

TB

AR

RO

SC

RU

AN

AM

IGSc

hism

us a

rabi

cus

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s*C

LE

34N

ICC

AT

35B

AR

CR

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hism

us b

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ll.*

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2014] CALIFORNIA ISLAND PLANT DISTRIBUTION PATTERNS 195A

PPE

ND

IX2.

Con

tinue

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ily/s

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196 MONOGRAPHS OF THE WESTERN NORTH AMERICAN NATURALIST [Volume 7A

PPE

ND

IX2.

Con

tinue

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Fam

ily/s

peci

esss

p./v

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3 66

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ober

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akar

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tace

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pril

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1

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2014] CALIFORNIA ISLAND PLANT DISTRIBUTION PATTERNS 197A

PPE

ND

IX2.

Con

tinue

d

Fam

ily/s

peci

esss

p./v

ar. a

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owe

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.L. H

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and

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198 MONOGRAPHS OF THE WESTERN NORTH AMERICAN NATURALIST [Volume 7A

PPE

ND

IX2.

Con

tinue

d

Fam

ily/s

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esss

p./v

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y) G

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eC

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69SB

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ak, S

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ak, K

. Kir

klan

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hman

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1998

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k H

oefs

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rops

t Mar

14

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hrys

anth

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cor

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75U

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.M. B

eauc

ham

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d H

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ven

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., M

ay 2

2 19

2080

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.A. J

unak

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1997

SC

l-78

4

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2014] CALIFORNIA ISLAND PLANT DISTRIBUTION PATTERNS 199A

PPE

ND

IX2.

Con

tinue

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ily/s

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esss

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ar. a

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74

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200 MONOGRAPHS OF THE WESTERN NORTH AMERICAN NATURALIST [Volume 7A

PPE

ND

IX2.

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tinue

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mor

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BA

RR

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teve

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ak, M

ary

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ochb

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. Fer

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l 28

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91 5

184

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2014] CALIFORNIA ISLAND PLANT DISTRIBUTION PATTERNS 201A

PPE

ND

IX2.

Con

tinue

d

Fam

ily/s

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esss

p./v

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y) G

reen

eva

r. in

term

edia

CL

EC

AT

97SB

BG

: R.N

. Phi

lbri

ck M

ar 1

8 19

68 B

68-2

098

Syno

nym

: Mic

rose

ris

hete

roca

rpa

(Nut

t.) K

.L. C

ham

bers

99R

SA: F

.R. F

osbe

rg, 4

460,

Apr

3 1

931

100 R

SA: O

rlan

do M

istr

etta

Apr

5 1

992

206

101 S

D: R

ober

t F. T

horn

e Se

p 15

, 196

6 36

698

102 S

D: R

ober

t F. T

horn

e, P

. Eve

rett

, 368

23, A

pr 1

8, 1

967

103 S

BB

G: M

.L. H

oefs

, R.F

. Tho

rne

Apr

6 1

996

2665

104 S

ynon

ym: M

icro

seri

s lin

eari

folia

(DC

.) Sc

h. B

ip.

105 N

ativ

e st

atus

on

isla

nds

unkn

own

106 N

ativ

e st

atus

on

isla

nds

unkn

own

107 U

CR

: J.R

. Ekh

off M

ay 1

6 20

01 s

.n.

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202 MONOGRAPHS OF THE WESTERN NORTH AMERICAN NATURALIST [Volume 7A

PPE

ND

IX2.

Con

tinue

d

Fam

ily/s

peci

esss

p./v

ar. a

nd in

fran

ame

CL

EN

ICC

AT

BA

RR

OS

CR

UA

NA

MIG

Cry

ptan

tha

clev

elan

dii G

reen

eC

LE

CA

TB

AR

RO

SC

RU

AN

AM

IGC

rypt

anth

a le

ioca

rpa

(Fis

ch. &

C.A

. Mey

.) G

reen

eR

OS

MIG

Cry

ptan

tha

mar

itim

a (G

reen

e) G

reen

eC

LE

NIC

CA

T10

8B

AR

Cry

ptan

tha

mic

rom

eres

(A. G

ray)

Gre

ene

CA

TR

OS

CR

UC

rypt

anth

a m

icro

stac

hys

(A. G

ray)

Gre

ene

CA

TC

rypt

anth

a m

uric

ata

(Hoo

k. &

Arn

.) C

RU

A. N

elso

n &

J.F.

Mac

br.

Cry

ptan

tha

tras

kiae

I.M

. Joh

nst.

CL

EN

ICC

AT

109

Cry

ptan

tha

wig

gins

ii I.

M. J

ohns

t.C

AT

110

Em

men

anth

e pe

ndul

iflor

a B

enth

.va

r. pe

ndul

iflor

aC

LE

CA

TC

RU

Eri

odic

tyon

tras

kiae

Eas

tw.

ssp.

tras

kiae

CA

TE

ucry

pta

chry

sant

hem

ifolia

(Ben

th.)

Gre

ene

var.

chry

sant

hem

ifolia

CL

EC

AT

BA

RR

OS

CR

UA

NA

MIG

Har

pago

nella

pal

mer

i A. G

ray

CA

TH

elio

trop

ium

cur

assa

vicu

m L

.va

r. oc

ulat

um (A

. Hel

ler)

C

LE

NIC

CA

TR

OS

CR

UA

NA

MIG

I.M

. Joh

nst.

ex T

ides

tr.N

ama

sten

ocar

pum

A. G

ray

CL

E11

1

Nem

ophi

la m

enzi

esii

Hoo

k. &

Arn

.C

AT

Nem

ophi

la p

edun

cula

ta B

enth

.N

ICR

OS

CR

UM

IGPe

ctoc

arya

line

aris

(Rui

z &

Pav

.) D

C.

ssp.

fero

cula

(I.M

. Joh

nst.)

Tho

rne

CL

E11

2N

ICC

AT

RO

SC

RU

AN

APe

ctoc

arya

pen

icill

ata

(Hoo

k. &

Arn

.) A

. DC

.C

AT

CR

UPh

acel

ia c

icut

aria

Gre

ene

var.

hisp

ida

J.T. H

owel

lC

LE

CA

TR

OS

CR

UA

NA

Phac

elia

hub

byi (

J.F. M

acbr

.) L

.M. G

arri

son1

13C

RU

Phac

elia

dis

tans

Ben

th.

CL

EN

ICC

AT

BA

RR

OS

CR

UA

NA

MIG

Phac

elia

flor

ibun

da G

reen

eC

LE

Phac

elia

gra

ndifl

ora

(Ben

th.)

A. G

ray

CA

TR

OS

CR

U11

4

Phac

elia

insu

lari

s M

unz

var.

insu

lari

sR

OS

MIG

Phac

elia

lyon

ii A

. Gra

yC

LE

CA

TPh

acel

ia r

amos

issi

ma

Dou

glas

ex

Leh

m.

RO

SC

RU

MIG

Phac

elia

vis

cida

(Lin

dl.)

Torr

.C

AT

RO

SC

RU

AN

AM

IGPh

olis

tom

a au

ritu

m (L

indl

.) L

ilja

var.

auri

tum

CL

E?

CA

TB

AR

CR

UPh

olis

tom

a ra

cem

osum

(A. G

ray)

Con

stan

ceC

LE

CA

TB

AR

CR

UPl

agio

both

rys

acan

thoc

arpu

s (P

iper

) I.M

. Joh

nst.

CA

T11

5C

RU

Plag

iobo

thry

s ca

nesc

ens

Ben

th.

CL

EC

AT

RO

SC

RU

AN

A

108 U

C: T

.S. B

rand

egee

May

20

1890

109 S

D: R

. Mitc

hel B

eauc

ham

p, M

. Dou

glas

Mar

24,

197

2 32

2211

0 UC

: Mic

hael

G. S

imps

on, L

ori L

. Sim

pson

Apr

il 21

201

2 36

8211

1 RSA

: E. K

ello

gg J

un 2

8 19

92 s

.n.

112 S

BB

G: S

.A. J

unak

Mar

7 1

997

SCl-

732

113 S

ynon

ym: P

hace

lia c

icut

aria

Gre

ene

var.

hubb

yi(J

.F. M

acbr

.) J.T

. How

ell

114 I

n W

alla

ce 1

985

but n

ot N

PS li

st11

5 UC

R: T

.S. R

oss

Apr

22

1993

689

5

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2014] CALIFORNIA ISLAND PLANT DISTRIBUTION PATTERNS 203A

PPE

ND

IX2.

Con

tinue

d

Fam

ily/s

peci

esss

p./v

ar. a

nd in

fran

ame

CL

EN

ICC

AT

BA

RR

OS

CR

UA

NA

MIG

Plag

iobo

thry

s co

llinu

s (P

hil.)

I.M

. Joh

nst.

var.

grac

ilis

(I.M

. Joh

nst.)

Hig

gins

CL

EC

AT

CR

UA

NA

MIG

Plag

iobo

thry

s co

llinu

s (P

hil.)

I.M

. Joh

nst.

var.

calif

orni

cus

(A. G

ray)

Hig

gins

C

LE

116

CA

T11

7C

RU

118

Bra

ssic

acea

e A

thys

anus

pus

illus

(Hoo

k.) G

reen

eC

LE

CA

TC

RU

Boe

cher

a ho

ffman

nii (

Mun

z) A

l-Sh

ehba

zR

OS

CR

UB

rass

ica

nigr

a (L

.) W

.D.J.

Koc

h*C

LE

NIC

CA

TB

AR

RO

SC

RU

AN

AM

IGB

rass

ica

tour

nefo

rtii

Gou

an*

NIC

Bra

ssic

a ra

pa L

.*C

LE

CA

TR

OS

CR

UM

IGC

akile

ede

ntul

a (B

igel

ow) H

ook.

*N

ICR

OS

MIG

Cak

ile m

ariti

ma

Scop

.*C

LE

NIC

CA

TR

OS

CR

UA

NA

MIG

Cap

sella

bur

sa-p

asto

ris

(L.)

Med

ik.*

CL

EC

AT

RO

SC

RU

MIG

Car

dam

ine

calif

orni

ca (N

utt.)

Gre

ene

CA

TR

OS

CR

UA

NA

MIG

Car

dam

ine

olig

ospe

rma

Nut

t.C

RU

Cau

lant

hus

lasi

ophy

llus

(Hoo

k. &

Arn

.) Pa

yson

119

CL

EC

AT

BA

RR

OS

CR

UA

NA

MIG

Des

cura

inia

pin

nata

(Wal

ter)

Bri

tton

CL

EC

AT

RO

SC

RU

AN

AD

ithyr

ea m

ariti

ma

(Dav

idso

n) D

avid

son

NIC

CA

T12

0M

IGD

raba

cun

eifo

lia N

utt.

ex T

orr.

& A

. Gra

yC

LE

NIC

CA

TC

RU

Ery

sim

um a

mm

ophi

lum

A. H

elle

rR

OS

MIG

Ery

sim

um c

heir

i (L

.) C

rant

z*C

AT

Ery

sim

um in

sula

re G

reen

eR

OS

CR

UA

NA

MIG

Hir

schf

eldi

a in

cana

(L.)

Lag

r.-Fo

ssat

121 *

CL

EN

ICC

AT

RO

SC

RU

MIG

Hor

nung

ia p

rocu

mbe

ns (L

.) H

ayek

CL

E? 1

22N

ICB

AR

RO

SC

RU

AN

AM

IGL

epid

ium

dra

ba L

.*C

AT

RO

SC

RU

Lep

idiu

m la

sioc

arpu

m N

utt.

CL

EN

ICC

AT

Lep

idiu

m la

tipes

Hoo

k.C

LE

CA

TC

RU

Lep

idiu

m n

itidu

m N

utt.

CL

EN

ICC

AT

BA

RR

OS

CR

UA

NA

MIG

Lep

idiu

m o

blon

gum

Sm

all

CL

EN

ICC

AT

RO

SC

RU

AN

AM

IGL

epid

ium

str

ictu

m (S

. Wat

son)

Rat

tan*

CA

T12

3

Lep

idiu

m v

irgi

nicu

m L

.124

*ss

p. m

enzi

esii

(DC

.) T

hell.

CL

EC

AT

CR

UL

obul

aria

mar

itim

a (L

.) D

esv.

*C

LE

125

NIC

CA

TM

IGN

astu

rtiu

m o

ffici

nale

W.T

. Aito

n126

NIC

CA

TC

RU

116 P

OM

: P.A

. Mun

z A

pr 1

0 19

23 6

705

117 R

SA: R

.F. T

horn

e, R

.C. R

ollin

s, D

. Pro

pst,

R. C

arol

in, 3

6761

, Mar

19

1967

118 U

C: T

.S. B

rand

egee

, Apr

188

811

9 Syn

onym

: Gui

lleni

a la

siop

hylla

(Hoo

k. &

Arn

.) G

reen

e12

0 No

spec

imen

s in

CC

H, e

xtir

pate

d fr

om C

atal

ina?

per

: htt

p://w

ww

.rar

epla

nts.

cnps

.org

/det

ail/5

71.h

tml

121 S

ynon

ym: B

rass

ica

geni

cula

ta(D

esf.)

Ben

th.

122 P

ers.

com

. E. H

owe

123 S

BB

G: S

.A. J

unak

, M.L

. Hoe

fs, J

. Tak

ara

Mar

28

1997

SC

a-38

812

4 Syn

onym

s: L

epid

ium

vir

gini

cum

var.

pube

scen

s(G

reen

e) T

hell.

; Lep

idiu

m v

irgi

nicu

mva

r. ro

bins

onii

(The

ll.) C

.L. H

itchc

.12

5 SB

BG

: S.A

. Jun

ak, M

.C. H

ochb

erg,

H. F

ergu

son

Jul 2

8 19

81 S

Cl-

1712

6 Nat

ive

stat

us o

n is

land

s un

know

n, d

oubt

ful

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204 MONOGRAPHS OF THE WESTERN NORTH AMERICAN NATURALIST [Volume 7A

PPE

ND

IX2.

Con

tinue

d

Fam

ily/s

peci

esss

p./v

ar. a

nd in

fran

ame

CL

EN

ICC

AT

BA

RR

OS

CR

UA

NA

MIG

Rap

hanu

s ra

phan

istr

um L

.*C

LE

CA

TR

OS

CR

UR

apha

nus

sativ

us L

.*C

LE

NIC

CA

TR

OS

CR

UM

IGSi

bara

filif

olia

(Gre

ene)

Gre

ene

CL

E12

7C

AT

CR

USi

napi

s ar

vens

is L

.128

*C

LE

? 129

CA

TC

RU

Sisy

mbr

ium

alti

ssim

um L

.*Si

sym

briu

m ir

io L

.*C

LE

NIC

CA

TA

NA

Sisy

mbr

ium

offi

cina

le (L

.) Sc

op.*

CA

TC

RU

MIG

Sisy

mbr

ium

ori

enta

le L

.*C

LE

130

NIC

CA

TA

NA

Thy

sano

carp

us c

onch

ulife

rus

Gre

ene

CR

UT

hysa

noca

rpus

cur

vipe

s H

ook.

CL

E13

1C

AT

CR

UT

hysa

noca

rpus

laci

niat

us N

utt.

CL

EC

AT

RO

SC

RU

Trop

idoc

arpu

m g

raci

le H

ook.

CL

EC

AT

Turr

itis

glab

ra L

.C

RU

Cac

tace

ae

Ber

gero

cact

us e

mor

yi (E

ngel

m.)

Bri

tton

& R

ose

CL

EC

AT

Cyl

indr

opun

tia p

rolif

era

(Eng

elm

.) F.

M. K

nuth

CL

EN

ICC

AT

BA

RR

OS

CR

UA

NA

Opu

ntia

ficu

s-in

dica

(L.)

Mill

.*C

LE

NIC

CA

TC

RU

AN

AO

punt

ia li

ttor

alis

(Eng

elm

.) C

ocke

rell

CL

EN

ICC

AT

BA

RR

OS

CR

UA

NA

MIG

Opu

ntia

ori

cola

Phi

lbri

ckC

LE

NIC

CA

TB

AR

RO

SC

RU

AN

AM

IGC

ampa

nula

ceae

Gith

opsi

s di

ffusa

A. G

ray

ssp.

diff

usa

CR

UTr

ioda

nis

biflo

ra (R

uiz

& P

av.)

Gre

ene

CA

TR

OS

CR

UC

apri

folia

ceae

L

onic

era

hisp

idul

a (L

indl

.) To

rr. &

A. G

ray

CL

EC

AT

RO

SC

RU

Lon

icer

a in

terr

upta

Ben

th.

CR

UL

onic

era

subs

pica

ta H

ook.

& A

rn.

CA

TR

OS

CR

USy

mph

oric

arpo

s m

ollis

Nut

t.C

AT

RO

SC

RU

Car

yoph

ylla

ceae

C

ardi

onem

a ra

mos

issi

mum

(Wei

nm.)

A. N

elso

nR

OS

CR

UM

IG&

J.F.

Mac

br.

Cer

astiu

m g

lom

erat

um T

huill

.*C

LE

CA

TR

OS

CR

UA

NA

MIG

Her

niar

ia h

irsu

ta L

.*va

r. ci

nere

a (D

C.)

Lor

et &

Bar

rand

onC

LE

NIC

CA

T13

2B

AR

RO

SM

IGL

oefli

ngia

squ

arro

sa N

utt.

NIC

Min

uart

ia d

ougl

asii

(Tor

r. &

A. G

ray)

Mat

tf.C

LE

CA

TR

OS

CR

UPo

lyca

rpon

dep

ress

um N

utt.

CL

EC

AT

CR

UM

IG

127 R

SA: O

rlan

do M

istr

etta

Apr

5 1

992

142

128 S

ynon

ym: B

rass

ica

kabe

r(D

C.)

L.C

. Whe

eler

129 P

ers.

com

. E. H

owe

130 S

BB

G: S

.A. J

unak

May

29

1996

SC

l-54

513

1 UC

: Tim

Ros

s A

pr 1

4 19

92 6

153

132 S

BB

G: S

.A. J

unak

Mar

27

1997

SC

a-34

6

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2014] CALIFORNIA ISLAND PLANT DISTRIBUTION PATTERNS 205A

PPE

ND

IX2.

Con

tinue

d

Fam

ily/s

peci

esss

p./v

ar. a

nd in

fran

ame

CL

EN

ICC

AT

BA

RR

OS

CR

UA

NA

MIG

Poly

carp

on te

trap

hyllu

m (L

.) L

.*C

AT

RO

SC

RU

AN

AM

IGSa

gina

ape

tala

Ard

.N

ICR

OS

CR

UM

IGSa

gina

dec

umbe

ns (E

lliot

t) To

rr. &

A. G

ray

ssp.

occ

iden

talis

(S. W

atso

n) G

.E. C

row

CA

TR

OS

CR

UM

IGSi

lene

ant

irrh

ina

L.

CL

EC

AT

RO

SC

RU

MIG

Sile

ne g

allic

a L

.*C

LE

NIC

CA

TB

AR

RO

SC

RU

AN

AM

IGSi

lene

laci

niat

a C

av.

ssp.

laci

niat

aC

LE

CA

TR

OS

CR

UA

NA

MIG

Sile

ne c

onifl

ora

Ott

hC

AT

CR

USp

ergu

la a

rven

sis

L.*

CA

TR

OS

Sper

gula

ria

bocc

oni (

Sche

ele)

Gra

ebn.

*C

LE

NIC

CA

TB

AR

CR

UA

NA

Sper

gula

ria

mac

roth

eca

(Cha

m. &

Sch

ltdl.)

Hey

nh.

ssp.

mac

roth

eca

CL

EN

ICC

AT

BA

RR

OS

CR

UA

NA

MIG

Sper

gula

ria

mar

ina

(L.)

Bes

ser

CL

EN

ICC

AT

BA

RR

OS

CR

UA

NA

Sper

gula

ria

villo

sa (P

ers.

) Cam

bess

.*C

LE

CA

TR

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ia (L

.) Vi

ll.*

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ism

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itoid

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ICC

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um s

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EC

AT

RO

SC

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iple

x ar

gent

ea N

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exp

ansa

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son)

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OS

CR

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lex

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EN

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OS

CR

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sh) N

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lex

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.) D

. Die

tr.C

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TR

OS

CR

UA

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MIG

Atr

iple

x le

ntifo

rmis

(Tor

r.) S

. Wat

son

(S. W

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n) H

.M. H

all &

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m.

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EN

ICC

AT

CR

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iple

x le

ucop

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(Moq

.) D

. Die

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LE

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CA

TR

OS

CR

UA

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iple

x pa

cific

a A

. Nel

son

CL

EN

ICC

AT

BA

RR

OS

CR

UA

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iple

x se

mib

acca

ta R

. Br.*

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EN

ICC

AT

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RR

OS

CR

UA

NA

MIG

Atr

iple

x se

rena

na A

bram

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vids

onii

(Sta

ndl.)

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zC

AT

133

RO

SC

RU

Atr

iple

x se

rena

na A

bram

sva

r. se

rena

naC

AT

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iple

x pr

ostr

ata

DC

.*N

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RU

MIG

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iple

x w

atso

nii A

bram

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LE

NIC

CA

TR

OS

CR

UA

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MIG

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sia

hyss

opifo

lia (P

all.)

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tze*

CL

EN

ICC

AT

Bet

a vu

lgar

is L

.*ss

p. m

ariti

ma

(L.)

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ang.

CL

EC

AT

RO

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RU

AN

AM

IGC

heno

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um b

erla

ndie

ri M

oq.

NIC

RO

SC

RU

AN

AC

heno

podi

um c

alifo

rnic

um (S

. Wat

son)

S. W

atso

nC

LE

NIC

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TB

AR

RO

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AM

IGC

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ural

e L

.*C

LE

NIC

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TB

AR

RO

SC

RU

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AM

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ania

am

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ioid

es (L

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osya

kin

& C

lem

ants

*N

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AT

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SC

RU

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phan

ia m

ultif

ida

(L.)

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yaki

n &

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man

ts*

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ER

OS

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olep

is n

utta

llian

a (S

chul

t.) G

reen

eC

LE

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RR

OS

CR

UM

IGSa

licor

nia

depr

essa

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ndl.

NIC

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orni

a pa

cific

a St

andl

.C

LE

CA

TR

OS

CR

UA

NA

MIG

133 R

SA: B

lanc

he T

rask

Mar

190

1 s.

n.

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206 MONOGRAPHS OF THE WESTERN NORTH AMERICAN NATURALIST [Volume 7A

PPE

ND

IX2.

Con

tinue

d

Fam

ily/s

peci

esss

p./v

ar. a

nd in

fran

ame

CL

EN

ICC

AT

BA

RR

OS

CR

UA

NA

MIG

Sals

ola

aust

ralis

R. B

r.134

*C

LE

NIC

CA

TR

OS

CR

USu

aeda

cal

ceol

iform

is (H

ook.

) Moq

.C

AT

135

Suae

da ta

xifo

lia (S

tand

l.) S

tand

l.C

LE

NIC

CA

TB

AR

RO

SC

RU

AN

AM

IGC

ista

ceae

H

elia

nthe

mum

gre

enei

B.L

. Rob

.C

AT

RO

SC

RU

MIG

&H

elia

nthe

mum

sco

pari

um N

utt.

CL

E? 1

36C

AT

RO

SC

RU

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omac

eae

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tom

a ar

bore

a (N

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. Ilti

sC

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CA

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OS

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volv

ulac

eae

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yste

gia

mac

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(Gre

ene)

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mm

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mpl

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ma

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mm

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LE

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BA

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alys

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a m

acro

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ia (G

reen

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mitt

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egia

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se) B

rum

mitt

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TC

alys

tegi

a m

acro

steg

ia (G

reen

e) B

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mitt

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rmed

ia (A

bram

s) B

rum

mitt

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TC

alys

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a m

acro

steg

ia (G

reen

e) B

rum

mitt

ssp.

mac

rost

egia

CA

TR

OS

CR

UA

NA

MIG

Cal

yste

gia

mal

acop

hylla

(Gre

ene)

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z*ss

p. p

edic

ella

ta (J

eps.

) Mun

zN

IC&

Cal

yste

gia

sold

anel

la (L

.) R

. Br.*

CL

EN

ICC

AT

RO

SC

RU

MIG

Con

volv

ulus

arv

ensi

s L

.*N

ICC

AT

RO

SC

RU

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volv

ulus

sim

ulan

s L

.M. P

erry

CL

EC

AT

CR

UC

ress

a tr

uxill

ensi

s K

unth

CL

EN

ICC

AT

RO

SC

RU

AN

AM

IGC

uscu

ta c

alifo

rnic

a H

ook.

& A

rn.

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ER

OS

CR

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7

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cuta

cam

pest

ris

Yunc

k.C

AT

138

Cus

cuta

occ

iden

talis

Mill

sp.

CA

TC

uscu

ta p

acifi

ca C

oste

a &

M. W

righ

tva

r. pa

cific

aR

OS

CR

UA

NA

Dic

hond

ra o

ccid

enta

lis H

ouse

CA

TR

OS

CR

UM

IGIp

omoe

a ca

iric

a (L

.) Sw

eet*

CA

TC

orna

ceae

C

ornu

s gl

abra

ta B

enth

.C

AT

Cra

ssul

acea

e C

rass

ula

aqua

tica

(L.)

Schö

nl.

CA

TC

rass

ula

conn

ata

(Rui

z &

Pav

.) A

. Ber

ger

CL

EN

ICC

AT

BA

RR

OS

CR

UA

NA

MIG

Cra

ssul

a ov

ata

(Mill

.) D

ruce

*C

LE

Dud

leya

blo

chm

ania

e (E

astw

.) M

oran

ssp.

blo

chm

ania

eC

RU

Dud

leya

blo

chm

ania

e (E

astw

.) M

oran

ssp.

insu

lari

s (M

oran

) Mor

anR

OS

Dud

leya

cae

spito

sa (H

aw.)

Bri

tton

& R

ose

AN

AD

udle

ya c

ande

labr

um R

ose

RO

SC

RU

MIG

Dud

leya

gre

enei

Ros

eC

AT

RO

SC

RU

MIG

Dud

leya

gno

ma

S.W

. McC

abe

RO

S

134 S

ynon

ym: S

also

la tr

agus

L.

135 S

BB

G: S

.A. J

unak

Oct

22

1998

SC

a-82

313

6 Per

s. c

om. E

. How

e13

7 SB

BG

: M.R

. Ben

edic

t Sep

30

1969

13

8 RSA

: Geo

rge

B. G

rant

454

1

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2014] CALIFORNIA ISLAND PLANT DISTRIBUTION PATTERNS 207A

PPE

ND

IX2.

Con

tinue

d

Fam

ily/s

peci

esss

p./v

ar. a

nd in

fran

ame

CL

EN

ICC

AT

BA

RR

OS

CR

UA

NA

MIG

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leya

nes

iotic

a (M

oran

) Mor

anC

RU

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leya

tras

kiae

(Ros

e) M

oran

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udle

ya v

iren

s (R

ose)

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anss

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asse

i (R

ose)

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vir

ens

(Ros

e) M

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ICC

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Dud

leya

vir

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(Ros

e) M

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vir

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CL

EC

ross

osom

atac

eae

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oma

calif

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t.C

LE

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CA

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ucur

bita

ceae

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sim

a K

unth

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UM

arah

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din)

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ene

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E13

9N

IC14

0C

AT

141

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UM

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2

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ah m

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a (G

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reen

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TB

AR

RO

SC

RU

AN

AM

IGE

rica

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A

rbut

us m

enzi

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hC

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Arc

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cata

linae

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. Wel

lsC

AT

Arc

tost

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conf

ertif

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Eas

tw.

RO

SA

rcto

stap

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s in

sula

ris

Parr

yC

RU

Arc

tost

aphy

los

crus

tace

a E

astw

.ss

p. in

sulic

ola

(P.V

. Wel

ls) V

.T. P

arke

r et

al.

RO

SC

RU

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tost

aphy

los

crus

tace

a E

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.ss

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ubco

rdat

a (E

astw

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arke

r et

al.

RO

SC

RU

Arc

tost

aphy

los

viri

diss

ima

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inn

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osta

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is d

iver

sifo

lia (P

arry

) Gre

ene

ssp.

pla

nifo

lia (J

eps.

) G.D

. Wal

lace

CA

TR

OS

CR

UA

NA

Vacc

iniu

m o

vatu

m P

ursh

RO

SC

RU

Xyl

ococ

cus

bico

lor

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t.C

AT

Eup

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ae

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ulat

a (L

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all*

CL

E14

3B

AR

CR

UA

NA

Cha

mae

syce

ser

pens

(Kun

th) S

mal

l*C

LE

Cha

mae

syce

ser

pylli

folia

(Per

s.) S

mal

lC

AT

Cro

ton

setig

erus

Hoo

k.C

LE

CA

TR

OS

CR

UE

upho

rbia

cre

nula

ta E

ngel

m.

CA

TE

upho

rbia

mis

era

Ben

th.

CL

EC

AT

CR

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upho

rbia

pep

lus

L.*

CL

EC

AT

RO

SC

RU

Eup

horb

ia s

path

ulat

a L

am.

CL

EC

AT

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inus

com

mun

is L

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LE

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CA

TSt

illin

gia

linea

rifo

lia S

. Wat

son

CR

UFa

bace

aeA

caci

a cy

clop

s G

. Don

*C

LE

? 144

139 R

SA: F

.H. E

lmor

e Fe

b 19

193

9 42

214

0 SB

BG

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. Dun

kle

Jul 2

6 19

39 8

359

141 S

D: D

arle

y F.

How

e Fe

b 24

, 196

8 44

8614

2 SB

BG

: M.B

. Dun

kle

Jul 3

1 19

39 8

402

143 R

SA: S

teve

n A

. Jun

ak N

ov 9

199

0 SC

I144

144 P

ers.

com

. E. H

owe

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208 MONOGRAPHS OF THE WESTERN NORTH AMERICAN NATURALIST [Volume 7A

PPE

ND

IX2.

Con

tinue

d

Fam

ily/s

peci

esss

p./v

ar. a

nd in

fran

ame

CL

EN

ICC

AT

BA

RR

OS

CR

UA

NA

MIG

Aca

cia

decu

rren

s W

illd.

*C

AT

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cia

deal

bata

Lin

k*C

RU

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cia

mel

anox

ylon

R. B

r.*C

AT

CR

UA

cmis

pon

amer

ican

us (N

utt.)

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eric

anus

CA

TC

RU

Acm

ispo

n ar

goph

yllu

s (A

. Gra

y) B

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llet

var.

adsu

rgen

s (D

unkl

e)C

LE

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ispo

n ar

goph

yllu

s (A

. Gra

y) B

roui

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enth

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ideu

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reen

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roid

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EC

AT

RO

SC

RU

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AA

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uille

tC

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n gr

andi

floru

s (B

enth

.) B

roui

llet

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gran

diflo

rus

CA

TR

OS

CR

UA

cmis

pon

heer

man

nii (

Dur

and

& H

ilg.)

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uille

tC

AT

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ispo

n m

ariti

mus

(Nut

t.) D

.D. S

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off1

49va

r. m

ariti

mus

NIC

CA

TR

OS

CR

UA

NA

MIG

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ispo

n m

icra

nthu

s (T

orr.

& A

. Gra

y) B

roui

llet1

50C

LE

CA

TR

OS

CR

UA

cmis

pon

parv

iflor

us (B

enth

.) D

.D. S

okol

off1

51C

RU

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ispo

n st

rigo

sus

(Nut

t.) B

roui

llet

CL

EC

AT

RO

SC

RU

AN

AM

IGA

cmis

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isch

. & C

.A. M

ey.)

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TR

OS

CR

UA

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D.D

. Sok

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th.*

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es A

. Gra

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OS

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raga

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didy

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s H

ook.

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LE

NIC

CA

TR

OS

CR

UA

NA

MIG

Ast

raga

lus

gam

belia

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held

.C

AT

CR

UA

stra

galu

s m

igue

lens

is G

reen

eC

LE

RO

SC

RU

AN

AM

IGA

stra

galu

s ne

vini

i A. G

ray

CL

EA

stra

galu

s tr

aski

ae E

astw

.N

ICB

AR

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raga

lus

tric

hopo

dus

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t.) A

. Gra

yva

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nchu

s (M

.E. J

ones

) Bar

neby

CA

TR

OS

CR

UA

NA

Ast

raga

lus

tric

hopo

dus

(Nut

t.) A

. Gra

yva

r. tr

icho

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sC

AT

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salp

inia

spi

nosa

(Mol

ina)

Kun

tze*

CA

T15

3

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onill

a va

lent

ina

L.*

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TG

enis

ta li

nifo

lia L

.*C

AT

Gen

ista

mon

spes

sula

na (L

.) L

.A.S

. Joh

nson

*C

AT

145 S

ynon

ym: L

otus

pur

shia

nus

Cle

m. &

E.G

. Cle

m.

146 C

AS:

L.R

. Abr

ams,

I.L

. Wig

gins

30

June

, 193

1 23

914

7 DS:

E.R

. Bla

kley

3 A

pril

1962

508

114

8 Syn

onym

: Lot

us h

umis

trat

usG

reen

e; W

alla

ce li

sted

L. h

umis

trat

uson

Cru

z, b

ut N

PS P

ark

Flo

ra d

id n

ot14

9 Syn

onym

: Lot

us s

alsu

gino

sus

Gre

ene

150 S

ynon

ym: L

otus

ham

atus

Gre

ene

151 S

ynon

ym: L

otus

mic

rant

hus

Ben

th.

152 S

ynon

ym: L

otus

wra

ngel

ianu

sFi

sch.

& C

.A. M

ey.;

Lot

us s

ubpi

nnat

usL

ag.,

mis

appl

.15

3 RSA

: Tim

Ros

s A

pr 2

3 19

93 6

934

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2014] CALIFORNIA ISLAND PLANT DISTRIBUTION PATTERNS 209A

PPE

ND

IX2.

Con

tinue

d

Fam

ily/s

peci

esss

p./v

ar. a

nd in

fran

ame

CL

EN

ICC

AT

BA

RR

OS

CR

UA

NA

MIG

Lat

hyru

s od

orat

us L

.*C

LE

154

Lat

hyru

s tin

gita

nus

L.*

CA

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itus

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r. al

efel

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T.G

. Whi

te) I

sley

CA

TL

athy

rus

vest

itus

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t.va

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stitu

sC

LE

CA

TR

OS

CR

UA

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us c

orni

cula

tus

L.*

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TC

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inus

alb

ifron

s B

enth

.va

r. do

ugla

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J. A

gard

h) C

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m.

NIC

CA

TR

OS

CR

UA

NA

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inus

arb

oreu

s Si

ms

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SC

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bic

olor

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dl.

CL

EN

ICC

AT

RO

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AM

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AT

RO

SC

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inus

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is G

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CA

TR

OS

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UA

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mic

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sR

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us s

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us K

. Koc

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CA

TR

OS

CR

UA

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inus

trun

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EC

AT

RO

SC

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us v

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ud.

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AM

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OS

CR

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EN

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AT

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ilotu

s in

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s (L

.) A

ll.*

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ICC

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BA

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OS

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UA

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erin

gia

mon

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m s

ativ

um L

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RU

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inia

pse

udoa

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a L

.*N

ICC

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tium

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eum

L.*

NIC

CA

TC

RU

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lium

alb

opur

pure

um T

orr.

& A

. Gra

yN

IC15

6C

AT

RO

SC

RU

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lium

bar

bige

rum

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r.R

OS

MIG

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lium

cili

olat

um B

enth

.C

AT

RO

SC

RU

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lium

dep

aupe

ratu

m D

esv.

var.

ampl

ecte

ns (T

orr.

& A

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y) M

cDer

mot

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LE

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epau

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tum

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v.va

r. tr

unca

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ene)

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lyC

LE

NIC

CA

TR

OS

CR

UA

NA

MIG

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lium

fuca

tum

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dl.

CL

EC

AT

RO

SC

RU

AN

AM

IGTr

ifoliu

m g

raci

lent

um T

orr.

& A

. Gra

yC

LE

NIC

CA

TB

AR

RO

SC

RU

AN

AM

IGTr

ifoliu

m h

irtu

m A

ll.C

LE

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lium

mac

raei

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k. &

Arn

.C

AT

RO

SC

RU

Trifo

lium

mic

roce

phal

um P

ursh

CL

EN

ICC

AT

RO

SC

RU

AN

AM

IGTr

ifoliu

m m

icro

don

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k. &

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.N

ICC

AT

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lium

pal

mer

i S. W

atso

nC

LE

NIC

CA

TB

AR

AN

ATr

ifoliu

m r

epen

s L

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AT

154 S

D: S

teve

n A

. Jun

ak M

ay 1

9, 1

997

SCl8

7715

5 SB

BG

: M.R

. Ben

edic

t Jun

26

1971

15

6 JE

PS93

564

(SN

-846

) det

. var

. dic

hoto

mum

(= T

. dic

hoto

mum

in T

JM2)

by

Mic

hael

Vin

cent

in 2

005.

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210 MONOGRAPHS OF THE WESTERN NORTH AMERICAN NATURALIST [Volume 7A

PPE

ND

IX2.

Con

tinue

d

Fam

ily/s

peci

esss

p./v

ar. a

nd in

fran

ame

CL

EN

ICC

AT

BA

RR

OS

CR

UA

NA

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lium

var

iega

tum

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t.va

r. m

ajor

Loj

a.C

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lium

will

deno

vii S

pren

g.C

LE

157

CA

T15

8B

AR

RO

SC

RU

AN

AM

IGV

icia

am

eric

ana

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d.R

OS

CR

UM

IGV

icia

ben

ghal

ensi

s L

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AT

159

CR

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Vic

ia h

asse

i S. W

atso

nC

LE

NIC

CA

TR

OS

CR

UA

NA

MIG

Vic

ia lu

dovi

cian

a To

rr. &

A. G

ray

ssp.

ludo

vici

ana

CL

E16

0C

AT

161

RO

SC

RU

AN

AV

icia

sat

iva

L.*

ssp.

nig

ra (L

.) E

rhar

tR

OS

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ia s

ativ

a L

.*ss

p. s

ativ

aN

ICR

OS

CR

UA

NA

Vic

ia v

illos

a R

oth

(Hos

t) C

orb.

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CA

T16

2

Que

rcus

agr

ifolia

Née

var.

agri

folia

RO

SC

RU

Que

rcus

chr

ysol

epis

Lie

bm.

CL

EC

AT

RO

SC

RU

AN

AQ

uerc

us d

ougl

asii

Hoo

k. &

Arn

.C

AT

CR

UQ

uerc

us e

ngel

man

nii G

reen

eC

AT

RO

SQ

uerc

us k

ello

ggii

New

b.C

RU

Que

rcus

loba

ta N

éeC

AT

RO

SC

RU

Que

rcus

×m

acdo

nald

ii G

reen

eC

AT

RO

SC

RU

Que

rcus

pac

ifica

Nix

on &

C.H

. Mul

l.C

AT

RO

SC

RU

Que

rcus

par

vula

Gre

ene

var.

parv

ula

CR

UQ

uerc

us to

men

tella

Eng

elm

.C

LE

CA

TR

OS

CR

UA

NA

Fran

keni

acea

eFr

anke

nia

salin

a (M

olin

a) I

.M. J

ohns

t.C

LE

NIC

CA

TR

OS

CR

UA

NA

MIG

Gar

ryac

eae

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rya

veat

chii

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logg

CR

UG

entia

nace

aeZ

eltn

era

davy

i (Je

ps.)

G. M

ans.

CL

ER

OS

CR

UZ

eltn

era

venu

sta

(A. G

ray)

G. M

ans.

NIC

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TG

eran

iace

aeC

alifo

rnia

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roph

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k. &

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. Ald

asor

oet

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3

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dium

bot

rys

(Cav

.) B

erol

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LE

NIC

CA

T16

4R

OS

CR

UE

rodi

um b

rach

ycar

pum

(God

r.) T

hell.

*C

LE

165

CA

T16

6R

OS

Ero

dium

cic

utar

ium

(L.)

Aito

n*C

LE

NIC

CA

TB

AR

RO

SC

RU

AN

AM

IGE

rodi

um m

osch

atum

(L.)

Aito

n*C

LE

NIC

CA

TB

AR

RO

SC

RU

AN

AM

IG

157 R

SA: P

.H. R

aven

May

8 1

962

1761

415

8 UC

: F.R

. Fos

berg

Mar

25

1931

s43

6715

9 SB

BG

: P.H

. Rav

en M

ay 2

0 19

62 1

7789

160 U

CR

: Ste

ve B

oyd,

T.S

. Ros

s, L

. Arn

seth

Apr

7 1

990

4260

161 S

BB

G: R

.F. T

horn

e M

ar 7

196

6 35

954

162 S

D: R

ober

t F. T

horn

e, D

. Pro

pst M

ay 2

9, 1

968

3769

116

3 GH

: W.A

. Wal

lace

164 S

BB

G: S

.A. J

unak

, K. K

irkl

and,

L. V

orob

ik M

ay 1

6 19

98 S

Ca-

697

165 S

BB

G: T

. Ros

s, O

. Mis

tret

ta, M

. Ham

mitt

May

18

1991

519

116

6 SB

BG

: S.A

. Jun

ak, M

.L. H

oefs

, K. K

irkl

and

Apr

5 1

998

SCa-

549

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2014] CALIFORNIA ISLAND PLANT DISTRIBUTION PATTERNS 211A

PPE

ND

IX2.

Con

tinue

d

Fam

ily/s

peci

esss

p./v

ar. a

nd in

fran

ame

CL

EN

ICC

AT

BA

RR

OS

CR

UA

NA

MIG

Ero

dium

texa

num

A. G

ray

CA

TG

eran

ium

car

olin

ianu

m L

.C

AT

RO

SC

RU

Ger

aniu

m d

isse

ctum

L.*

RO

SG

eran

ium

mol

le L

.*C

RU

Pela

rgon

ium

×ho

rtor

um L

.H. B

aile

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LE

167

NIC

CA

TR

OS

CR

UM

IG16

8

Pela

rgon

ium

pel

tatu

m (L

.) L’

Hér

.*N

ICG

ross

ular

iace

ae

Rib

es m

alva

ceum

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.va

r. m

alva

ceum

CL

ER

OS

CR

UA

NA

Rib

es v

ibur

nifo

lium

A. G

ray

CA

TR

ibes

thac

heri

anum

(Jep

s.) M

unz

CR

UJu

glan

dace

ae

Jugl

ans

calif

orni

ca S

. Wat

son*

CA

TJu

glan

s re

gia

L.*

CR

UL

amia

ceae

C

linop

odiu

m d

ougl

asii

(Ben

th.)

Kun

tze

CA

TR

OS

CR

UL

amiu

m a

mpl

exic

aule

L.*

CR

UL

epec

hini

a fr

agra

ns (G

reen

e) E

plin

gC

AT

RO

SC

RU

Mar

rubi

um v

ulga

re L

.*C

LE

NIC

CA

TR

OS

CR

UM

IGM

enth

a aq

uatic

a L

.*C

AT

Men

tha

spic

ata

L.*

CA

TC

RU

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eta

cata

ria

L.*

CA

TSa

lvia

api

ana

Jeps

.C

AT

Salv

ia b

rand

egee

i Mun

zR

OS

Salv

ia c

olum

bari

ae B

enth

.C

LE

CA

TR

OS

CR

USa

lvia

leuc

ophy

lla G

reen

eC

RU

Salv

ia m

ellif

era

Gre

ene

CL

E? 1

69C

AT

CR

UA

NA

Scut

ella

ria

tube

rosa

Ben

th.

RO

SC

RU

Stac

hys

ajug

oide

s B

enth

.M

IGSt

achy

s bu

llata

Ben

th.

RO

SC

RU

AN

ATr

icho

stem

a la

nceo

latu

m B

enth

.C

AT

Lin

acea

eH

espe

rolin

on m

icra

nthu

m (A

. Gra

y) S

mal

lC

AT

Loa

sace

ae

Men

tzel

ia a

ffini

s G

reen

eC

LE

NIC

CA

TR

OS

CR

UM

entz

elia

mic

rant

ha (H

ook.

& A

rn.)

Torr

. & A

.C

LE

CA

TC

RU

Lyt

hrac

eae

Am

man

nia

robu

sta

Hee

r &

Reg

elC

AT

167 S

BB

G: T

. Ros

s M

ay 2

2 19

91 5

435

168 S

BB

G: R

.N. P

hilb

rick

Mar

25

1966

B66

-91

169 P

ers.

com

. E. H

owe

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212 MONOGRAPHS OF THE WESTERN NORTH AMERICAN NATURALIST [Volume 7A

PPE

ND

IX2.

Con

tinue

d

Fam

ily/s

peci

esss

p./v

ar. a

nd in

fran

ame

CL

EN

ICC

AT

BA

RR

OS

CR

UA

NA

MIG

Lyt

hrum

cal

iforn

icum

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r. &

A. G

ray

CR

UL

ythr

um h

ysso

pifo

lia L

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LE

NIC

CA

T17

0R

OS

CR

UM

alva

ceae

A

lcea

ros

ea L

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AT

CR

UE

rem

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e ex

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ray)

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avat

era

assu

rgen

tiflo

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p. a

ssur

gent

iflor

a C

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IC*

RO

SC

RU

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AM

IGL

avat

era

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EC

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CR

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acot

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orr.

& A

. Gra

y) G

reen

eva

r. ne

siot

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(B.L

. Rob

.) K

earn

eyC

RU

Mal

va p

seud

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ater

a W

ebb

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erth

el.*

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E17

1R

OS

AN

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alva

nic

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sis

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OS

CR

UM

alva

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viflo

ra L

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LE

NIC

CA

TB

AR

RO

SC

RU

AN

AM

IGM

alve

lla le

pros

a (O

rteg

a) K

rapo

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LE

CA

TR

OS

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lcea

mal

viflo

ra (D

C.)

A. G

ray

ssp.

mal

viflo

raR

OS

CR

UM

IGM

ontia

ceae

C

alan

drin

ia b

rew

eri S

. Wat

son

RO

SC

RU

Cal

andr

inia

cili

ata

Fisc

h. &

C.A

. Mey

.C

LE

NIC

CA

TB

AR

RO

SC

RU

AN

AM

IGC

ista

nthe

mar

itim

a (N

utt.)

Her

shk.

CL

EC

AT

BA

RR

OS

CR

UA

NA

Cla

yton

ia p

arvi

flora

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k.ss

p. p

arvi

flora

NIC

CA

T17

2B

AR

CR

UM

IGC

layt

onia

per

folia

ta W

illd.

ssp.

mex

ican

a (R

ydb.

) Joh

n M

. Mill

. C

LE

173

NIC

CA

T17

4B

AR

RO

SC

RU

AN

AM

IG&

K.L

. Cha

mbe

rsC

layt

onia

per

folia

ta W

illd.

ssp.

per

folia

taC

LE

CA

TR

OS

CR

UM

ontia

font

ana

L.

CA

TC

RU

Mor

acea

e Fi

cus

cari

ca L

.*C

LE

175

CA

TC

RU

MIG

Myr

sina

ceae

A

naga

llis

arve

nsis

L.*

CL

EN

ICC

AT

BA

RR

OS

CR

UA

NA

MIG

Ana

galli

s m

inim

a (L

.) E

.H.L

. Kra

use

RO

SM

yrta

ceae

E

ucal

yptu

s ca

mal

dule

nsis

Deh

nh.*

CA

T17

6R

OS

CR

UE

ucal

yptu

s cl

adoc

alyx

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uell.

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AT

177

Euc

alyp

tus

glob

ulus

Lab

ill.*

CL

EC

AT

BA

RR

OS

CR

UA

NA

170 R

SA: M

ark

Hoe

fs, S

teve

n A

. Jun

ak, J

anet

Tak

ara,

M. G

ay J

ul 1

4 19

95 2

401

171 S

D: S

teve

n A

. Jun

ak A

pr 2

3, 1

997

SCI8

5817

2 SB

BG

: S.A

. Jun

ak, K

. Kir

klan

d A

pr 1

3 20

01 S

Ca-

1421

173 S

BB

G: T

im R

oss,

Orl

ando

Mis

tret

ta, M

ike

Ham

mitt

May

19

1991

528

617

4 SB

BG

: E.R

. Bla

kley

Apr

7 1

963

5537

175 S

BB

G: S

.A. J

unak

May

20

1996

176 U

CR

: T.S

. Ros

s A

pr 2

3 19

93 6

926

177 U

CR

: O.F

. Cla

rke

Feb

25 1

968

s.n.

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2014] CALIFORNIA ISLAND PLANT DISTRIBUTION PATTERNS 213A

PPE

ND

IX2.

Con

tinue

d

Fam

ily/s

peci

esss

p./v

ar. a

nd in

fran

ame

CL

EN

ICC

AT

BA

RR

OS

CR

UA

NA

MIG

Euc

alyp

tus

poly

anth

emos

Sch

auer

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AT

178

Euc

alyp

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side

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lon

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CR

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retic

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s Sm

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UN

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bron

ia la

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schs

ch.

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onia

mar

itim

a S.

Wat

son

CL

EN

ICC

AT

RO

SC

RU

AN

AM

IGA

bron

ia u

mbe

llata

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.C

LE

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TR

OS

CR

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enth

.) C

urra

nva

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edia

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TC

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ner

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och

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sis

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rant

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pren

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EC

AT

RO

SC

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AN

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agne

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aven

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LE

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ner

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eps.

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stra

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is &

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pure

a (C

urtis

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n &

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qua

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ulne

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indl

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AT

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H. L

ewis

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AT

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178 U

CR

: T.S

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s, J

anet

Tak

ara,

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cey

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e A

pr 2

4 19

93 6

964

179 P

ers.

com

. E. H

owe

180 T

his

taxa

unr

esol

ved

181 S

DSU

: R.M

. Bea

ucha

mp

Aug

ust 8

, 196

6 54

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214 MONOGRAPHS OF THE WESTERN NORTH AMERICAN NATURALIST [Volume 7A

PPE

ND

IX2.

Con

tinue

d

Fam

ily/s

peci

esss

p./v

ar. a

nd in

fran

ame

CL

EN

ICC

AT

BA

RR

OS

CR

UA

NA

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Epi

lobi

um c

iliat

um R

af.

ssp.

cili

atum

CA

TC

RU

MIG

Eul

obus

cal

iforn

icus

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r. &

A. G

ray

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TR

OS

CR

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igia

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loid

es (K

unth

) P.H

. Rav

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ides

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ogau

ra W

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agne

r &

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h*C

RU

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othe

ra s

inuo

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agne

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th.

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elle

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OS

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m. &

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icul

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t.C

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TR

OS

CR

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banc

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aris

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.) H

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rach

ylob

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AT

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SC

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arfo

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logg

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EC

AT

RO

SC

RU

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endo

rfer

ia o

chro

leuc

a (E

ngel

m.)

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chol

zia

calif

orni

ca C

ham

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LE

184

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OS

CR

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ticul

ata

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yR

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tero

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th.)

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EC

AT

BA

RR

OS

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mon

cal

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icus

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th.

NIC

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TB

AR

RO

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AN

AM

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omne

ya c

oulte

ri H

arv.

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AT

182 P

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com

. E. H

owe

183 R

SA: S

teve

n A

. Jun

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ay 1

4 19

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198

3

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2014] CALIFORNIA ISLAND PLANT DISTRIBUTION PATTERNS 215A

PPE

ND

IX2.

Con

tinue

d

Fam

ily/s

peci

esss

p./v

ar. a

nd in

fran

ame

CL

EN

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RR

OS

CR

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Phry

mac

eae

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ulus

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185

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orus

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ene)

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. Tho

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us a

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Mim

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ttos

pora

ceae

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poru

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ndul

atum

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AT

188

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nace

ae

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irrh

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reen

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um m

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m P

enne

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nut

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nata

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sia

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ham

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ER

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cios

a N

utt.

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EC

AT

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RK

ecki

ella

cor

difo

lia (B

enth

.) St

raw

CL

EC

AT

RO

SC

RU

AN

AK

ickx

ia e

latin

e (L

.) D

umor

t.*C

AT

189

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ria

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rtita

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t.) W

illd.

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s te

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s (S

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le)

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AM

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utto

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TA

NA

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elo

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RO

SC

RU

MIG

Plan

tago

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LE

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TR

OS

CR

UA

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tago

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CA

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AT

CR

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go m

ariti

ma

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anta

go o

vata

For

ssk.

var.

insu

lari

s (E

astw

.) S.

C. M

eyer

s &

A. L

isto

n C

LE

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AR

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SC

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AM

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anta

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SC

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BI:

Joh

n K

napp

, Den

ise

Kna

pp, 7

, Apr

il 13

, 200

318

6 SB

BG

: S. B

oyd,

T. R

oss,

L. A

rnse

th A

pr 8

199

0 42

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e M

. Tho

mps

on 2

012.

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ulus

, in

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on F

lora

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ject

(eds

.) Je

pson

eF

lora

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esse

d on

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7 2

014

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H: R

.F. T

horn

e &

A.D

. Pro

pst 1

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09-1

2 45

100

189 S

BB

G: M

.L. H

oefs

, R.F

. Tho

rne,

J. T

akar

a Ju

l 25

1995

243

819

0 SB

BG

: M.L

. Hoe

fs, R

.F. T

horn

e A

pr 6

199

6 26

71

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216 MONOGRAPHS OF THE WESTERN NORTH AMERICAN NATURALIST [Volume 7A

PPE

ND

IX2.

Con

tinue

d

Fam

ily/s

peci

esss

p./v

ar. a

nd in

fran

ame

CL

EN

ICC

AT

BA

RR

OS

CR

UA

NA

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nace

ae

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itim

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ill.)

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d.ss

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oiss

.) A

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ium

sin

uatu

m (L

.) M

ill.*

CA

TA

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ae

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phyl

lum

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enth

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trum

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ilia

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. Gra

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Gili

a ca

pita

ta S

ims

ssp.

abr

otan

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ene)

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ilia

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orum

(Jep

s.) V

.E. G

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TR

OS

CR

UA

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i A. G

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CL

EN

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CR

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enth

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OS

CR

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on p

ygm

aeus

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nd)

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pyg

mae

usC

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orte

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L.A

. Joh

nson

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anth

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us (B

enth

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enth

.) C

LE

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OS

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ook.

& A

rn.

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arre

tia h

amat

a G

reen

ess

p. le

ptan

tha

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ason

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EC

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gala

ceae

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lyga

la c

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utt.

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AT

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nthe

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eler

i S. W

atso

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ores

cens

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RO

SC

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AN

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m B

enth

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IC19

2*R

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scic

ulat

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r. fa

scic

ulat

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AT

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scic

ulat

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r. po

lifol

ium

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th.)

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. & A

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riog

onum

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um S

. Wat

son

var.

com

pact

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unkl

eB

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ogon

um g

igan

teum

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atso

nva

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nde

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gran

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LE

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NA

191 P

ers.

com

. E. H

owe

192 S

BB

G: S

.A. J

unak

, T. M

urph

ey, S

N-4

32, F

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1 19

9019

3 HSC

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rne,

D. P

rops

t, M

. Hae

fs S

ep 1

2 19

74 4

5110

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2014] CALIFORNIA ISLAND PLANT DISTRIBUTION PATTERNS 217A

PPE

ND

IX2.

Con

tinue

d

Fam

ily/s

peci

esss

p./v

ar. a

nd in

fran

ame

CL

EN

ICC

AT

BA

RR

OS

CR

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orum

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eal

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Las

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cor

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oodm

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icar

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path

ifolia

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ICC

AT

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OS

CR

UA

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ex a

ceto

sella

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SC

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Rum

ex c

ongl

omer

atus

Mur

ray*

CL

E19

4C

AT

RO

SC

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ex c

risp

us L

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NIC

CA

TR

OS

CR

UA

NA

MIG

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ex fu

egin

us P

hil.

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umex

obt

usifo

lius

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Rum

ex p

ulch

er L

.*C

RU

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ex s

alic

ifoliu

s W

einm

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NIC

CA

TR

OS

CR

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IGPo

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ulac

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erac

ea L

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AT

CR

UPr

imul

acea

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odec

athe

on c

leve

land

ii G

reen

ess

p. in

sula

re H

.J. T

hom

ps.

CL

EN

ICC

AT

RO

SC

RU

AN

AM

IGR

anun

cula

ceae

C

lem

atis

lasi

anth

a N

utt.

CR

UC

lem

atis

ligu

stic

ifolia

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t.C

AT

RO

SC

RU

Cle

mat

is p

auci

flora

Nut

t.C

RU

Del

phin

ium

par

ryi A

. Gra

yss

p. m

ariti

mum

(Dav

idso

n) M

.J. W

arno

ckC

AT

RO

SC

RU

AN

AM

IGD

elph

iniu

m p

arry

i A. G

ray

ssp.

par

ryi

CA

TC

RU

Del

phin

ium

var

iega

tum

Tor

r. &

A. G

ray

ssp.

kin

kien

se (M

unz)

M.J.

War

nock

CL

ED

elph

iniu

m v

arie

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m T

orr.

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. Gra

yss

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orne

i Mun

zC

LE

Ran

uncu

lus

calif

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cus

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th.

CA

TR

OS

CR

UM

IGR

anun

culu

s he

beca

rpus

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k. &

Arn

.C

AT

Res

edac

eae

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omer

is li

nifo

lia (H

orne

m.)

J.F. M

acbr

.C

LE

NIC

CA

TB

AR

RO

SC

RU

AN

AM

IGR

esed

a od

orat

a L

.*C

AT

Rha

mna

ceae

C

eano

thus

arb

oreu

s G

reen

eC

AT

RO

SC

RU

Cea

noth

us m

egac

arpu

s N

utt.

var.

insu

lari

s (E

astw

.) M

unz

CL

EC

AT

RO

SC

RU

AN

AM

IG&

Cea

noth

us m

egac

arpu

s N

utt.

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meg

acar

pus

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EC

AT

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angu

la c

alifo

rnic

a (E

schs

ch.)

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ray

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cal

iforn

ica

CR

UR

ham

nus

piri

folia

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ene

CL

EC

AT

RO

SC

RU

MIG

&R

osac

eae

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nost

oma

fasc

icul

atum

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k. &

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.va

r. fa

scic

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LE

CA

TR

OS

CR

U

194 S

D: S

teve

n A

. Jun

ak M

ay 1

8, 1

996

SCI4

88a

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218 MONOGRAPHS OF THE WESTERN NORTH AMERICAN NATURALIST [Volume 7A

PPE

ND

IX2.

Con

tinue

d

Fam

ily/s

peci

esss

p./v

ar. a

nd in

fran

ame

CL

EN

ICC

AT

BA

RR

OS

CR

UA

NA

MIG

Ade

nost

oma

fasc

icul

atum

Hoo

k. &

Arn

.va

r. pr

ostr

atum

Dun

kle

CA

TR

OS

CR

U

Aph

anes

occ

iden

talis

(Nut

t.) R

ydb.

CL

EC

AT

RO

SC

RU

Cer

coca

rpus

bet

uloi

des

Nut

t.va

r. be

tulo

ides

CA

TC

erco

carp

us b

etul

oide

s N

utt.

var.

blan

chea

e (C

.K. S

chne

id.)

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leC

AT

RO

SC

RU

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coca

rpus

tras

kiae

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tw.

CA

TD

rym

ocal

lis g

land

ulos

a (L

indl

.) R

ydb.

195

CA

TH

eter

omel

es a

rbut

ifolia

(Lin

dl.)

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oem

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LE

CA

TR

OS

CR

UA

NA

MIG

Hol

odis

cus

disc

olor

(Pur

sh) M

axim

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AT

CR

UL

yono

tham

nus

flori

bund

us A

. Gra

yss

p. a

sple

niifo

lius

(Gre

ene)

P.H

. Rav

enC

LE

RO

SC

RU

Lyo

noth

amnu

s flo

ribu

ndus

A. G

ray

ssp.

flor

ibun

dus

CA

TPo

tent

illa

anse

rina

L.

ssp.

pac

ifica

(How

ell)

Rou

siC

RU

MIG

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rium

san

guis

orba

L.*

CA

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6

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us il

icifo

lia (H

ook.

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ietr.

ss

p. ly

onii

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tw.)

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. Rav

enC

LE

CA

TR

OS

CR

UA

NA

Ros

a ca

lifor

nica

Cha

m. &

Sch

ltdl.

CA

TR

OS

CR

UR

ubus

urs

inus

Cha

m. &

Sch

ltdl.

CA

TR

OS

CR

UM

IGR

ubia

ceae

G

aliu

m a

ngus

tifol

ium

sub

sp.

ssp.

folio

sum

(Hile

nd &

J.T.

How

ell)

RO

SC

RU

AN

AD

emps

ter

& S

tebb

ins

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ium

ang

ustif

oliu

m A

. Gra

yss

p. a

ngus

tifol

ium

CA

TG

aliu

m a

pari

ne L

.C

LE

NIC

CA

TB

AR

RO

SC

RU

AN

AM

IGG

aliu

m b

uxifo

lium

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ene

CR

UM

IGG

aliu

m c

alifo

rnic

um H

ook.

& A

rn.

ssp.

flac

cidu

m (G

reen

e) D

emps

ter

& S

tebb

ins

CR

UG

aliu

m c

alifo

rnic

um H

ook.

& A

rn.

ssp.

mig

uele

nse

(Gre

ene)

R

OS

MIG

Dem

pste

r &

Ste

bbin

sG

aliu

m c

atal

inen

se A

. Gra

yss

p. a

cris

pum

Dem

pste

rC

LE

Gal

ium

cat

alin

ense

A. G

ray

ssp.

cat

alin

ense

CA

TG

aliu

m n

utta

llii A

. Gra

yss

p. in

sula

re F

erri

sC

AT

RO

SC

RU

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ium

par

isie

nse

L.*

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7R

OS

Gal

ium

por

rige

ns D

emps

ter

var.

porr

igen

sC

AT

RO

SC

RU

Rut

acea

e R

uta

chal

epen

sis

L.*

CA

TSa

licae

ae

Popu

lus

frem

ontii

S. W

atso

nss

p. fr

emon

tiiC

AT

CR

UPo

pulu

s tr

icho

carp

a H

ook.

CA

TR

OS

CR

USa

lix e

xigu

a N

utt.

NIC

CR

USa

lix g

oodd

ingi

i C.R

. Bal

lC

LE

Salix

laev

igat

a B

ebb

CA

TC

RU

195 S

ynon

ym: P

oten

tilla

gla

ndul

osa

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dl.

196 U

C: T

im R

oss,

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pr 2

2 19

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BG

: S.A

. Jun

ak, M

.L. H

oefs

, K. K

irkl

and

Jul 1

7 19

98 S

Ca-

769

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2014] CALIFORNIA ISLAND PLANT DISTRIBUTION PATTERNS 219A

PPE

ND

IX2.

Con

tinue

d

Fam

ily/s

peci

esss

p./v

ar. a

nd in

fran

ame

CL

EN

IC

CA

T

BA

R

RO

SC

RU

AN

AM

IG

Salix

lasi

andr

a B

enth

.va

r. la

sian

dra

CR

USa

lix la

siol

epis

Ben

th.

CL

E? 1

98N

ICC

AT

RO

SC

RU

AN

AM

IGSa

pind

acea

e A

cer

mac

roph

yllu

m P

ursh

CR

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xifr

agac

eae

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cher

a m

axim

a G

reen

eR

OS

CR

UA

NA

Jeps

onia

mal

vifo

lia (G

reen

e) S

mal

lC

LE

NIC

CA

TR

OS

CR

ULi

thop

hrag

ma

affin

e A

. Gra

yC

AT

Lith

ophr

agm

a cy

mba

lari

a To

rr. &

A. G

ray

RO

SC

RU

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ophr

agm

a m

axim

um B

acig

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LE

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rant

hes

calif

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reen

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mal

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OS

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ular

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ae

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201

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CR

U&

Scro

phul

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cal

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RO

SC

ham

. & S

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ular

ia v

illos

a Pe

nnel

lC

LE

CA

TVe

rbas

cum

thap

sus

L.*

CR

USo

lana

ceae

D

atur

a w

righ

tii R

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CL

E? 2

02*

CA

TR

OS

CR

UL

yciu

m b

revi

pes

Ben

th.

CL

EN

ICC

AT

&L

yciu

m c

alifo

rnic

um N

utt.

CL

EN

ICC

AT

BA

RC

RU

AN

AM

IGL

yciu

m fr

emon

tii A

. Gra

yR

OS

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ium

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ruco

sum

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tw.

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cum

esc

ulen

tum

Mill

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LE

? 203

NIC

CA

T20

4B

AR

&C

RU

AN

A&

MIG

&N

icot

iana

cle

vela

ndii

A. G

ray

CA

T20

5B

AR

CR

UA

NA

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otia

na g

lauc

a G

raha

m*

CL

E? 2

06N

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AT

CR

UN

icot

iana

qua

driv

alvi

s Pu

rsh

CA

T20

7

Petu

nia

parv

iflor

a Ju

ss.

RO

SSo

lanu

m a

mer

ican

um M

ill.*

CL

E20

8N

ICR

OS2

09A

NA

Sola

num

dou

glas

ii D

unal

CL

EN

ICC

AT

BA

RR

OS

CR

UA

NA

MIG

198 P

ers.

com

. E. H

owe

199 S

ynon

ym: S

axifr

aga

calif

orni

caG

reen

e20

0 SB

BG

: S.A

. Jun

ak M

ar 1

3 19

97 S

Cl-

770

201 S

BB

G: S

.A. J

unak

Jul

31

1981

SC

l-53

202 P

ers.

com

. E. H

owe

203 P

ers.

com

. E. H

owe

204 R

SA: R

.F. T

horn

e, 4

7381

, Sep

4 1

975

205 R

SA: B

lanc

he T

rask

Mar

190

1 s.

n.20

6 Per

s. c

om. E

. How

e20

7 SD

: Rob

ert F

. Tho

rne,

P. E

vere

tt, 3

4883

, Jun

22,

196

520

8 SB

BG

: H.L

. Fer

guso

n, R

.M. B

eauc

ham

p Se

p 15

197

9 20

9 SB

BG

: S.A

. Jun

ak S

ep 2

3 19

98 S

R-8

96

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220 MONOGRAPHS OF THE WESTERN NORTH AMERICAN NATURALIST [Volume 7A

PPE

ND

IX2.

Con

tinue

d

Fam

ily/s

peci

esss

p./v

ar. a

nd in

fran

ame

CL

EN

IC

CA

T

BA

R

RO

SC

RU

AN

AM

IG

Sola

num

ela

eagn

ifoliu

m C

av.*

CA

TR

OS

CR

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lanu

m w

alla

cei (

A. G

ray)

Par

ish

CA

TR

OS

CR

UTa

mar

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eae

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arix

aph

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(L.)

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arst

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RO

SC

RU

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ia h

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ra H

into

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LE

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CA

TB

AR

RO

SC

RU

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AM

IGSo

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sol

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lii (R

eq.)

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ca L

.ss

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TR

OS

CR

UA

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s L

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RO

SC

RU

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uber

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sa (G

reen

e) J

eps.

CR

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e Ph

yla

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ene*

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3C

AT

CR

U21

4

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ena

brac

teat

a L

ag. &

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r.C

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215

CA

TR

OS

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ena

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osta

chys

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kva

r. la

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tach

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LE

NIC

CA

TVe

rben

a la

sios

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ys L

ink

var.

scab

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Mol

denk

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RO

SC

RU

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acea

e V

iola

ped

uncu

lata

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r. &

A. G

ray

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EC

AT

RO

SC

RU

Vita

ceae

V

itis

gird

iana

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son

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TZ

ygop

hylla

ceae

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ibul

us te

rres

tris

L.*

CA

T21

6

NU

MB

ER

OF

SPE

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627

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315

051

966

227

129

6

210 P

ers.

com

. E. H

owe

211 R

SA: R

.F. T

horn

e, 4

5759

, Mar

22

1975

212 U

CR

: T.S

. Ros

s, 5

434,

May

22

1991

213 S

BB

G: H

.L. F

ergu

son

Mar

12

1981

234

214 S

BB

G: S

.A. J

unak

, R.N

. Phi

lbri

ck, M

.C. H

ochb

erg,

SC

-63,

May

9 1

979

215 R

SA: E

. Kel

logg

Jun

28

1992

s.n

.21

6 SB

BG

: S.A

. Jun

ak, K

. Kir

klan

d, M

. Bus

hman

Oct

21

1998

SC

a-81

5

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