UNITED STATES SECURITIES AND EXCHANGE...

The Securities and Exchange Commission has not necessarily reviewed the information in this filing and has not determined if itis accurate and complete.

The reader should not assume that the information is accurate and complete.

UNITED STATES SECURITIES AND EXCHANGE COMMISSION OMB APPROVAL

Washington, D.C. 20549 FORM 13F

OMB Number: 3235-0006 Expires: July 31, 2015 Estimated average burden hours per response: 23.8

FORM 13F COVER PAGE

Report for the Calendar Year or Quarter Ended: 06-30-2019 Check here if Amendment: Amendment Number: This Amendment (Check only one.): is a restatement.

adds new holdings entries.

Institutional Investment Manager Filing this Report:

Name: Invesco Ltd.

Address: 1555 Peachtree Street NE

Suite 1800

Atlanta GA 30309

Form 13F File Number: 028-11246

The institutional investment manager filing this report and the person by whom it is signed hereby represent that the person signing the report isauthorized to submit it, that all information contained herein is true, correct and complete, and that it is understood that all required items,statements, schedules, lists, and tables, are considered integral parts of this form.

Person Signing this Report on Behalf of Reporting Manager:

Name: Nancy L. Tomassone Title: Global Assurance Officer

Phone: 404-479-2926

Signature, Place, and Date of Signing:

Nancy L. Tomassone, Global Assurance Officer Atlanta, GEORGIA 08-14-2019

[Signature] [City, State] [Date]

Do you wish to provide information pursuant to Special Instruction 5? Yes X No

Report Type (Check only one.):

X 13F HOLDINGS REPORT. (Check here if all holdings of this reporting manager are reported in this report.)

13F NOTICE. (Check here if no holdings reported are in this report, and all holdings are reported by other reporting manager(s).)

13F COMBINATION REPORT. (Check here if a portion of the holdings for this reporting manager are reported in this report and a portion arereported by other reporting manager(s).)

Form 13F Summary Page

Report Summary:

Number of Other Included Managers: 13

Form 13F Information table Entry Total: 12289

Form 13F Information table Value Total: 419,132,888 (thousands)

List of Other Included Managers:

Provide a numbered list of the name(s) and Form 13F number(s) of all institutional investement managers with respect to which this report is filed,other than the manager filing this report.

No. Form 13F File Number Name CIK

1 028-00255 Invesco Advisers, Inc.

2 028-06137 Invesco Canada Ltd

3 028-12270 Invesco Trust Co.

4 Invesco Hong Kong Limited

5 Invesco Asset Management Deutschland GmbH

6 028-05683 Invesco Asset Management Ltd

8 Invesco Asset Management Osterreich GmbH

9 Invesco Management S.A.

11 Invesco Asset Management Japan Limited

12 Invesco Global Asset Management DAC

13 Invesco Australia Ltd.

14 028-11365 Invesco Investment Advisers, LLC

15 028-11936 Invesco Capital Management LLC

FORM 13F INFORMATION TABLE

NAME OF ISSUER TITLE OF CLASS CUSIP VALUE (x$1000)SHRS OR PRN AMT

SH / PRN

PUT / CALL

INVESTMENT DISCRETION

OTHER MANAGER

VOTING AUTHORITY

AAC HLDGS INC COM 000307108 68 78,508 SH - DFND 15 0 78,508 0

AAON INC COM PAR $0.004 000360206 46 923 SH - DFND 1,15 0 923 0

AAON INC COM PAR $0.004 000360206 5,056 100,753 SH - DFND 15 0 100,753 0

AAR CORP COM 000361105 330 8,966 SH - DFND 14 0 8,966 0

AAR CORP COM 000361105 6,566 178,473 SH - DFND 15 0 178,473 0

ABB LTD SPONSORED ADR 000375204 13,999 698,905 SH - DFND 2 0 698,905 0



ACCO BRANDS CORP COM 00081T108 25,190 3,200,728 SH - DFND 1 0 3,200,728 0

ACCO BRANDS CORP COM 00081T108 3,082 391,647 SH - DFND 15 0 391,647 0

ACNB CORP COM 000868109 286 7,240 SH - DFND 1 0 7,240 0

ACNB CORP COM 000868109 63 1,600 SH - DFND 1,3 0 1,600 0

ACNB CORP COM 000868109 113 2,856 SH - DFND 5 0 0 2,856

ADT INC COM 00090Q103 1,471 240,298 SH - DFND 15 0 240,298 0

ABM INDS INC COM 000957100 14,024 350,593 SH - DFND 1 0 350,593 0

ABM INDS INC COM 000957100 1,107 27,673 SH - DFND 1,11 0 27,673 0



ABM INDS INC COM 000957100 1,026 25,652 SH - DFND 6,2 0 25,652 0

ABM INDS INC COM 000957100 655 16,368 SH - DFND 6,9 0 16,368 0

AFLAC INC COM 001055102 66,403 1,211,514 SH - DFND 14 0 1,211,514 0

AFLAC INC COM 001055102 44 800 SH Call DFND 14 0 0 0

AFLAC INC COM 001055102 132 2,400 SH Call DFND 14 0 0 0

AFLAC INC COM 001055102 219 4,000 SH Call DFND 14 0 0 0

AFLAC INC COM 001055102 23,267 424,497 SH - DFND 1 0 406,247 18,250

AFLAC INC COM 001055102 2,033 37,100 SH - DFND 1,4 0 37,100 0

AFLAC INC COM 001055102 176 3,220 SH - DFND 1,9 0 3,220 0



AFLAC INC COM 001055102 212,357 3,874,417 SH - DFND 15 0 3,855,435 18,982

AFLAC INC COM 001055102 32,145 586,488 SH - DFND 5 0 356,056 230,432



AFLAC INC COM 001055102 6,079 110,911 SH - DFND 5,6 0 16,284 94,627

AFLAC INC COM 001055102 202 3,688 SH - DFND 11 0 3,688 0

AFLAC INC COM 001055102 914 16,668 SH - DFND 6 0 16,668 0

AFLAC INC COM 001055102 5,615 102,436 SH - DFND 13 0 102,436 0

AGCO CORP COM 001084102 2,188 28,201 SH - DFND 14 0 28,201 0



AGCO CORP COM 001084102 28 355 SH - DFND 5 0 0 355

AGCO CORP COM 001084102 497 6,408 SH - DFND 5,6 0 627 5,781

AG MTG INVT TR INC COM 001228105 2,069 130,100 SH - DFND 1 0 91,600 38,500

AG MTG INVT TR INC COM 001228105 196 12,300 SH - DFND 1,3 0 12,300 0

AG MTG INVT TR INC COM 001228105 11,808 742,658 SH - DFND 15 0 742,658 0

AGNC INVT CORP COM 00123Q104 555 32,983 SH - DFND 14 0 32,983 0

AGNC INVT CORP COM 00123Q104 33,574 1,996,083 SH - DFND 1 0 1,996,083 0

AGNC INVT CORP COM 00123Q104 12,564 746,943 SH - DFND 15 0 746,943 0

AGNC INVT CORP COM 00123Q104 612 36,381 SH - DFND 5,6 0 4,307 32,074



AES CORP COM 00130H105 210 12,531 SH - DFND 2,15 0 12,531 0

AES CORP COM 00130H105 271 16,155 SH - DFND 14 0 16,155 0

COLUMN 1 COLUMN 2 COLUMN 3 COLUMN 4 COLUMN 5 COLUMN 6 COLUMN 7 COLUMN 8

SOLE SHARED NONE

AES CORP COM 00130H105 15,724 938,213 SH - DFND 1 0 938,213 0

AES CORP COM 00130H105 1,489 88,829 SH - DFND 1,15 0 88,829 0

AES CORP COM 00130H105 110,538 6,595,350 SH - DFND 15 0 6,595,350 0

AES CORP COM 00130H105 594 35,438 SH - DFND 5 0 0 35,438

AES CORP COM 00130H105 54 3,216 SH - DFND 11 0 3,216 0

AK STL HLDG CORP COM 001547108 5,112 2,157,060 SH - DFND 15 0 2,157,060 0

ALPS ETF TR ALERIAN MLP 00162Q866 4,028 408,934 SH - DFND 14 0 408,934 0

AMAG PHARMACEUTICALS INC COM 00163U106 100 10,000 SH - DFND 1 0 10,000 0

AMAG PHARMACEUTICALS INC COM 00163U106 3,148 315,150 SH - DFND 15 0 315,150 0

AMC NETWORKS INC CL A 00164V103 5,098 93,559 SH - DFND 1 0 93,559 0

AMC NETWORKS INC CL A 00164V103 10,615 194,812 SH - DFND 15 0 194,812 0

AMC NETWORKS INC CL A 00164V103 883 16,202 SH - DFND 5 0 16,202 0

AMC NETWORKS INC CL A 00164V103 92 1,681 SH - DFND 5,9 0 1,681 0

AMC ENTMT HLDGS INC CL A COM 00165C104 150 16,119 SH - DFND 14 0 16,119 0

AMC ENTMT HLDGS INC CL A COM 00165C104 4,011 429,898 SH - DFND 15 0 429,898 0

AMN HEALTHCARE SERVICES INC COM 001744101 754 13,905 SH - DFND 14 0 13,905 0

AMN HEALTHCARE SERVICES INC COM 001744101 6,449 118,870 SH - DFND 1 0 118,870 0

AMN HEALTHCARE SERVICES INC COM 001744101 20,367 375,421 SH - DFND 15 0 375,421 0

ANI PHARMACEUTICALS INC COM 00182C103 17,611 214,240 SH - DFND 15 0 214,240 0

ARC DOCUMENT SOLUTIONS INC COM 00191G103 364 178,671 SH - DFND 15 0 178,671 0

ASGN INC COM 00191U102 37 604 SH - DFND 14 0 604 0

ASGN INC COM 00191U102 37,877 625,031 SH - DFND 1 0 625,031 0

ASGN INC COM 00191U102 9,931 163,882 SH - DFND 15 0 163,882 0

AT&T INC COM 00206R102 425 12,689 SH - DFND 2,15 0 12,689 0

AT&T INC COM 00206R102 40,048 1,195,104 SH - DFND 14 0 1,195,104 0

AT&T INC COM 00206R102 47 1,400 SH Call DFND 14 0 0 0



AT&T INC COM 00206R102 925,126 27,607,470 SH - DFND 1 0 27,421,692 185,778



AT&T INC COM 00206R102 3,177 94,800 SH - DFND 1,4 0 94,800 0




AT&T INC COM 00206R102 378,967 11,309,078 SH - DFND 15 0 11,309,078 0

AT&T INC COM 00206R102 7,785 232,312 SH - DFND 5 0 0 232,312


AT&T INC COM 00206R102 5,394 160,976 SH - DFND 5,6 0 17,598 143,378

AT&T INC COM 00206R102 1,184 35,340 SH - DFND 11 0 35,340 0

AT&T INC COM 00206R102 4,081 121,777 SH - DFND 13 0 121,777 0

A10 NETWORKS INC COM 002121101 203 29,795 SH - DFND 15 0 29,795 0

ATN INTL INC COM 00215F107 5,428 94,024 SH - DFND 15 0 94,024 0

ASE TECHNOLOGY HOLDING CO LT SPONSORED ADS 00215W100 470 119,056 SH - DFND 14 0 119,056 0

ASE TECHNOLOGY HOLDING CO LT SPONSORED ADS 00215W100 730 184,735 SH - DFND 15 0 184,735 0

AU OPTRONICS CORP SPONSORED ADR 002255107 660 222,980 SH - DFND 14 0 222,980 0

AVX CORP NEW COM 002444107 1,264 76,122 SH - DFND 15 0 76,122 0

AXT INC COM 00246W103 266 67,055 SH - DFND 15 0 67,055 0

AZZ INC COM 002474104 2,917 63,391 SH - DFND 15 0 63,391 0

AARONS INC COM PAR $0.50 002535300 4,392 71,514 SH - DFND 1 0 55,314 16,200

AARONS INC COM PAR $0.50 002535300 718 11,700 SH - DFND 1,3 0 11,700 0



AARONS INC COM PAR $0.50 002535300 1,897 30,889 SH - DFND 5,9 0 30,889 0

AARONS INC COM PAR $0.50 002535300 1,095 17,835 SH - DFND 5,6 0 1,889 15,946

AARONS INC COM PAR $0.50 002535300 748 12,178 SH - DFND 13 0 12,178 0

ABBOTT LABS COM 002824100 95,767 1,138,732 SH - DFND 14 0 1,138,732 0

ABBOTT LABS COM 002824100 42 500 SH Call DFND 14 0 0 0

ABBOTT LABS COM 002824100 151 1,800 SH Call DFND 14 0 0 0

ABBOTT LABS COM 002824100 227 2,700 SH Call DFND 14 0 0 0

ABBOTT LABS COM 002824100 313,570 3,728,537 SH - DFND 1 0 3,528,100 200,437

ABBOTT LABS COM 002824100 8,795 104,575 SH - DFND 1,12 0 104,575 0


ABBOTT LABS COM 002824100 76 900 SH - DFND 1,3 0 900 0


ABBOTT LABS COM 002824100 160,125 1,903,989 SH - DFND 15 0 1,903,989 0

ABBOTT LABS COM 002824100 4,903 58,296 SH - DFND 5 0 0 58,296

ABBOTT LABS COM 002824100 1,777 21,128 SH - DFND 5,6 0 2,648 18,480

ABBOTT LABS COM 002824100 721 8,570 SH - DFND 11 0 8,570 0

ABBOTT LABS COM 002824100 1,564 18,592 SH - DFND 13 0 18,592 0

ABBVIE INC COM 00287Y109 21,624 297,364 SH - DFND 14 0 297,364 0

ABBVIE INC COM 00287Y109 27,909 383,781 SH - DFND 1 0 376,981 6,800

ABBVIE INC COM 00287Y109 1,673 23,000 SH Call DFND 1 0 0 0

ABBVIE INC COM 00287Y109 1,483 20,400 SH - DFND 1,4 0 20,400 0

ABBVIE INC COM 00287Y109 143 1,960 SH - DFND 1,9 0 1,960 0


ABBVIE INC COM 00287Y109 30 419 SH - DFND 1,15 0 419 0

ABBVIE INC COM 00287Y109 3,436 47,245 SH - DFND 1,5 0 47,245 0

ABBVIE INC COM 00287Y109 221,545 3,046,547 SH - DFND 15 0 3,046,547 0

ABBVIE INC COM 00287Y109 32,717 449,906 SH - DFND 5 0 239,792 210,114




ABBVIE INC COM 00287Y109 3,429 47,152 SH - DFND 5,6 0 5,228 41,924

ABBVIE INC COM 00287Y109 516 7,101 SH - DFND 11 0 7,101 0

ABBVIE INC COM 00287Y109 99 1,362 SH - DFND 6 0 1,362 0

ABBVIE INC COM 00287Y109 131 1,800 SH Call DFND 6 0 0 0

ABBVIE INC COM 00287Y109 2,836 39,002 SH - DFND 13 0 39,002 0

ABERCROMBIE & FITCH CO CL A 002896207 751 46,800 SH - DFND 1 0 0 46,800

ABERCROMBIE & FITCH CO CL A 002896207 29 1,795 SH - DFND 1,15 0 1,795 0

ABERCROMBIE & FITCH CO CL A 002896207 4,589 286,068 SH - DFND 15 0 286,068 0

ABERDEEN ASIA PACIFIC INCOM COM 003009107 15,506 3,709,646 SH - DFND 15 0 3,709,646 0

ABERDEEN GLOBAL INCOME FD IN COM 003013109 250 29,879 SH - DFND 14 0 29,879 0

ABERDEEN GLOBAL PREMIER PPTY COM SH BEN INT 00302L108 1,181 188,605 SH - DFND 14 0 188,605 0

ABERDEEN GLOBAL DYNAMIC DIVI COM 00302M106 2,905 301,387 SH - DFND 14 0 301,387 0

ABERDEEN INCOME CR STRATEGIE COM 003057106 4,346 351,624 SH - DFND 14 0 351,624 0

ABERDEEN TOTAL DYNAMIC DIVD COM SH BEN INT 00326L100 2,774 332,584 SH - DFND 14 0 332,584 0

ABIOMED INC COM 003654100 1,423 5,462 SH - DFND 14 0 5,462 0


ABIOMED INC COM 003654100 3 13 SH - DFND 1,15 0 13 0


ABIOMED INC COM 003654100 337 1,292 SH - DFND 5 0 0 1,292

ABIOMED INC COM 003654100 52 200 SH - DFND 11 0 200 0

ABRAXAS PETE CORP COM 003830106 191 185,128 SH - DFND 15 0 185,128 0

ACACIA RESH CORP ACACIA TCHCOM 003881307 7,400 2,500,000 SH - DFND 1 0 2,500,000 0

ACACIA RESH CORP ACACIA TCHCOM 003881307 0 93 SH - DFND 15 0 93 0

ACACIA COMMUNICATIONS INC COM 00401C108 2,854 60,510 SH - DFND 1 0 60,510 0

ACACIA COMMUNICATIONS INC COM 00401C108 1,014 21,500 SH - DFND 1,3 0 21,500 0

ACACIA COMMUNICATIONS INC COM 00401C108 1,730 36,686 SH - DFND 15 0 36,686 0

ACADIA HEALTHCARE COMPANY IN COM 00404A109 208 5,965 SH - DFND 14 0 5,965 0

ACADIA HEALTHCARE COMPANY IN COM 00404A109 8,326 238,234 SH - DFND 1 0 238,234 0

ACADIA HEALTHCARE COMPANY IN COM 00404A109 5,370 153,657 SH - DFND 15 0 123,227 30,430

ACADIA HEALTHCARE COMPANY IN COM 00404A109 3,142 89,914 SH - DFND 6 0 89,914 0

ACADIA HEALTHCARE COMPANY IN COM 00404A109 679 19,439 SH - DFND 6,2 0 19,439 0

ACADIA PHARMACEUTICALS INC COM 004225108 103,797 3,883,170 SH - DFND 1 0 3,597,120 286,050

ACADIA PHARMACEUTICALS INC COM 004225108 1,494 55,880 SH - DFND 1,12 0 55,880 0

ACADIA PHARMACEUTICALS INC COM 004225108 700 26,190 SH - DFND 1,3 0 26,190 0

ACADIA PHARMACEUTICALS INC COM 004225108 1,343 50,242 SH - DFND 15 0 50,242 0

ACADIA RLTY TR COM SH BEN INT 004239109 21,154 772,887 SH - DFND 15 0 772,887 0

ACCELERON PHARMA INC COM 00434H108 323 7,852 SH - DFND 15 0 7,852 0

ACCURAY INC COM 004397105 285 73,655 SH - DFND 15 0 73,655 0

ACHILLION PHARMACEUTICALS IN COM 00448Q201 428 159,678 SH - DFND 15 0 159,678 0

ACI WORLDWIDE INC COM 004498101 343 10,000 SH - DFND 1 0 10,000 0

ACI WORLDWIDE INC COM 004498101 17 500 SH - DFND 1,3 0 500 0

ACI WORLDWIDE INC COM 004498101 26,217 763,454 SH - DFND 15 0 763,454 0

ACLARIS THERAPEUTICS INC COM 00461U105 66 30,162 SH - DFND 15 0 30,162 0

ACORDA THERAPEUTICS INC COM 00484M106 2,565 334,456 SH - DFND 15 0 334,456 0

ACTIVISION BLIZZARD INC COM 00507V109 1,114 23,610 SH - DFND 14 0 23,610 0

ACTIVISION BLIZZARD INC COM 00507V109 498,112 10,553,213 SH - DFND 1 0 10,553,213 0

ACTIVISION BLIZZARD INC COM 00507V109 493 10,452 SH - DFND 1,2 0 10,452 0

ACTIVISION BLIZZARD INC COM 00507V109 40,356 855,000 SH - DFND 1,9 0 855,000 0

ACTIVISION BLIZZARD INC COM 00507V109 10 216 SH - DFND 1,15 0 216 0

ACTIVISION BLIZZARD INC COM 00507V109 31 663 SH - DFND 1,11 0 663 0

ACTIVISION BLIZZARD INC COM 00507V109 371,180 7,863,978 SH - DFND 15 0 7,863,978 0

ACTIVISION BLIZZARD INC COM 00507V109 1,141 24,166 SH - DFND 5 0 0 24,166

ACTIVISION BLIZZARD INC COM 00507V109 168 3,568 SH - DFND 11 0 3,568 0

ACTUANT CORP CL A NEW 00508X203 3,031 122,161 SH - DFND 15 0 122,161 0

ACUITY BRANDS INC COM 00508Y102 82 598 SH - DFND 14 0 598 0

ACUITY BRANDS INC COM 00508Y102 3,694 26,783 SH - DFND 15 0 26,783 0

ACUITY BRANDS INC COM 00508Y102 28 200 SH - DFND 11 0 200 0

ACUSHNET HOLDINGS CORP COM 005098108 1,361 51,834 SH - DFND 15 0 51,834 0

ADAMS RES & ENERGY INC COM NEW 006351308 295 8,610 SH - DFND 1 0 8,610 0

ADAMS RES & ENERGY INC COM NEW 006351308 51 1,500 SH - DFND 1,3 0 1,500 0

ADAPTIVE BIOTECHNOLOGIES COR COM 00650F109 10,320 213,663 SH - DFND 1 0 213,663 0

ADAPTIVE BIOTECHNOLOGIES COR COM 00650F109 548 11,337 SH - DFND 1,12 0 11,337 0

ADDUS HOMECARE CORP COM 006739106 17,154 228,876 SH - DFND 1 0 228,876 0

ADDUS HOMECARE CORP COM 006739106 7,606 101,486 SH - DFND 15 0 101,486 0

ADOBE INC COM 00724F101 627 2,128 SH - DFND 4 0 2,128 0

ADOBE INC COM 00724F101 7,497 25,445 SH - DFND 14 0 25,445 0

ADOBE INC COM 00724F101 881,536 2,991,806 SH - DFND 1 0 2,813,631 178,175

ADOBE INC COM 00724F101 619 2,100 SH - DFND 1,4 0 2,100 0

ADOBE INC COM 00724F101 4,826 16,380 SH - DFND 1,3 0 16,380 0

ADOBE INC COM 00724F101 40 137 SH - DFND 1,15 0 137 0

ADOBE INC COM 00724F101 1,445,154 4,904,646 SH - DFND 15 0 4,904,646 0

ADOBE INC COM 00724F101 4,614 15,658 SH - DFND 5 0 0 15,658

ADOBE INC COM 00724F101 319 1,083 SH - DFND 5,4 0 1,083 0

ADOBE INC COM 00724F101 1,910 6,483 SH - DFND 5,6 0 680 5,803

ADOBE INC COM 00724F101 682 2,313 SH - DFND 11 0 2,313 0

ADOBE INC COM 00724F101 1,646 5,587 SH - DFND 13 0 5,587 0

ADTALEM GLOBAL ED INC COM 00737L103 73 1,617 SH - DFND 14 0 1,617 0

ADTALEM GLOBAL ED INC COM 00737L103 45,713 1,014,715 SH - DFND 1 0 1,000,687 14,028

ADTALEM GLOBAL ED INC COM 00737L103 6,876 152,621 SH - DFND 15 0 152,621 0

ADTRAN INC COM 00738A106 17 1,112 SH - DFND 1,15 0 1,112 0

ADTRAN INC COM 00738A106 5,879 385,486 SH - DFND 15 0 385,486 0

ADURO BIOTECH INC COM 00739L101 93 60,233 SH - DFND 15 0 60,233 0

ADVANCE AUTO PARTS INC COM 00751Y106 26,779 173,729 SH - DFND 2 0 173,729 0


ADVANCE AUTO PARTS INC COM 00751Y106 108,223 702,111 SH - DFND 1 0 693,322 8,789

ADVANCE AUTO PARTS INC COM 00751Y106 9,440 61,246 SH - DFND 1,2 0 61,246 0

ADVANCE AUTO PARTS INC COM 00751Y106 208 1,350 SH - DFND 1,4 0 1,350 0

ADVANCE AUTO PARTS INC COM 00751Y106 224 1,452 SH - DFND 1,9 0 1,452 0


ADVANCE AUTO PARTS INC COM 00751Y106 78 509 SH - DFND 5,4 0 509 0

ADVANCE AUTO PARTS INC COM 00751Y106 8 50 SH - DFND 5,6 0 50 0

ADVANCE AUTO PARTS INC COM 00751Y106 52 335 SH - DFND 11 0 335 0


ADVENT CLAYMORE CV SECS & IN COM 00764C109 2,318 153,928 SH - DFND 14 0 153,928 0

AECOM COM 00766T100 3,901 103,073 SH - DFND 14 0 103,073 0

AECOM COM 00766T100 107,080 2,829,058 SH - DFND 1 0 2,829,058 0

AECOM COM 00766T100 17,870 472,134 SH - DFND 15 0 472,134 0

ADVANCED EMISSIONS SOLUTS IN COM 00770C101 279 22,034 SH - DFND 15 0 22,034 0

AEGION CORP COM 00770F104 1,407 76,446 SH - DFND 14 0 76,446 0

AEGION CORP COM 00770F104 10,537 572,671 SH - DFND 15 0 572,671 0

AERIE PHARMACEUTICALS INC COM 00771V108 26,925 911,153 SH - DFND 1 0 898,868 12,285

AERIE PHARMACEUTICALS INC COM 00771V108 1 34 SH - DFND 15 0 34 0

ADVANSIX INC COM 00773T101 416 17,030 SH - DFND 14 0 17,030 0

ADVANSIX INC COM 00773T101 10 413 SH - DFND 1 0 413 0

ADVANSIX INC COM 00773T101 15 595 SH - DFND 1,15 0 595 0

ADVANSIX INC COM 00773T101 3,553 145,427 SH - DFND 15 0 145,427 0

ADVERUM BIOTECHNOLOGIES INC COM 00773U108 109 9,126 SH - DFND 1 0 9,126 0

ADVERUM BIOTECHNOLOGIES INC COM 00773U108 3,427 288,195 SH - DFND 15 0 288,195 0

AEROJET ROCKETDYNE HLDGS INC COM 007800105 791 17,660 SH - DFND 14 0 17,660 0

AEROJET ROCKETDYNE HLDGS INC COM 007800105 45,017 1,005,513 SH - DFND 1 0 907,155 98,358

AEROJET ROCKETDYNE HLDGS INC COM 007800105 81 1,800 SH - DFND 1,3 0 1,800 0

AEROJET ROCKETDYNE HLDGS INC COM 007800105 15,266 340,978 SH - DFND 15 0 340,978 0

ADVANCED MICRO DEVICES INC COM 007903107 1,077 35,471 SH - DFND 14 0 35,471 0

ADVANCED MICRO DEVICES INC COM 007903107 702,771 23,140,309 SH - DFND 1 0 23,140,309 0

ADVANCED MICRO DEVICES INC COM 007903107 2,506 82,500 SH Call DFND 1 0 0 0

ADVANCED MICRO DEVICES INC COM 007903107 358 11,778 SH - DFND 1,3 0 11,778 0

ADVANCED MICRO DEVICES INC COM 007903107 9 289 SH - DFND 1,15 0 289 0

ADVANCED MICRO DEVICES INC COM 007903107 406,935 13,399,240 SH - DFND 15 0 13,399,240 0

ADVANCED MICRO DEVICES INC COM 007903107 758 24,974 SH - DFND 5 0 0 24,974

ADVANCED MICRO DEVICES INC COM 007903107 140 4,600 SH - DFND 11 0 4,600 0

ADVANCED MICRO DEVICES INC NOTE 2.125% 9/0 007903BD8 4,260 1,100,000 PRN - DFND 1 0 1,000,000 100,000

ADVANCED DRAIN SYS INC DEL COM 00790R104 1,421 43,330 SH - DFND 1 0 7,330 36,000

ADVANCED DRAIN SYS INC DEL COM 00790R104 197 6,000 SH - DFND 1,3 0 6,000 0

ADVANCED DRAIN SYS INC DEL COM 00790R104 18,586 566,804 SH - DFND 15 0 566,804 0

ADVANCED DISP SVCS INC DEL COM 00790X101 20,114 630,349 SH - DFND 1 0 630,349 0

ADVANCED DISP SVCS INC DEL COM 00790X101 3,946 123,663 SH - DFND 15 0 123,663 0

AEGON N V NY REGISTRYSHS 007924103 83 16,809 SH - DFND 15 0 16,809 0

ADVANCED ENERGY INDS COM 007973100 540 9,605 SH - DFND 1 0 0 9,605

ADVANCED ENERGY INDS COM 007973100 16,406 291,560 SH - DFND 15 0 291,560 0

AEROVIRONMENT INC COM 008073108 153 2,694 SH - DFND 14 0 2,694 0

AEROVIRONMENT INC COM 008073108 35 620 SH - DFND 1,15 0 620 0

AEROVIRONMENT INC COM 008073108 5,777 101,770 SH - DFND 15 0 101,770 0

AFFILIATED MANAGERS GROUP IN COM 008252108 40,053 434,700 SH - DFND 2 0 434,700 0


AFFILIATED MANAGERS GROUP IN COM 008252108 7,247 78,655 SH - DFND 1,2 0 78,655 0


AFFILIATED MANAGERS GROUP IN COM 008252108 28 300 SH - DFND 11 0 300 0

AGILENT TECHNOLOGIES INC COM 00846U101 2,829 37,888 SH - DFND 14 0 37,888 0

AGILENT TECHNOLOGIES INC COM 00846U101 622,943 8,342,615 SH - DFND 1 0 7,970,229 372,386

AGILENT TECHNOLOGIES INC COM 00846U101 1,892 25,337 SH - DFND 1,12 0 25,337 0

AGILENT TECHNOLOGIES INC COM 00846U101 523 7,000 SH - DFND 1,4 0 7,000 0

AGILENT TECHNOLOGIES INC COM 00846U101 2,069 27,711 SH - DFND 1,3 0 27,711 0

AGILENT TECHNOLOGIES INC COM 00846U101 7 89 SH - DFND 1,15 0 89 0

AGILENT TECHNOLOGIES INC COM 00846U101 58,029 777,142 SH - DFND 15 0 777,142 0

AGILENT TECHNOLOGIES INC COM 00846U101 825 11,055 SH - DFND 5 0 0 11,055

AGILENT TECHNOLOGIES INC COM 00846U101 10 140 SH - DFND 5,6 0 140 0

AGILENT TECHNOLOGIES INC COM 00846U101 116 1,549 SH - DFND 11 0 1,549 0

AGNICO EAGLE MINES LTD COM 008474108 1,045 20,388 SH - DFND 2,15 0 20,388 0

AGNICO EAGLE MINES LTD COM 008474108 63,383 1,236,989 SH - DFND 1 0 1,236,989 0



AGNICO EAGLE MINES LTD COM 008474108 861 16,809 SH - DFND 1,6 0 16,809 0

AGNICO EAGLE MINES LTD COM 008474108 675 13,179 SH - DFND 15 0 13,179 0

AGNICO EAGLE MINES LTD COM 008474108 582 11,367 SH - DFND 5 0 0 11,367

AGNICO EAGLE MINES LTD COM 008474108 59 1,150 SH - DFND 11 0 1,150 0

AGNICO EAGLE MINES LTD COM 008474108 64,090 1,250,780 SH - DFND 6 0 1,250,780 0

AGNICO EAGLE MINES LTD COM 008474108 794 15,505 SH - DFND 6,9 0 15,505 0

AGILYSYS INC COM 00847J105 3,442 160,330 SH - DFND 15 0 160,330 0

AGIOS PHARMACEUTICALS INC COM 00847X104 68,691 1,377,127 SH - DFND 1 0 1,363,406 13,721

AGIOS PHARMACEUTICALS INC COM 00847X104 980 19,646 SH - DFND 15 0 19,646 0

AGREE REALTY CORP COM 008492100 76 1,184 SH - DFND 14 0 1,184 0

AGREE REALTY CORP COM 008492100 146,942 2,294,183 SH - DFND 1 0 1,114,586 1,179,597

AGREE REALTY CORP COM 008492100 960 14,985 SH - DFND 1,2 0 14,985 0


AGREE REALTY CORP COM 008492100 10,060 157,067 SH - DFND 1,3 0 157,067 0



AGREE REALTY CORP COM 008492100 25,939 404,986 SH - DFND 15 0 389,590 15,396

AGROFRESH SOLUTIONS COM 00856G109 176 116,476 SH - DFND 15 0 116,476 0

AIR LEASE CORP CL A 00912X302 1,776 42,963 SH - DFND 15 0 42,963 0

AIR PRODS & CHEMS INC COM 009158106 78,165 345,297 SH - DFND 14 0 345,297 0

AIR PRODS & CHEMS INC COM 009158106 152,430 673,367 SH - DFND 1 0 673,167 200

AIR PRODS & CHEMS INC COM 009158106 928 4,100 SH - DFND 1,4 0 4,100 0

AIR PRODS & CHEMS INC COM 009158106 2,013 8,891 SH - DFND 1,15 0 8,891 0

AIR PRODS & CHEMS INC COM 009158106 241,463 1,066,676 SH - DFND 15 0 1,066,676 0

AIR PRODS & CHEMS INC COM 009158106 1,940 8,572 SH - DFND 5 0 0 8,572

AIR PRODS & CHEMS INC COM 009158106 11 50 SH - DFND 5,6 0 50 0

AIR PRODS & CHEMS INC COM 009158106 233 1,030 SH - DFND 11 0 1,030 0

AIR TRANSPORT SERVICES GRP I COM 00922R105 2,018 82,700 SH - DFND 15 0 82,700 0

AIR TRANSPORT SERVICES GRP I NOTE 1.125%10/1 00922RAB1 10,307 10,120,000 PRN - DFND 1 0 9,000,000 1,120,000

AKAMAI TECHNOLOGIES INC COM 00971T101 636 7,932 SH - DFND 14 0 7,932 0

AKAMAI TECHNOLOGIES INC COM 00971T101 16,680 208,133 SH - DFND 1 0 207,783 350

AKAMAI TECHNOLOGIES INC COM 00971T101 601 7,500 SH - DFND 1,4 0 7,500 0

AKAMAI TECHNOLOGIES INC COM 00971T101 4 46 SH - DFND 1,15 0 46 0

AKAMAI TECHNOLOGIES INC COM 00971T101 3,169 39,545 SH - DFND 1,5 0 39,545 0

AKAMAI TECHNOLOGIES INC COM 00971T101 91,688 1,144,098 SH - DFND 15 0 1,144,098 0

AKAMAI TECHNOLOGIES INC COM 00971T101 28,538 356,097 SH - DFND 5 0 294,373 61,724




AKAMAI TECHNOLOGIES INC COM 00971T101 2,697 33,655 SH - DFND 5,6 0 3,707 29,948

AKAMAI TECHNOLOGIES INC COM 00971T101 60 744 SH - DFND 11 0 744 0

AKAMAI TECHNOLOGIES INC COM 00971T101 675 8,420 SH - DFND 6 0 8,420 0

AKAMAI TECHNOLOGIES INC COM 00971T101 2,663 33,228 SH - DFND 13 0 33,228 0

AKAMAI TECHNOLOGIES INC NOTE 0.125% 5/0 00971TAJ0 11,041 10,335,000 PRN - DFND 1 0 9,575,000 760,000

AKAMAI TECHNOLOGIES INC NOTE 0.125% 5/0 00971TAJ0 160 150,000 PRN - DFND 6 0 150,000 0

AKORN INC COM 009728106 5,721 1,110,960 SH - DFND 15 0 1,110,960 0

ALAMO GROUP INC COM 011311107 16,795 168,072 SH - DFND 15 0 168,072 0

ALAMOS GOLD INC NEW COM CL A 011532108 67 11,017 SH - DFND 2,15 0 11,017 0

ALAMOS GOLD INC NEW COM CL A 011532108 11,828 1,955,000 SH - DFND 1 0 1,955,000 0

ALAMOS GOLD INC NEW COM CL A 011532108 7,477 1,235,805 SH - DFND 1,2 0 1,235,805 0

ALAMOS GOLD INC NEW COM CL A 011532108 2,705 447,105 SH - DFND 1,9 0 447,105 0

ALAMOS GOLD INC NEW COM CL A 011532108 389 64,275 SH - DFND 15 0 64,275 0

ALARM COM HLDGS INC COM 011642105 21,400 400,000 SH - DFND 1 0 400,000 0

ALARM COM HLDGS INC COM 011642105 43 800 SH - DFND 1,3 0 800 0

ALARM COM HLDGS INC COM 011642105 9,558 178,646 SH - DFND 15 0 159,074 19,572

ALASKA AIR GROUP INC COM 011659109 16,007 250,455 SH - DFND 1 0 250,455 0

ALASKA AIR GROUP INC COM 011659109 45,266 708,275 SH - DFND 15 0 690,975 17,300

ALBANY INTL CORP CL A 012348108 52,737 636,071 SH - DFND 1 0 636,071 0

ALBANY INTL CORP CL A 012348108 1,110 13,394 SH - DFND 1,11 0 13,394 0



ALBANY INTL CORP CL A 012348108 1,312 15,829 SH - DFND 6,2 0 15,829 0

ALBANY INTL CORP CL A 012348108 655 7,896 SH - DFND 6,9 0 7,896 0

ALBEMARLE CORP COM 012653101 722 10,250 SH - DFND 2 0 10,250 0

ALBEMARLE CORP COM 012653101 406 5,766 SH - DFND 14 0 5,766 0

ALBEMARLE CORP COM 012653101 31,627 449,182 SH - DFND 1 0 449,182 0

ALBEMARLE CORP COM 012653101 7,825 111,141 SH - DFND 1,2 0 111,141 0

ALBEMARLE CORP COM 012653101 1,883 26,745 SH - DFND 1,9 0 26,745 0

ALBEMARLE CORP COM 012653101 44,231 628,193 SH - DFND 15 0 628,193 0

ALBEMARLE CORP COM 012653101 35 500 SH - DFND 11 0 500 0

ALCOA CORP COM 013872106 46 1,952 SH - DFND 14 0 1,952 0

ALCOA CORP COM 013872106 2,588 110,560 SH - DFND 1 0 110,560 0

ALCOA CORP COM 013872106 6,537 279,250 SH - DFND 15 0 279,250 0

ALDER BIOPHARMACEUTICALS INC COM 014339105 236 20,039 SH - DFND 15 0 20,039 0

ALEXANDER & BALDWIN INC NEW COM 014491104 4,965 214,933 SH - DFND 15 0 214,933 0

ALEXANDERS INC COM 014752109 914 2,468 SH - DFND 15 0 2,468 0

ALEXANDRIA REAL ESTATE EQ IN COM 015271109 1,434 10,164 SH - DFND 14 0 10,164 0

ALEXANDRIA REAL ESTATE EQ IN COM 015271109 313,925 2,224,998 SH - DFND 1 0 1,328,933 896,065

ALEXANDRIA REAL ESTATE EQ IN COM 015271109 1,679 11,902 SH - DFND 1,2 0 11,902 0

ALEXANDRIA REAL ESTATE EQ IN COM 015271109 222 1,570 SH - DFND 1,9 0 1,570 0



ALEXANDRIA REAL ESTATE EQ IN COM 015271109 873 6,191 SH - DFND 1,13 0 6,191 0

ALEXANDRIA REAL ESTATE EQ IN COM 015271109 137,811 976,756 SH - DFND 15 0 976,756 0

ALEXANDRIA REAL ESTATE EQ IN COM 015271109 71 500 SH - DFND 11 0 500 0

ALEXANDRIA REAL ESTATE EQ IN PFD CONV SER D 015271505 1,415 37,121 SH - DFND 1,11 0 37,121 0

ALEXION PHARMACEUTICALS INC COM 015351109 2,848 21,744 SH - DFND 14 0 21,744 0

ALEXION PHARMACEUTICALS INC COM 015351109 81,129 619,400 SH - DFND 1 0 617,900 1,500

ALEXION PHARMACEUTICALS INC COM 015351109 5,942 45,369 SH - DFND 1,12 0 45,369 0

ALEXION PHARMACEUTICALS INC COM 015351109 498 3,800 SH - DFND 1,4 0 3,800 0



ALEXION PHARMACEUTICALS INC COM 015351109 8 63 SH - DFND 1,15 0 63 0

ALEXION PHARMACEUTICALS INC COM 015351109 333,835 2,548,749 SH - DFND 15 0 2,548,749 0

ALEXION PHARMACEUTICALS INC COM 015351109 3,799 29,006 SH - DFND 5 0 0 29,006


ALEXION PHARMACEUTICALS INC COM 015351109 425 3,248 SH - DFND 5,6 0 373 2,875

ALEXION PHARMACEUTICALS INC COM 015351109 144 1,100 SH - DFND 11 0 1,100 0

ALEXION PHARMACEUTICALS INC COM 015351109 400 3,052 SH - DFND 13 0 3,052 0

ALGONQUIN PWR UTILS CORP COM 015857105 21,076 1,740,419 SH - DFND 2 0 1,740,419 0

ALGONQUIN PWR UTILS CORP COM 015857105 14,954 1,234,844 SH - DFND 2,15 0 1,234,844 0

ALGONQUIN PWR UTILS CORP COM 015857105 40,059 3,307,947 SH - DFND 15 0 3,307,947 0

ALIBABA GROUP HLDG LTD SPONSORED ADS 01609W102 76,611 452,118 SH - DFND 2 0 452,118 0

ALIBABA GROUP HLDG LTD SPONSORED ADS 01609W102 294,341 1,737,037 SH - DFND 4 0 1,737,037 0

ALIBABA GROUP HLDG LTD SPONSORED ADS 01609W102 224,171 1,322,931 SH - DFND 4,9 0 1,322,931 0

ALIBABA GROUP HLDG LTD SPONSORED ADS 01609W102 4,236,050 24,998,821 SH - DFND 1 0 24,817,776 181,045

ALIBABA GROUP HLDG LTD SPONSORED ADS 01609W102 41,149 242,837 SH - DFND 1,2 0 242,837 0



ALIBABA GROUP HLDG LTD SPONSORED ADS 01609W102 187,594 1,107,076 SH - DFND 1,3 0 1,107,076 0

ALIBABA GROUP HLDG LTD SPONSORED ADS 01609W102 158,114 933,100 SH - DFND 15 0 615,530 317,570

ALIBABA GROUP HLDG LTD SPONSORED ADS 01609W102 32 186 SH - DFND 11,2 0 186 0

ALIBABA GROUP HLDG LTD SPONSORED ADS 01609W102 244,532 1,443,095 SH - DFND 6 0 820,353 622,742

ALIBABA GROUP HLDG LTD SPONSORED ADS 01609W102 329 1,940 SH - DFND 6,9 0 1,940 0

ALIBABA GROUP HLDG LTD SPONSORED ADS 01609W102 7,178 42,362 SH - DFND 13 0 42,362 0

ALIGN TECHNOLOGY INC COM 016255101 132,908 485,599 SH - DFND 1 0 485,599 0

ALIGN TECHNOLOGY INC COM 016255101 2,901 10,599 SH - DFND 1,12 0 10,599 0

ALIGN TECHNOLOGY INC COM 016255101 6 21 SH - DFND 1,15 0 21 0

ALIGN TECHNOLOGY INC COM 016255101 276,249 1,009,312 SH - DFND 15 0 1,009,312 0

ALIGN TECHNOLOGY INC COM 016255101 417 1,522 SH - DFND 5 0 0 1,522

ALIGN TECHNOLOGY INC COM 016255101 109 400 SH - DFND 11 0 400 0

ALLEGHANY CORP DEL COM 017175100 49,649 72,894 SH - DFND 15 0 72,894 0

ALLEGHANY CORP DEL COM 017175100 68 100 SH - DFND 11 0 100 0

ALLEGHENY TECHNOLOGIES INC COM 01741R102 519 20,583 SH - DFND 14 0 20,583 0

ALLEGHENY TECHNOLOGIES INC COM 01741R102 75,113 2,980,687 SH - DFND 1 0 2,980,687 0

ALLEGHENY TECHNOLOGIES INC COM 01741R102 12,691 503,624 SH - DFND 15 0 503,624 0

ALLEGIANCE BANCSHARES INC COM 01748H107 308 9,228 SH - DFND 15 0 9,228 0

ALLEGIANT TRAVEL CO COM 01748X102 499 3,477 SH - DFND 14 0 3,477 0

ALLEGIANT TRAVEL CO COM 01748X102 4,294 29,926 SH - DFND 15 0 29,926 0

ALLETE INC COM NEW 018522300 2,106 25,311 SH - DFND 14 0 25,311 0

ALLETE INC COM NEW 018522300 60 720 SH - DFND 1 0 720 0

ALLETE INC COM NEW 018522300 25 300 SH - DFND 1,3 0 300 0

ALLETE INC COM NEW 018522300 40,381 485,287 SH - DFND 15 0 485,287 0

ALLIANCE DATA SYSTEMS CORP COM 018581108 8,537 60,922 SH - DFND 2 0 60,922 0



ALLIANCE DATA SYSTEMS CORP COM 018581108 14,266 101,804 SH - DFND 1,2 0 101,804 0


ALLIANCE DATA SYSTEMS CORP COM 018581108 336 2,400 SH - DFND 5 0 0 2,400

ALLIANCE DATA SYSTEMS CORP COM 018581108 28 200 SH - DFND 11 0 200 0

ALLIANCEBERNSTEIN NATL MUNI COM 01864U106 600 44,628 SH - DFND 14 0 44,628 0

ALLIANCE RES PARTNER L P UT LTD PART 01877R108 6,962 410,013 SH - DFND 1 0 410,013 0

ALLIANCEBERNSTEIN GBL HGH IN COM 01879R106 249 21,117 SH - DFND 14 0 21,117 0

ALLIANCEBERNSTEIN GBL HGH IN COM 01879R106 14,592 1,238,677 SH - DFND 15 0 1,238,677 0

ALLIANT ENERGY CORP COM 018802108 5,325 108,495 SH - DFND 14 0 108,495 0

ALLIANT ENERGY CORP COM 018802108 15,763 321,177 SH - DFND 1 0 321,177 0

ALLIANT ENERGY CORP COM 018802108 191,540 3,902,605 SH - DFND 15 0 3,902,605 0

ALLIANT ENERGY CORP COM 018802108 581 11,834 SH - DFND 5 0 0 11,834

ALLIANT ENERGY CORP COM 018802108 58 1,176 SH - DFND 11 0 1,176 0

ALLIANZGI EQUITY & CONV INCO COM 018829101 4,570 210,622 SH - DFND 14 0 210,622 0

ALLIANZGI NFJ DIVID INT & PR COM 01883A107 16,913 1,363,983 SH - DFND 15 0 1,363,983 0

ALLIANZGI CON INCM 2024 TARG COM 01883H102 743 80,511 SH - DFND 14 0 80,511 0

ALLIANZGI CON INCM 2024 TARG COM 01883H102 1,920 208,028 SH - DFND 15 0 208,028 0

ALLIANZGI DIVERS INC & CNV F COM 01883J108 348 14,899 SH - DFND 14 0 14,899 0

ALLIED MOTION TECHNOLOGIES I COM 019330109 220 5,794 SH - DFND 15 0 5,794 0

ALLISON TRANSMISSION HLDGS I COM 01973R101 148 3,184 SH - DFND 14 0 3,184 0

ALLISON TRANSMISSION HLDGS I COM 01973R101 16,008 345,372 SH - DFND 1 0 277,622 67,750

ALLISON TRANSMISSION HLDGS I COM 01973R101 161 3,480 SH - DFND 1,9 0 3,480 0


ALLISON TRANSMISSION HLDGS I COM 01973R101 1,576 33,994 SH - DFND 1,5 0 33,994 0

ALLISON TRANSMISSION HLDGS I COM 01973R101 4,428 95,536 SH - DFND 15 0 95,536 0

ALLISON TRANSMISSION HLDGS I COM 01973R101 10,178 219,597 SH - DFND 5 0 67,991 151,606



ALLISON TRANSMISSION HLDGS I COM 01973R101 2,307 49,763 SH - DFND 5,6 0 5,647 44,116

ALLISON TRANSMISSION HLDGS I COM 01973R101 148 3,197 SH - DFND 6 0 3,197 0

ALLISON TRANSMISSION HLDGS I COM 01973R101 1,606 34,647 SH - DFND 13 0 34,647 0

ALLSCRIPTS HEALTHCARE SOLUTN COM 01988P108 285 24,501 SH - DFND 14 0 24,501 0

ALLSCRIPTS HEALTHCARE SOLUTN COM 01988P108 4,349 373,973 SH - DFND 15 0 373,973 0

ALLSTATE CORP COM 020002101 5,848 57,505 SH - DFND 14 0 57,505 0

ALLSTATE CORP COM 020002101 224,496 2,207,647 SH - DFND 1 0 2,160,587 47,060

ALLSTATE CORP COM 020002101 193 1,900 SH - DFND 1,4 0 1,900 0


ALLSTATE CORP COM 020002101 1,883 18,521 SH - DFND 1,15 0 18,521 0

ALLSTATE CORP COM 020002101 203,980 2,005,901 SH - DFND 15 0 1,995,757 10,144

ALLSTATE CORP COM 020002101 1,938 19,062 SH - DFND 5 0 0 19,062


ALLSTATE CORP COM 020002101 1,330 13,077 SH - DFND 5,6 0 1,592 11,485

ALLSTATE CORP COM 020002101 160 1,578 SH - DFND 11 0 1,578 0

ALLSTATE CORP COM 020002101 871 8,562 SH - DFND 13 0 8,562 0

ALLY FINL INC COM 02005N100 449 14,499 SH - DFND 14 0 14,499 0

ALLY FINL INC COM 02005N100 145,574 4,697,466 SH - DFND 1 0 4,521,758 175,708

ALLY FINL INC COM 02005N100 734 23,694 SH - DFND 1,9 0 23,694 0


ALLY FINL INC COM 02005N100 3 110 SH - DFND 1,15 0 110 0

ALLY FINL INC COM 02005N100 7,894 254,742 SH - DFND 1,5 0 254,742 0

ALLY FINL INC COM 02005N100 14,061 453,733 SH - DFND 15 0 453,733 0

ALLY FINL INC COM 02005N100 36,027 1,162,523 SH - DFND 5 0 792,710 369,813



ALLY FINL INC COM 02005N100 4,001 129,118 SH - DFND 5,6 0 41,215 87,903



ALLY FINL INC COM 02005N100 2,929 94,508 SH - DFND 13 0 94,508 0

ALNYLAM PHARMACEUTICALS INC COM 02043Q107 33,834 466,284 SH - DFND 1 0 466,284 0

ALNYLAM PHARMACEUTICALS INC COM 02043Q107 2,167 29,864 SH - DFND 15 0 29,864 0

ALNYLAM PHARMACEUTICALS INC COM 02043Q107 29 400 SH - DFND 11 0 400 0

ALPHABET INC CAP STK CL C 02079K107 111,273 102,944 SH - DFND 2 0 102,944 0

ALPHABET INC CAP STK CL C 02079K107 829 767 SH - DFND 4 0 767 0


ALPHABET INC CAP STK CL C 02079K107 2,059 1,905 SH - DFND 1,2 0 1,905 0


ALPHABET INC CAP STK CL C 02079K107 49 45 SH - DFND 1,3 0 45 0



ALPHABET INC CAP STK CL C 02079K107 3,280,412 3,034,861 SH - DFND 15 0 3,034,861 0

ALPHABET INC CAP STK CL C 02079K107 13,664 12,641 SH - DFND 5 0 1,409 11,232


ALPHABET INC CAP STK CL C 02079K107 2,559 2,367 SH - DFND 5,6 0 279 2,088



ALPHABET INC CAP STK CL C 02079K107 139 129 SH - DFND 6 0 129 0


ALPHABET INC CAP STK CL A 02079K305 134,298 124,028 SH - DFND 2 0 124,028 0


ALPHABET INC CAP STK CL A 02079K305 2,055,275 1,898,111 SH - DFND 1 0 1,787,747 110,364

ALPHABET INC CAP STK CL A 02079K305 1,045 965 SH - DFND 1,2 0 965 0

ALPHABET INC CAP STK CL A 02079K305 4,737 4,375 SH - DFND 1,4 0 4,375 0



ALPHABET INC CAP STK CL A 02079K305 187 173 SH - DFND 1,15 0 173 0

ALPHABET INC CAP STK CL A 02079K305 2,887,614 2,666,803 SH - DFND 15 0 2,666,803 0

ALPHABET INC CAP STK CL A 02079K305 12,085 11,161 SH - DFND 5 0 897 10,264

ALPHABET INC CAP STK CL A 02079K305 7,813 7,216 SH - DFND 5,6 0 797 6,419


ALPHABET INC CAP STK CL A 02079K305 474 438 SH - DFND 11,2 0 438 0

ALPHABET INC CAP STK CL A 02079K305 902 833 SH - DFND 13 0 833 0

ALTA MESA RES INC CL A 02133L109 44 300,000 SH - DFND 15 0 300,000 0

ALTABA INC COM 021346101 67,166 968,231 SH - DFND 1 0 942,410 25,821

ALTABA INC COM 021346101 291 4,202 SH - DFND 1,9 0 4,202 0

ALTAIR ENGR INC COM CL A 021369103 993 24,574 SH - DFND 15 0 24,574 0

ALTERYX INC COM CL A 02156B103 37,518 343,820 SH - DFND 1 0 343,820 0

ALTERYX INC COM CL A 02156B103 26,218 240,272 SH - DFND 15 0 240,272 0

ALTICE USA INC CL A 02156K103 55 2,250 SH - DFND 14 0 2,250 0

ALTICE USA INC CL A 02156K103 34,540 1,418,500 SH - DFND 1 0 1,374,500 44,000

ALTICE USA INC CL A 02156K103 27,637 1,135,000 SH - DFND 1,9 0 1,135,000 0

ALTICE USA INC CL A 02156K103 3,726 153,021 SH - DFND 15 0 153,021 0

ALTICE USA INC CL A 02156K103 16 640 SH - DFND 5,6 0 640 0

ALTICE USA INC CL A 02156K103 17 700 SH - DFND 11 0 700 0

ALTRA INDL MOTION CORP COM 02208R106 12,083 336,775 SH - DFND 1 0 221,126 115,649

ALTRA INDL MOTION CORP COM 02208R106 1,604 44,698 SH - DFND 15 0 44,698 0

ALTRIA GROUP INC COM 02209S103 368 7,767 SH - DFND 2,15 0 7,767 0

ALTRIA GROUP INC COM 02209S103 4,806 101,505 SH - DFND 14 0 101,505 0

ALTRIA GROUP INC COM 02209S103 439,340 9,278,568 SH - DFND 1 0 9,160,255 118,313


ALTRIA GROUP INC COM 02209S103 1,489 31,451 SH - DFND 1,9 0 31,451 0


ALTRIA GROUP INC COM 02209S103 25 531 SH - DFND 1,15 0 531 0


ALTRIA GROUP INC COM 02209S103 202,109 4,268,397 SH - DFND 15 0 4,268,397 0

ALTRIA GROUP INC COM 02209S103 3,017 63,708 SH - DFND 5 0 0 63,708


ALTRIA GROUP INC COM 02209S103 1,768 37,338 SH - DFND 5,6 0 4,186 33,152

ALTRIA GROUP INC COM 02209S103 434 9,160 SH - DFND 11 0 9,160 0

ALTRIA GROUP INC COM 02209S103 117,166 2,474,467 SH - DFND 6 0 2,474,467 0


AMARIN CORP PLC SPONS ADR NEW 023111206 160,995 8,303,000 SH - DFND 1 0 8,303,000 0

AMARIN CORP PLC SPONS ADR NEW 023111206 1,793 92,484 SH - DFND 1,12 0 92,484 0

AMARIN CORP PLC SPONS ADR NEW 023111206 301 15,500 SH - DFND 1,3 0 15,500 0

AMARIN CORP PLC SPONS ADR NEW 023111206 77 3,959 SH - DFND 15 0 3,959 0

AMAZON COM INC COM 023135106 377 199 SH - DFND 4 0 199 0

AMAZON COM INC COM 023135106 9,159 4,837 SH - DFND 14 0 4,837 0

AMAZON COM INC COM 023135106 2,867,388 1,514,228 SH - DFND 1 0 1,479,715 34,513

AMAZON COM INC COM 023135106 1,515 800 SH Call DFND 1 0 0 0

AMAZON COM INC COM 023135106 9,648 5,095 SH - DFND 1,4 0 5,095 0



AMAZON COM INC COM 023135106 225 119 SH - DFND 1,15 0 119 0


AMAZON COM INC COM 023135106 7,889,861 4,166,527 SH - DFND 15 0 4,166,527 0

AMAZON COM INC COM 023135106 33,744 17,820 SH - DFND 5 0 3,108 14,712


AMAZON COM INC COM 023135106 16,376 8,648 SH - DFND 5,6 0 951 7,697


AMAZON COM INC COM 023135106 267 141 SH - DFND 6 0 141 0

AMAZON COM INC COM 023135106 189 100 SH Call DFND 6 0 0 0


AMBAC FINL GROUP INC COM NEW 023139884 6,025 357,545 SH - DFND 15 0 357,545 0

AMBER RD INC COM 02318Y108 211 16,145 SH - DFND 1 0 16,145 0

AMBEV SA SPONSORED ADR 02319V103 94 20,166 SH - DFND 14 0 20,166 0

AMBEV SA SPONSORED ADR 02319V103 247,561 53,010,833 SH - DFND 1 0 53,010,833 0

AMBEV SA SPONSORED ADR 02319V103 22,747 4,870,858 SH - DFND 1,2 0 4,870,858 0

AMBEV SA SPONSORED ADR 02319V103 33,394 7,150,723 SH - DFND 1,3 0 7,150,723 0

AMBEV SA SPONSORED ADR 02319V103 3,706 793,660 SH - DFND 15 0 487,262 306,398

AMBEV SA SPONSORED ADR 02319V103 161 34,483 SH - DFND 5,4 0 34,483 0

AMBEV SA SPONSORED ADR 02319V103 908 194,383 SH - DFND 13 0 194,383 0

AMEDISYS INC COM 023436108 21,716 178,867 SH - DFND 1 0 178,867 0

AMEDISYS INC COM 023436108 219 1,800 SH - DFND 1,3 0 1,800 0

AMEDISYS INC COM 023436108 21,880 180,212 SH - DFND 15 0 180,212 0

AMEDISYS INC COM 023436108 829 6,826 SH - DFND 5 0 0 6,826

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AMERCO COM 023586100 4,552 12,024 SH - DFND 15 0 12,024 0

AMEREN CORP COM 023608102 4,543 60,480 SH - DFND 14 0 60,480 0

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AMEREN CORP COM 023608102 737 9,810 SH - DFND 5 0 0 9,810

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AMERESCO INC CL A 02361E108 6,001 407,370 SH - DFND 15 0 407,370 0

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AMERICA MOVIL SAB DE CV SPON ADR L SHS 02364W105 164 11,250 SH - DFND 5,4 0 11,250 0

AMERICA MOVIL SAB DE CV SPON ADR L SHS 02364W105 3,713 255,000 SH - DFND 6 0 255,000 0

AMERICA MOVIL SAB DE CV SPON ADR L SHS 02364W105 115 7,900 SH - DFND 6,9 0 7,900 0

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AMERICAN AIRLS GROUP INC COM 02376R102 23 700 SH - DFND 11 0 700 0

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AMERICAN ASSETS TR INC COM 024013104 427 9,066 SH - DFND 1,2 0 9,066 0

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AMERICAN ELEC PWR CO INC COM 025537101 449 5,100 SH - DFND 1,4 0 5,100 0

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AMERICAN ELEC PWR CO INC COM 025537101 208,730 2,371,667 SH - DFND 15 0 2,371,667 0

AMERICAN ELEC PWR CO INC COM 025537101 1,598 18,155 SH - DFND 5 0 0 18,155



AMERICAN ELEC PWR CO INC COM 025537101 208 2,366 SH - DFND 11 0 2,366 0

AMERICAN ELEC PWR CO INC UNIT 03/15/2022 025537127 14,635 273,500 SH - DFND 1 0 249,788 23,712

AMERICAN ELEC PWR CO INC UNIT 03/15/2022 025537127 144 2,700 SH - DFND 6 0 2,700 0

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AMERICAN EAGLE OUTFITTERS IN COM 02553E106 690 40,855 SH - DFND 1,11 0 40,855 0

AMERICAN EAGLE OUTFITTERS IN COM 02553E106 5,900 349,088 SH - DFND 15 0 349,088 0

AMERICAN EAGLE OUTFITTERS IN COM 02553E106 3,674 217,424 SH - DFND 6 0 217,424 0



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AMERICAN EXPRESS CO COM 025816109 78 634 SH - DFND 1,11 0 634 0


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AMERICAN EXPRESS CO COM 025816109 40 324 SH - DFND 6,2 0 324 0


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AMERICAN FINL GROUP INC OHIO COM 025932104 63,690 621,549 SH - DFND 1 0 614,681 6,868


AMERICAN FINL GROUP INC OHIO COM 025932104 41 400 SH - DFND 11 0 400 0


AMERICAN FINL GROUP INC OHIO COM 025932104 1,015 9,903 SH - DFND 6,2 0 9,903 0

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AMERICAN HOMES 4 RENT CL A 02665T306 1,068 43,913 SH - DFND 1,2 0 43,913 0

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AMERICAN INTL GROUP INC COM NEW 026874784 1,708 32,063 SH - DFND 1,9 0 32,063 0

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AMERICAN INTL GROUP INC COM NEW 026874784 95,026 1,783,528 SH - DFND 15 0 1,783,528 0

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AMERICAN INTL GROUP INC COM NEW 026874784 231 4,330 SH - DFND 11 0 4,330 0

AMERICAN MIDSTREAM PARTNERS COM UNITS 02752P100 32,194 6,227,012 SH - DFND 1 0 6,227,012 0

AMERICAN NATL BANKSHARES INC COM 027745108 749 19,330 SH - DFND 15 0 98 19,232

AMERICAN NATL INS CO COM 028591105 2,872 24,663 SH - DFND 15 0 24,663 0

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AMERICAN PUBLIC EDUCATION IN COM 02913V103 2,244 75,870 SH - DFND 15 0 75,870 0

AMERICAN RLTY CAP PPTYS INC NOTE 3.750%12/1 02917TAB0 629 625,000 PRN - DFND 1 0 625,000 0

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AMERICAN SOFTWARE INC CL A 029683109 499 37,928 SH - DFND 15 0 37,928 0

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AMERICAN STS WTR CO COM 029899101 56,223 747,252 SH - DFND 15 0 747,252 0

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AMERICAN TOWER CORP NEW COM 03027X100 422 2,065 SH - DFND 1,9 0 2,065 0

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AMERICAN TOWER CORP NEW COM 03027X100 5,238 25,620 SH - DFND 1,15 0 25,620 0

AMERICAN TOWER CORP NEW COM 03027X100 208,154 1,018,116 SH - DFND 15 0 1,018,116 0

AMERICAN TOWER CORP NEW COM 03027X100 31 150 SH - DFND 5,6 0 150 0

AMERICAN TOWER CORP NEW COM 03027X100 444 2,170 SH - DFND 11 0 2,170 0

AMERICAN TOWER CORP NEW COM 03027X100 1,682 8,225 SH - DFND 13 0 8,225 0

AMERICAN VANGUARD CORP COM 030371108 1,100 71,359 SH - DFND 15 0 71,359 0

AMERICAN WTR WKS CO INC NEW COM 030420103 6,420 55,348 SH - DFND 14 0 55,348 0

AMERICAN WTR WKS CO INC NEW COM 030420103 17,529 151,109 SH - DFND 1 0 140,871 10,238

AMERICAN WTR WKS CO INC NEW COM 030420103 6 51 SH - DFND 1,15 0 51 0

AMERICAN WTR WKS CO INC NEW COM 030420103 335,834 2,895,119 SH - DFND 15 0 2,895,119 0

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AMERICAN WTR WKS CO INC NEW COM 030420103 104 900 SH - DFND 11 0 900 0

AMERICAN WOODMARK CORPORATIO COM 030506109 3,333 39,385 SH - DFND 15 0 39,385 0

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AMERICOLD RLTY TR COM 03064D108 82,218 2,536,027 SH - DFND 1 0 1,214,751 1,321,276

AMERICOLD RLTY TR COM 03064D108 578 17,834 SH - DFND 1,2 0 17,834 0

AMERICOLD RLTY TR COM 03064D108 5,901 182,031 SH - DFND 1,3 0 182,031 0



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AMERISAFE INC COM 03071H100 1,254 19,662 SH - DFND 14 0 19,662 0

AMERISAFE INC COM 03071H100 32 507 SH - DFND 1,15 0 507 0

AMERISAFE INC COM 03071H100 21,869 342,943 SH - DFND 15 0 342,943 0

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AMERISOURCEBERGEN CORP COM 03073E105 648 7,600 SH - DFND 1,4 0 7,600 0

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AMERISOURCEBERGEN CORP COM 03073E105 81,426 955,037 SH - DFND 15 0 955,037 0

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AMERISOURCEBERGEN CORP COM 03073E105 247 2,901 SH - DFND 5,4 0 2,901 0

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AMERISOURCEBERGEN CORP COM 03073E105 69 806 SH - DFND 11 0 806 0

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AMERIPRISE FINL INC COM 03076C106 1,586 10,927 SH - DFND 14 0 10,927 0

AMERIPRISE FINL INC COM 03076C106 24,580 169,328 SH - DFND 1 0 168,178 1,150

AMERIPRISE FINL INC COM 03076C106 1,060 7,300 SH - DFND 1,4 0 7,300 0

AMERIPRISE FINL INC COM 03076C106 6 38 SH - DFND 1,15 0 38 0


AMERIPRISE FINL INC COM 03076C106 6,484 44,669 SH - DFND 5 0 31,588 13,081

AMERIPRISE FINL INC COM 03076C106 358 2,467 SH - DFND 5,4 0 2,467 0

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AMERIPRISE FINL INC COM 03076C106 88 608 SH - DFND 11 0 608 0

AMERIPRISE FINL INC COM 03076C106 779 5,366 SH - DFND 6 0 5,366 0


AMERIS BANCORP COM 03076K108 2,723 69,475 SH - DFND 15 0 69,475 0

AMERIGAS PARTNERS L P UNIT L P INT 030975106 136,623 3,921,429 SH - DFND 1 0 3,921,429 0

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AMETEK INC NEW COM 031100100 24,422 268,849 SH - DFND 2 0 268,849 0


AMETEK INC NEW COM 031100100 52,337 576,144 SH - DFND 1 0 572,194 3,950

AMETEK INC NEW COM 031100100 1,172 12,900 SH - DFND 1,4 0 12,900 0

AMETEK INC NEW COM 031100100 228 2,515 SH - DFND 1,3 0 2,515 0

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AMETEK INC NEW COM 031100100 385 4,239 SH - DFND 5 0 0 4,239

AMETEK INC NEW COM 031100100 22 240 SH - DFND 5,6 0 240 0

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AMGEN INC COM 031162100 9,691 52,590 SH - DFND 14 0 52,590 0

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AMGEN INC COM 031162100 1,391 7,550 SH - DFND 1,4 0 7,550 0

AMGEN INC COM 031162100 184 1,000 SH - DFND 1,9 0 1,000 0



AMGEN INC COM 031162100 1,269,894 6,891,111 SH - DFND 15 0 6,891,111 0

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AMICUS THERAPEUTICS INC COM 03152W109 455 36,444 SH - DFND 15 0 36,444 0

AMKOR TECHNOLOGY INC COM 031652100 7,889 1,057,520 SH - DFND 15 0 1,057,520 0

AMNEAL PHARMACEUTICALS INC COM STK CL A 03168L105 1,453 202,689 SH - DFND 15 0 50,670 152,019

AMPCO-PITTSBURGH CORP COM 032037103 208 51,690 SH - DFND 15 0 51,690 0

AMPHENOL CORP NEW CL A 032095101 50,169 522,921 SH - DFND 1 0 522,921 0

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ANADARKO PETE CORP COM 032511107 459 6,500 SH - DFND 1,4 0 6,500 0

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ANALOG DEVICES INC COM 032654105 205,927 1,824,461 SH - DFND 2 0 1,824,461 0

ANALOG DEVICES INC COM 032654105 22,699 201,105 SH - DFND 14 0 201,105 0

ANALOG DEVICES INC COM 032654105 70,330 623,103 SH - DFND 1 0 622,903 200

ANALOG DEVICES INC COM 032654105 508 4,500 SH - DFND 1,4 0 4,500 0

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ANALOG DEVICES INC COM 032654105 458,558 4,062,707 SH - DFND 15 0 4,062,707 0

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ANALOG DEVICES INC COM 032654105 11 100 SH - DFND 5,6 0 100 0

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ANAPTYSBIO INC COM 032724106 60,332 1,069,330 SH - DFND 1 0 984,890 84,440

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ANDEAVOR LOGISTICS LP COM UNIT LP INT 03350F106 2,830 77,893 SH - DFND 15 0 77,893 0

ANDERSONS INC COM 034164103 8,975 329,490 SH - DFND 15 0 329,490 0

ANGIODYNAMICS INC COM 03475V101 7,276 369,504 SH - DFND 15 0 369,504 0

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ANHEUSER BUSCH INBEV SA/NV SPONSORED ADR 03524A108 159,432 1,801,293 SH - DFND 2 0 1,801,293 0

ANHEUSER BUSCH INBEV SA/NV SPONSORED ADR 03524A108 1,369 15,471 SH - DFND 14 0 15,471 0



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ANIXTER INTL INC COM 035290105 12,423 208,058 SH - DFND 15 0 208,058 0

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ANNALY CAP MGMT INC COM 035710409 4 439 SH - DFND 1,15 0 439 0

ANNALY CAP MGMT INC COM 035710409 16,598 1,817,909 SH - DFND 15 0 1,817,909 0

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ANNALY CAP MGMT INC COM 035710409 62 6,800 SH - DFND 11 0 6,800 0

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ANSYS INC COM 03662Q105 57,342 279,962 SH - DFND 1 0 276,717 3,245

ANSYS INC COM 03662Q105 195 954 SH - DFND 1,3 0 954 0

ANSYS INC COM 03662Q105 127,015 620,129 SH - DFND 15 0 620,129 0

ANSYS INC COM 03662Q105 82 400 SH - DFND 11 0 400 0

ANTARES PHARMA INC COM 036642106 1,156 351,440 SH - DFND 15 0 117 351,323

ANTERO RES CORP COM 03674X106 34 6,208 SH - DFND 14 0 6,208 0

ANTERO RES CORP COM 03674X106 2,960 535,263 SH - DFND 15 0 535,263 0

ANTHEM INC COM 036752103 96,291 341,203 SH - DFND 2 0 341,203 0


ANTHEM INC COM 036752103 1,316,479 4,664,891 SH - DFND 1 0 4,394,568 270,323

ANTHEM INC COM 036752103 2,879 10,200 SH Call DFND 1 0 0 0

ANTHEM INC COM 036752103 7,311 25,907 SH - DFND 1,12 0 25,907 0




ANTHEM INC COM 036752103 20 72 SH - DFND 1,15 0 72 0

ANTHEM INC COM 036752103 417 1,477 SH - DFND 1,6 0 1,477 0


ANTHEM INC COM 036752103 3,936 13,946 SH - DFND 5 0 354 13,592


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ANTHEM INC COM 036752103 338 1,196 SH - DFND 11 0 1,196 0


ANTHEM INC COM 036752103 169 600 SH Call DFND 6 0 0 0



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ANTERO MIDSTREAM CORP COM 03676B102 248,120 21,650,951 SH - DFND 1 0 21,650,951 0

ANTERO MIDSTREAM CORP COM 03676B102 186 16,219 SH - DFND 1,11 0 16,219 0

ANTERO MIDSTREAM CORP COM 03676B102 1,312 114,442 SH - DFND 15 0 114,442 0

ANWORTH MORTGAGE ASSET CP COM 037347101 1,771 467,312 SH - DFND 1 0 280,812 186,500

ANWORTH MORTGAGE ASSET CP COM 037347101 233 61,500 SH - DFND 1,3 0 61,500 0

ANWORTH MORTGAGE ASSET CP COM 037347101 1,288 339,951 SH - DFND 15 0 339,951 0

APACHE CORP COM 037411105 179 6,182 SH - DFND 14 0 6,182 0

APACHE CORP COM 037411105 24,728 853,573 SH - DFND 1 0 853,573 0

APACHE CORP COM 037411105 3 110 SH - DFND 1,15 0 110 0

APACHE CORP COM 037411105 68,992 2,381,493 SH - DFND 15 0 2,381,493 0

APACHE CORP COM 037411105 474 16,369 SH - DFND 5 0 0 16,369

APACHE CORP COM 037411105 54 1,856 SH - DFND 11 0 1,856 0

APACHE CORP COM 037411105 9,161 316,219 SH - DFND 6 0 316,219 0

APARTMENT INVT & MGMT CO CL A 03748R754 15,876 316,761 SH - DFND 1 0 315,461 1,300

APARTMENT INVT & MGMT CO CL A 03748R754 7 140 SH - DFND 1,9 0 140 0

APARTMENT INVT & MGMT CO CL A 03748R754 660 13,161 SH - DFND 1,15 0 13,161 0

APARTMENT INVT & MGMT CO CL A 03748R754 178,000 3,551,485 SH - DFND 15 0 3,551,485 0

APARTMENT INVT & MGMT CO CL A 03748R754 48 966 SH - DFND 5 0 0 966

APARTMENT INVT & MGMT CO CL A 03748R754 24 480 SH - DFND 5,6 0 480 0

APELLIS PHARMACEUTICALS INC COM 03753U106 1,033 40,779 SH - DFND 15 0 40,779 0

APERGY CORP COM 03755L104 16,665 496,872 SH - DFND 1 0 496,522 350

APERGY CORP COM 03755L104 701 20,891 SH - DFND 1,11 0 20,891 0


APERGY CORP COM 03755L104 155 4,625 SH - DFND 5 0 0 4,625

APERGY CORP COM 03755L104 4 120 SH - DFND 5,6 0 120 0




APOGEE ENTERPRISES INC COM 037598109 1,006 23,169 SH - DFND 14 0 23,169 0

APOGEE ENTERPRISES INC COM 037598109 4,013 92,388 SH - DFND 15 0 92,388 0

APOLLO GLOBAL MGMT LLC CL A SHS 037612306 40 1,169 SH - DFND 14 0 1,169 0

APOLLO GLOBAL MGMT LLC CL A SHS 037612306 51,018 1,487,401 SH - DFND 1 0 1,487,401 0

APOLLO GLOBAL MGMT LLC CL A SHS 037612306 7,551 220,138 SH - DFND 15 0 220,138 0

APOLLO INVT CORP COM NEW 03761U502 1,580 99,995 SH - DFND 15 0 99,995 0

APOLLO COML REAL EST FIN INC COM 03762U105 5,114 278,100 SH - DFND 14 0 278,100 0

APOLLO COML REAL EST FIN INC COM 03762U105 2,987 162,441 SH - DFND 1 0 162,441 0

APOLLO COML REAL EST FIN INC COM 03762U105 26 1,400 SH - DFND 1,3 0 1,400 0

APOLLO COML REAL EST FIN INC COM 03762U105 33,301 1,810,823 SH - DFND 15 0 1,810,823 0

APOLLO COML REAL EST FIN INC NOTE 4.750% 8/2 03762UAB1 250 250,000 PRN - DFND 1 0 250,000 0

APOLLO SR FLOATING RATE FD I COM 037636107 257 17,208 SH - DFND 14 0 17,208 0

APOLLO TACTICAL INCOME FD IN COM 037638103 2,975 201,695 SH - DFND 14 0 201,695 0

APPLE INC COM 037833100 5,802 29,313 SH - DFND 2 0 29,313 0



APPLE INC COM 037833100 1,076,076 5,436,924 SH - DFND 1 0 5,252,081 184,843

APPLE INC COM 037833100 1,324 6,691 SH - DFND 1,2 0 6,691 0


APPLE INC COM 037833100 158 800 SH - DFND 1,9 0 800 0


APPLE INC COM 037833100 258 1,303 SH - DFND 1,15 0 1,303 0


APPLE INC COM 037833100 8,068,442 40,766,179 SH - DFND 15 0 40,766,179 0

APPLE INC COM 037833100 47,655 240,781 SH - DFND 5 0 59,851 180,930


APPLE INC COM 037833100 62 314 SH - DFND 5,11 0 314 0

APPLE INC COM 037833100 20,085 101,481 SH - DFND 5,6 0 11,018 90,463


APPLE INC COM 037833100 459 2,319 SH - DFND 6 0 2,319 0


APPFOLIO INC COM CL A 03783C100 10 100 SH - DFND 1 0 100 0

APPFOLIO INC COM CL A 03783C100 8,591 84,005 SH - DFND 15 0 84,005 0

APPLE HOSPITALITY REIT INC COM NEW 03784Y200 3,686 232,382 SH - DFND 1 0 232,382 0

APPLE HOSPITALITY REIT INC COM NEW 03784Y200 195 12,300 SH - DFND 1,9 0 12,300 0

APPLE HOSPITALITY REIT INC COM NEW 03784Y200 13,513 852,017 SH - DFND 15 0 852,017 0

APPLIED INDL TECHNOLOGIES IN COM 03820C105 67 1,093 SH - DFND 14 0 1,093 0

APPLIED INDL TECHNOLOGIES IN COM 03820C105 6,937 112,749 SH - DFND 15 0 112,749 0

APPLIED MATLS INC COM 038222105 2,632 58,608 SH - DFND 14 0 58,608 0

APPLIED MATLS INC COM 038222105 644,730 14,356,039 SH - DFND 1 0 14,184,441 171,598

APPLIED MATLS INC COM 038222105 341 7,600 SH - DFND 1,4 0 7,600 0

APPLIED MATLS INC COM 038222105 11 256 SH - DFND 1,15 0 256 0

APPLIED MATLS INC COM 038222105 500,624 11,147,271 SH - DFND 15 0 11,147,271 0

APPLIED MATLS INC COM 038222105 1,510 33,633 SH - DFND 5 0 0 33,633

APPLIED MATLS INC COM 038222105 133 2,956 SH - DFND 5,4 0 2,956 0

APPLIED MATLS INC COM 038222105 8 180 SH - DFND 5,6 0 180 0

APPLIED MATLS INC COM 038222105 204 4,538 SH - DFND 11 0 4,538 0

APPLIED OPTOELECTRONICS INC COM 03823U102 1,256 122,190 SH - DFND 15 0 122,190 0

APTARGROUP INC COM 038336103 71,849 577,842 SH - DFND 15 0 577,842 0

APPROACH RESOURCES INC COM 03834A103 61 211,627 SH - DFND 15 0 211,627 0

AQUA AMERICA INC COM 03836W103 2,746 66,369 SH - DFND 14 0 66,369 0

AQUA AMERICA INC COM 03836W103 1,483 35,859 SH - DFND 1 0 35,859 0

AQUA AMERICA INC COM 03836W103 114,173 2,759,808 SH - DFND 15 0 2,759,808 0

AQUA AMERICA INC UNIT 04/30/2022 03836W202 13,801 242,200 SH - DFND 1 0 219,500 22,700

AQUANTIA CORP COM 03842Q108 231 17,724 SH - DFND 1 0 17,724 0

ARAMARK COM 03852U106 33 922 SH - DFND 14 0 922 0

ARAMARK COM 03852U106 3,963 109,896 SH - DFND 15 0 109,896 0

ARAMARK COM 03852U106 43 1,200 SH - DFND 11 0 1,200 0

ARBOR RLTY TR INC COM 038923108 2,800 231,054 SH - DFND 1 0 231,054 0

ARBOR RLTY TR INC COM 038923108 2,037 168,058 SH - DFND 15 0 168,058 0

ARCBEST CORP COM 03937C105 470 16,720 SH - DFND 14 0 16,720 0

ARCBEST CORP COM 03937C105 1,769 62,930 SH - DFND 1 0 26,330 36,600

ARCBEST CORP COM 03937C105 354 12,600 SH - DFND 1,3 0 12,600 0

ARCBEST CORP COM 03937C105 5,917 210,491 SH - DFND 15 0 210,491 0

ARCH COAL INC CL A 039380407 308,304 3,272,518 SH - DFND 1 0 3,247,218 25,300

ARCH COAL INC CL A 039380407 867 9,200 SH - DFND 1,3 0 9,200 0

ARCH COAL INC CL A 039380407 4,162 44,179 SH - DFND 15 0 44,179 0

ARCELORMITTAL SA LUXEMBOURG NY REGISTRY SH 03938L203 379 21,017 SH - DFND 1 0 21,017 0

ARCELORMITTAL SA LUXEMBOURG NY REGISTRY SH 03938L203 138 7,661 SH - DFND 15 0 7,661 0

ARCHER DANIELS MIDLAND CO COM 039483102 5,159 126,453 SH - DFND 14 0 126,453 0

ARCHER DANIELS MIDLAND CO COM 039483102 201,580 4,940,698 SH - DFND 1 0 4,829,949 110,749

ARCHER DANIELS MIDLAND CO COM 039483102 734 17,993 SH - DFND 1,9 0 17,993 0

ARCHER DANIELS MIDLAND CO COM 039483102 1,689 41,386 SH - DFND 1,15 0 41,386 0

ARCHER DANIELS MIDLAND CO COM 039483102 189,134 4,635,628 SH - DFND 15 0 4,635,628 0

ARCHER DANIELS MIDLAND CO COM 039483102 864 21,176 SH - DFND 5 0 0 21,176

ARCHER DANIELS MIDLAND CO COM 039483102 109 2,681 SH - DFND 11 0 2,681 0

ARCHROCK INC COM 03957W106 1,031 97,289 SH - DFND 14 0 97,289 0

ARCHROCK INC COM 03957W106 75,351 7,108,613 SH - DFND 1 0 7,108,613 0

ARCHROCK INC COM 03957W106 23 2,200 SH - DFND 1,3 0 2,200 0

ARCHROCK INC COM 03957W106 7,639 720,619 SH - DFND 15 0 720,619 0

ARCOSA INC COM 039653100 4,891 129,975 SH - DFND 15 0 129,975 0

ARCONIC INC COM 03965L100 98,919 3,831,116 SH - DFND 1 0 3,748,184 82,932

ARCONIC INC COM 03965L100 348 13,484 SH - DFND 1,9 0 13,484 0

ARCONIC INC COM 03965L100 3 124 SH - DFND 1,15 0 124 0

ARCONIC INC COM 03965L100 52,580 2,036,390 SH - DFND 15 0 2,036,390 0

ARCONIC INC COM 03965L100 55 2,132 SH - DFND 11 0 2,132 0

ARES MANAGEMENT CORPORATION CL A COM STK 03990B101 40,861 1,561,369 SH - DFND 1 0 1,561,369 0

ARES MANAGEMENT CORPORATION CL A COM STK 03990B101 11 438 SH - DFND 15 0 438 0

ARENA PHARMACEUTICALS INC COM NEW 040047607 4,182 71,330 SH - DFND 1 0 26,130 45,200

ARENA PHARMACEUTICALS INC COM NEW 040047607 885 15,100 SH - DFND 1,3 0 15,100 0

ARENA PHARMACEUTICALS INC COM NEW 040047607 7,762 132,382 SH - DFND 15 0 132,382 0

ARENA PHARMACEUTICALS INC COM NEW 040047607 282 4,807 SH - DFND 5 0 0 4,807

ARGAN INC COM 04010E109 1,002 24,697 SH - DFND 15 0 24,697 0

ARES CAP CORP COM 04010L103 2,333 130,071 SH - DFND 14 0 130,071 0

ARES CAP CORP COM 04010L103 8,251 459,923 SH - DFND 15 0 459,923 0

ARES CAP CORP NOTE 4.625% 3/0 04010LAW3 11,919 11,600,000 PRN - DFND 1 0 10,600,000 1,000,000

ARES COML REAL ESTATE CORP COM 04013V108 751 50,571 SH - DFND 1 0 50,571 0

ARES COML REAL ESTATE CORP COM 04013V108 53 3,600 SH - DFND 1,3 0 3,600 0

ARES COML REAL ESTATE CORP COM 04013V108 2,113 142,183 SH - DFND 15 0 72,892 69,291

ARES DYNAMIC CR ALLOCATION F COM 04014F102 4,983 329,325 SH - DFND 14 0 329,325 0

ARES DYNAMIC CR ALLOCATION F COM 04014F102 710 46,927 SH - DFND 15 0 46,927 0

ARGENX SE SPONSORED ADR 04016X101 58 410 SH - DFND 15 0 410 0

ARGENX SE SPONSORED ADR 04016X101 1,424 10,058 SH - DFND 6 0 10,058 0

ARGENX SE SPONSORED ADR 04016X101 421 2,973 SH - DFND 6,9 0 2,973 0

ARISTA NETWORKS INC COM 040413106 3,402 13,105 SH - DFND 14 0 13,105 0


ARISTA NETWORKS INC COM 040413106 4 17 SH - DFND 1,15 0 17 0


ARISTA NETWORKS INC COM 040413106 358 1,380 SH - DFND 5 0 0 1,380

ARISTA NETWORKS INC COM 040413106 78 300 SH - DFND 11 0 300 0

ARLINGTON ASSET INVT CORP CL A NEW 041356205 12,008 1,745,326 SH - DFND 15 0 1,745,326 0

ARLO TECHNOLOGIES INC COM 04206A101 1,133 282,623 SH - DFND 15 0 282,623 0

ARMADA HOFFLER PPTYS INC COM 04208T108 22,537 1,361,777 SH - DFND 15 0 1,361,777 0

ARMOUR RESIDENTIAL REIT INC COM NEW 042315507 347 18,611 SH - DFND 14 0 18,611 0

ARMOUR RESIDENTIAL REIT INC COM NEW 042315507 1,125 60,380 SH - DFND 1 0 34,780 25,600

ARMOUR RESIDENTIAL REIT INC COM NEW 042315507 220 11,800 SH - DFND 1,3 0 11,800 0

ARMOUR RESIDENTIAL REIT INC COM NEW 042315507 36,474 1,956,754 SH - DFND 15 0 1,956,754 0

ARMSTRONG FLOORING INC COM 04238R106 1,050 106,636 SH - DFND 15 0 106,636 0

ARMSTRONG WORLD INDS INC NEW COM 04247X102 87 900 SH - DFND 1 0 900 0

ARMSTRONG WORLD INDS INC NEW COM 04247X102 10,303 106,002 SH - DFND 15 0 106,002 0

AROTECH CORP COM NEW 042682203 105 52,898 SH - DFND 15 0 52,898 0

ARQULE INC COM 04269E107 4,254 386,400 SH - DFND 1 0 386,400 0

ARQULE INC COM 04269E107 880 79,941 SH - DFND 1,12 0 79,941 0

ARQULE INC COM 04269E107 135 12,300 SH - DFND 1,3 0 12,300 0

ARQULE INC COM 04269E107 3,978 361,353 SH - DFND 15 0 361,353 0

ARRAY BIOPHARMA INC COM 04269X105 40,034 864,109 SH - DFND 1 0 864,109 0

ARRAY BIOPHARMA INC COM 04269X105 1,331 28,739 SH - DFND 1,11 0 28,739 0



ARRAY BIOPHARMA INC COM 04269X105 1,558 33,639 SH - DFND 6,2 0 33,639 0

ARRAY BIOPHARMA INC COM 04269X105 678 14,637 SH - DFND 6,9 0 14,637 0

ARROW ELECTRS INC COM 042735100 18,166 254,896 SH - DFND 15 0 239,872 15,024

ARROW ELECTRS INC COM 042735100 32 446 SH - DFND 11 0 446 0

ARROWHEAD PHARMACEUTICALS IN COM 04280A100 107,210 4,045,660 SH - DFND 1 0 4,045,660 0

ARROWHEAD PHARMACEUTICALS IN COM 04280A100 27 1,000 SH - DFND 1,3 0 1,000 0

ARROWHEAD PHARMACEUTICALS IN COM 04280A100 20,510 773,947 SH - DFND 15 0 773,947 0

ARTESIAN RESOURCES CORP CL A 043113208 1,023 27,509 SH - DFND 15 0 27,509 0

ARTISAN PARTNERS ASSET MGMT CL A 04316A108 10,798 392,387 SH - DFND 15 0 392,387 0

ASBURY AUTOMOTIVE GROUP INC COM 043436104 7,028 83,325 SH - DFND 15 0 83,325 0

ASCENA RETAIL GROUP INC COM 04351G101 3,453 5,661,002 SH - DFND 15 0 5,661,002 0

ASCENDIS PHARMA A S SPONSORED ADR 04351P101 113,632 986,817 SH - DFND 1 0 986,817 0

ASCENDIS PHARMA A S SPONSORED ADR 04351P101 1,822 15,826 SH - DFND 1,12 0 15,826 0

ASCENDIS PHARMA A S SPONSORED ADR 04351P101 1,313 11,400 SH - DFND 1,3 0 11,400 0

ASCENDIS PHARMA A S SPONSORED ADR 04351P101 35 303 SH - DFND 15 0 303 0

ASHFORD HOSPITALITY TR INC COM SHS 044103109 1,470 494,789 SH - DFND 15 0 494,789 0

ASHLAND GLOBAL HLDGS INC COM 044186104 97 1,208 SH - DFND 14 0 1,208 0

ASHLAND GLOBAL HLDGS INC COM 044186104 39,326 491,757 SH - DFND 15 0 491,757 0

ASPEN TECHNOLOGY INC COM 045327103 80,075 644,311 SH - DFND 1 0 633,211 11,100

ASPEN TECHNOLOGY INC COM 045327103 14,988 120,595 SH - DFND 15 0 120,595 0

ASSERTIO THERAPEUTICS INC COM 04545L107 796 230,670 SH - DFND 1 0 135,270 95,400

ASSERTIO THERAPEUTICS INC COM 04545L107 132 38,200 SH - DFND 1,3 0 38,200 0

ASSERTIO THERAPEUTICS INC COM 04545L107 2,742 794,669 SH - DFND 15 0 794,669 0

ASSOCIATED BANC CORP COM 045487105 820 38,800 SH - DFND 1 0 0 38,800

ASSOCIATED BANC CORP COM 045487105 5,526 261,386 SH - DFND 15 0 261,386 0

ASSURANT INC COM 04621X108 26,597 250,015 SH - DFND 1 0 224,468 25,547

ASSURANT INC COM 04621X108 1,688 15,872 SH - DFND 1,15 0 15,872 0

ASSURANT INC COM 04621X108 56,927 535,131 SH - DFND 15 0 535,131 0

ASSURANT INC COM 04621X108 32 300 SH - DFND 11 0 300 0

ASSURANT INC 6.50% CONV PFDD 04621X207 14,568 130,700 SH - DFND 1 0 118,600 12,100

ASTEC INDS INC COM 046224101 2,980 91,525 SH - DFND 15 0 91,525 0

ASTRAZENECA PLC SPONSORED ADR 046353108 204,782 4,960,801 SH - DFND 1 0 4,866,111 94,690

ASTRAZENECA PLC SPONSORED ADR 046353108 11,570 280,293 SH - DFND 1,12 0 280,293 0

ASTRAZENECA PLC SPONSORED ADR 046353108 2,811 68,087 SH - DFND 15 0 68,087 0

ASTRONICS CORP COM 046433108 1,075 26,721 SH - DFND 15 0 26,721 0

AT HOME GROUP INC COM 04650Y100 341 51,172 SH - DFND 15 0 51,172 0

ATHENEX INC COM 04685N103 1,077 54,376 SH - DFND 15 0 54,376 0

ATKORE INTL GROUP INC COM 047649108 16,130 623,494 SH - DFND 1 0 623,494 0

ATKORE INTL GROUP INC COM 047649108 383 14,800 SH - DFND 1,3 0 14,800 0

ATKORE INTL GROUP INC COM 047649108 2,775 107,248 SH - DFND 15 0 107,248 0

ATLANTIC CAP BANCSHARES INC COM 048269203 455 26,557 SH - DFND 15 0 26,557 0

ATLANTIC UN BANKSHARES CORP COM 04911A107 3,204 90,675 SH - DFND 15 0 60,550 30,125

ATLAS AIR WORLDWIDE HLDGS IN COM NEW 049164205 4,505 100,914 SH - DFND 15 0 100,914 0

ATLAS AIR WORLDWIDE HLDGS IN NOTE 2.250% 6/0 049164BH8 7,903 8,000,000 PRN - DFND 1 0 8,000,000 0

ATLAS AIR WORLDWIDE HLDGS IN NOTE 1.875% 6/0 049164BJ4 883 860,000 PRN - DFND 1 0 0 860,000

ATMOS ENERGY CORP COM 049560105 12,977 122,933 SH - DFND 14 0 122,933 0

ATMOS ENERGY CORP COM 049560105 17,961 170,152 SH - DFND 1 0 157,052 13,100

ATMOS ENERGY CORP COM 049560105 61 575 SH - DFND 1,11 0 575 0

ATMOS ENERGY CORP COM 049560105 193,368 1,831,831 SH - DFND 15 0 1,831,831 0

ATMOS ENERGY CORP COM 049560105 625 5,917 SH - DFND 5 0 0 5,917

ATMOS ENERGY CORP COM 049560105 63 600 SH - DFND 11 0 600 0

ATRICURE INC COM 04963C209 11,149 373,630 SH - DFND 1 0 373,630 0

ATRICURE INC COM 04963C209 257 8,626 SH - DFND 15 0 8,626 0

ATRION CORP COM 049904105 4,511 5,290 SH - DFND 15 0 5,290 0

AURORA MOBILE LTD ADS 051857100 127 26,783 SH - DFND 15 0 26,783 0

AUTODESK INC COM 052769106 40 248 SH - DFND 14 0 248 0

AUTODESK INC COM 052769106 51,847 318,272 SH - DFND 1 0 253,018 65,254

AUTODESK INC COM 052769106 10 63 SH - DFND 1,15 0 63 0

AUTODESK INC COM 052769106 445,341 2,733,833 SH - DFND 15 0 2,733,833 0

AUTODESK INC COM 052769106 1,167 7,163 SH - DFND 5 0 0 7,163

AUTODESK INC COM 052769106 180 1,104 SH - DFND 11 0 1,104 0

AUTOHOME INC SP ADR RP CL A 05278C107 63,593 742,734 SH - DFND 4 0 742,734 0

AUTOHOME INC SP ADR RP CL A 05278C107 59,331 692,958 SH - DFND 4,9 0 692,958 0

AUTOHOME INC SP ADR RP CL A 05278C107 4,119 48,112 SH - DFND 1,4 0 48,112 0

AUTOHOME INC SP ADR RP CL A 05278C107 409 4,775 SH - DFND 1,5 0 4,775 0

AUTOHOME INC SP ADR RP CL A 05278C107 28,507 332,953 SH - DFND 15 0 104,978 227,975

AUTOHOME INC SP ADR RP CL A 05278C107 6,542 76,408 SH - DFND 5 0 64,857 11,551

AUTOHOME INC SP ADR RP CL A 05278C107 445 5,194 SH - DFND 5,9 0 5,194 0

AUTOHOME INC SP ADR RP CL A 05278C107 1,461 17,059 SH - DFND 13 0 17,059 0

AUTOLIV INC COM 052800109 34 478 SH - DFND 14 0 478 0

AUTOLIV INC COM 052800109 1,230 17,441 SH - DFND 1,6 0 17,441 0

AUTOLIV INC COM 052800109 2,118 30,043 SH - DFND 15 0 30,043 0

AUTOLIV INC COM 052800109 324 4,600 SH - DFND 5 0 0 4,600

AUTOLIV INC COM 052800109 28 400 SH - DFND 11 0 400 0


AUTOLIV INC COM 052800109 57 814 SH - DFND 6,2 0 814 0

AUTOLIV INC COM 052800109 4,035 57,220 SH - DFND 6,9 0 57,220 0

AUTOLIV INC COM 052800109 333 4,728 SH - DFND 6,3 0 4,728 0


AUTOMATIC DATA PROCESSING IN COM 053015103 73,467 444,366 SH - DFND 14 0 444,366 0

AUTOMATIC DATA PROCESSING IN COM 053015103 33 200 SH Call DFND 14 0 0 0

AUTOMATIC DATA PROCESSING IN COM 053015103 99 600 SH Call DFND 14 0 0 0

AUTOMATIC DATA PROCESSING IN COM 053015103 265 1,600 SH Call DFND 14 0 0 0

AUTOMATIC DATA PROCESSING IN COM 053015103 280,523 1,696,743 SH - DFND 1 0 1,687,143 9,600

AUTOMATIC DATA PROCESSING IN COM 053015103 83 502 SH - DFND 1,2 0 502 0

AUTOMATIC DATA PROCESSING IN COM 053015103 1,653 10,000 SH - DFND 1,4 0 10,000 0




AUTOMATIC DATA PROCESSING IN COM 053015103 780,111 4,718,508 SH - DFND 15 0 4,718,508 0

AUTOMATIC DATA PROCESSING IN COM 053015103 11,118 67,247 SH - DFND 5 0 44,094 23,153

AUTOMATIC DATA PROCESSING IN COM 053015103 310 1,877 SH - DFND 5,4 0 1,877 0

AUTOMATIC DATA PROCESSING IN COM 053015103 2,993 18,105 SH - DFND 5,6 0 2,048 16,057

AUTOMATIC DATA PROCESSING IN COM 053015103 347 2,101 SH - DFND 11 0 2,101 0

AUTOMATIC DATA PROCESSING IN COM 053015103 561 3,391 SH - DFND 6 0 3,391 0

AUTOMATIC DATA PROCESSING IN COM 053015103 2,652 16,040 SH - DFND 13 0 16,040 0

AUTONATION INC COM 05329W102 71,619 1,707,660 SH - DFND 1 0 1,707,660 0

AUTONATION INC COM 05329W102 12,

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