Turfgrass: Biology, Use, and Management › resources › files › Downloads › pdf ›...
Transcript of Turfgrass: Biology, Use, and Management › resources › files › Downloads › pdf ›...
Turfgrass: Biology, Use, and Management
John C. Stier, Brian P. Horgan, and Stacy A. Bonos, Editors
Book and Multimedia Publishing Committee
April Ulery, ChairWarren Dick, ASA Editor-in-ChiefE. Charles Brummer, CSSA Editor-in-ChiefAndrew Sharpley, SSSA Editor-in-ChiefMary Savin, ASA RepresentativeMike Casler, CSSA RepresentativeDavid Clay, SSSA Representative
Managing Editor: Gail K. Schmitt
Agronomy Monograph 56
Copyright © 2013 by American Society of AgronomySoil Science Society of America Crop Science Society of America
ALL RIGHTS RESERVED. No part of this publication may be reproduced or transmi ed in any form or by any means, electronic or mechanical, including photocopying, recording, or any information storage and retrieval system, without permission in writing from the publisher.
The views expressed in this publication represent those of the individual Editors and Authors. These views do not necessarily refl ect endorsement by the Publisher(s). In addition, trade names are some-times mentioned in this publication. No endorsement of these products by the Publisher(s) is intended, nor is any criticism implied of similar products not mentioned.
American Society of AgronomySoil Science Society of America Crop Science Society of America, Inc.5585 Guilford Road, Madison, WI 53711-5801 USA
agronomy.org • soils.org • crops.orgdl.sciencesocieties.orgSocietyStore.org
Agronomy Monograph Series ISSN 0065-4663 (print)ISSN 2156-3276 (online)
ISBN: 978-0-89118-613-7 (print)ISBN: 978-0-89118-614-4 (electronic) doi:10.2134/agronmonogr56Library of Congress Control Number: 2013931617
Cover design: Patricia Scullion Cover image: Turfgrass growing in heavy shade at Caves Valley Golf Club, outside of Baltimore, MD. Photo courtesy of John Kaminski.
Printed in the United States of America.
v
Contents
Foreword xiPreface xiiiContributors xv
1Origins of North American Turfgrasses 1
James B Beard
2The Turfgrass Industry 37
James M. BreuningerMark S. WelterlenBruce J. AugustinVan ClineKevin Morris
3Turfgrass Benefits and Issues 105
John C. StierKurt SteinkeErik H. ErvinFrancis R. HigginsonPeter E. McMaugh
4Turfgrass Extension and Outreach Programming 147
Aaron J. PattonLaurie E. TrenholmF. Clint Waltz, Jr.
5Synthetic Turf 179
Thomas J. SerensitsAndrew S. McNittJohn C. Sorochan
6Management of Turfgrass in Shade 219
David S. GardnerRyan M. Goss
vi
7Heat-Stress Physiology and Management 249
Michelle DaCostaBingru Huang
8Low-Temperature Stress 279
Annick BertrandYves CastonguayAïda AzaiezJulie Dionne
9Turfgrass Water Use and Physiology 319
Kelly L. KoppYiwei Jiang
10Turfgrass Ecology 347
Kurt Steinke Erik H. Ervin
11Turfgrass Root Zones: Management, Construction Methods, Amendment Characterization, and Use 383
Cale A. BigelowDouglas J. Soldat
12Characterization, Development, and Management of Organic Matter in Turfgrass Systems 425
Roch E. Gaussoin William L. BerndtColm A. DockrellRhae A. Drijber
13Nitrogen Research in Turfgrass 457
Kevin W. FrankElizabeth A. Guertal
14Potassium and Phosphorus Research in Turfgrass 493
Kevin W. FrankElizabeth A. Guertal
Turfgrass: Biology, Use, and Management
vii Contents
15Secondary Nutrients and Micronutrient Fertilization 521
Rodney A. St. JohnNick E. ChristiansHaibo LiuNicholas A. Menchyk
16Warm-Season Grasses: Biology and Breeding 543
Wayne HannaPaul RaymerBrian Schwartz
17Cool-Season Grasses: Biology and Breeding 591
Stacy A. BonosDavid R. Huff
18Development of Native Grasses for Turf 661
Eric WatkinsLeah BrilmanDavid Kopec
19Genetic and Genomic Approaches for Improving Turfgrass 683
B. Shaun BushmanScott E. Warnke
20Biology and Applications of Fungal Endophytes in Turfgrasses 713
William A. MeyerMónica S. TorresJames F. White, Jr.
21Advances in Turfgrass Pathology since 1990 733
James P. KernsLane P. Tredway
22Turfgrass Weed Management 777
J. Scott McElroy Prasanta C. Bhowmik
viii Turfgrass: Biology, Use, and Management
23Turfgrass Insect Pests 809
R. Chris WilliamsonDavid W. HeldRick BrandenburgFred Baxendale
24Insecticides, Fungicides, Herbicides, and Growth Regulators Used in Turfgrass Systems 891
Zachary J. ReicherPeter H. DernoedenDouglas S. Richmond
25Integrated Pest Management 933
Albrecht M. KoppenhöferRichard LatinBenjamin A. McGrawJames T. BrosnanWilliam T. Crow
26Sustainable Turfgrass Management in an Increasingly Urbanized World 1007
Paul G. JohnsonFrank S. RossiBrian P. Horgan
27Study and Management of Turfgrass Traffic Stress 1029
James A. MurphyJ. Scott Ebdon
28Irrigation Science and Technology 1075
Bernd LeinauerDale A. Devitt
29Digital Image Analysis in Turfgrass Research 1133
Douglas E. KarcherMichael D. Richardson
ix Contents
30The Evolution of Spectral Sensing and Advances in Precision Turfgrass Management 1151
Gregory E. BellJason K. KruseJoseph M. Krum
31Research Tools and Technologies for Turfgrass Establishment 1189
Deying LiJason HendersonJ. Tim VaniniJohn N. Rogers, III
Index 1241
xi
Foreword
Turfgrass in a well-maintained lawn, on a sports playing fi eld, or underfoot at a public park did not grow there by accident. A large body of science has informed its development, management, and appearance. Science has pro-vided for turfgrasses that are tolerant of environmental stresses, diseases,
and insect pests and that also have a beautiful appearance. Basic and applied research has led to scientifi cally sound technologies that minimize management inputs to turfgrasses, while maximizing obvious and less noticeable benefi ts, enabling turf to be grown in many diff erent contexts. That science is on ready display in this excellent monograph, which greatly expands on two previous edi-tions to provide the most comprehensive overview of turfgrass science published to date. Complete with information on the latest advances, this monograph cov-ers numerous topics barely discussed twenty years ago when the last edition was issued and updates existing topics with the latest information. All aspects of turf are covered, from basic biology and plant breeding to applied turf management and how Extension promotes sustainable turfgrass management in an increas-ingly urbanized world.
This comprehensive book is an invaluable source of information for research-ers, students, and practitioners, as well as an excellent aid to teaching worldwide. As this book shows, turfgrass science is alive and well, producing valuable infor-mation to make turfgrass more resilient in the face of climate change, more beautiful, and more valuable to the world. The entire turfgrass community should be proud of these accomplishments; society at large appreciates your eff ort.
As editors-in-chief of the Societies, we are indebted to the editors of this vol-ume, John Stier, Brian Horgan, and Stacy Bonos for their outstanding eff orts to bring this book to completion and to the authors for their synthesis of an ever-increasing body of literature. This book collectively refl ects their considerable hard work and dedication to furthering the fi eld of turfgrass. This will be the go-to turfgrass book for years to come.
E. Charles BrummerEditor-in-Chief, Crop Science Society of America
Warren DickEditor-in-Chief, American Society of Agronomy
Andrew SharpleyEditor-in-Chief, Soil Science Society of America
xiii
Preface
This book was produced for students, researchers, consultants, regulators, and turfgrass management professionals. Many chapters are extensive reviews of the literature and will be helpful to those seeking a comprehensive view of a subject such as turfgrass heat tolerance or management of shaded turf.
Other chapters, such as “Turfgrass Extension and Outreach Programming,” summarize examples of key issues and programs, along with the need for, and outcomes of, state and federal investment in turfgrass science. Chapters were developed to serve as stand-alone entities for teaching purposes and are avail-able electronically from the publisher. This feature may prove to be a harbinger of the publishing future: before this book’s publication, we were already receiving requests for specifi c chapters.
The previous two versions of this book, both titled Turfgrass, were published in 1969 and 1992. During each of the approximately 20 yr between versions, the number of turfgrass researchers and the amount of data produced increased exponentially. The depth and breadth of knowledge expanded, and new areas of research were developed. On the plant side, the advent of molecular biology gave researchers tools for understanding the genetic underpinnings of turfgrass behavior. Advancements in engineering technologies allowed researchers to more precisely, and quickly, measure such aspects of turf as soil moisture, temperature, and traction for sporting purposes. Knowledge about turf in the landscape from a social sciences perspective exploded, too, with greater awareness of the value of turf to diff erent social factors, ranging from human behavior to direct economic activity and employment. Since 1992, the ability to share knowledge as a result of computers and the Internet has catalyzed enormous amounts of research. In the United States, the Turfgrass Information File database was transformed from a DOS-based, dial-up system accessible to only a few researchers in 1992 to a full-fl edged web-based, key-word searchable database called “the best database for a subject” by a librarian at the University of Wisconsin–Madison. By the late 1990s, the Internet’s various search engines allowed people across the world immedi-ate access to publications on turfgrass science, management, and news. By the mid-2000s, government agencies and municipalities were also readily shar-ing information on turf management and regulations. Bans on turf pesticides in Canada emboldened pesticide-free lawn activists in the United States, while lawn phosphorus restrictions in Minnesota initiated a chain reaction copied and expanded by such states as Wisconsin and New Jersey. Turfgrass researchers and extension (outreach) specialists are increasingly being asked to provide data to regulatory agencies on topics as varied as invasive grasses in the United States to drought and irrigation in Australia. Data sharing among researchers from a face-to-face perspective grew, too. In 1992, the International Turfgrass Research Society was the only international, formal venue for researchers to present data. During the 2000s, the International Sports Turf Conference and the European
xiv Turfgrass: Biology, Use, and Management
Turfgrass Society formed, providing additional venues for sharing data with international colleagues.
Readers of the previous monographs in the series will recognize updated chapters from previous versions, such as those on synthetic turf, shade, and the turfgrass industry. Other topics have been greatly expanded. For example, the 1992 Turfgrass monograph had three chapters on turfgrass soil, nutrients, and water; the new monograph has seven chapters on the three subjects. “Turfgrass Benefi ts and Issues” broadens the theme of turfgrass benefi ts fi rst published by J.B Beard and R.L. Green in a 1994 Journal of Environmental Quality article. A num-ber of new topics are being included for the fi rst time. “Turfgrass Extension and Outreach Programming” is the fi rst documentation of its kind devoted to turf-grass. The chapters on native grasses and genomics address research areas that were in their infancy in 1992. Chapters on irrigation technology, digital analysis, and spectral sensing document how research and technology are changing the way turf is managed. In accord with the public movement for more sustainable industries, “Sustainable Turfgrass Management in an Increasingly Urbanized World” describes strategies for reducing reliance on synthetic products for turf management while maintaining well-kept turf for pleasure, recreation, and envi-ronmental benefi ts.
Numerous books and other publications have been produced for students, homeowners, and other groups in the past 20 yr as society has become increas-ingly urbanized, and turf, primarily in the form of lawns, has garnered more public a ention due to its maintenance costs and perceived environmental impacts. Our mission was to have some of the world’s best turfgrass researchers and educators summarize the state of knowledge and activities in their particu-lar areas. The outcome of this eff ort was meant to accomplish a special vision: enhanced public awareness of the scientifi c underpinnings of turfgrass use and management, thereby allowing people to make the best possible decisions regarding turfgrass as part of the human environment.
John C. Stier, co-editorPlant Sciences Department University of Tennessee–KnoxvilleKnoxville, TN 37996
Brian P. Horgan, co-editorDepartment of Horticultural ScienceUniversity of MinnesotaSt. Paul, MN 55108
Stacy A. Bonos, co-editorDepartment of Plant Biology and PathologyRutgers UniversityNew Brunswick, NJ 08901
xv
Contributors
Augustin, B.J.Sco s MiracleGro Company 17578 Raymond RoadMarysville, OH 43040 ([email protected])
Azaiez, A.So ils and Crops Research and
Development CentreAgriculture and Agri-Food Canada2560 Hochelaga Blvd.Québec City, QuébecCanada G1V 2J3 ([email protected])
Baxendale, F.Dep. of EntomologyUniv. of Nebraska–Lincoln202 Plant Industry Bldg.Lincoln, NE 68583 ( [email protected])
Beard, J.BProfessor emeritusTe xas A&M Univ. and International
Sports Turf InstituteCollege Station, TX 77840 ([email protected])
Bell, G.E.De p. of Horticulture and
Landscape ArchitectureOklahoma State Univ.358 Ag HallStillwater, OK 74078 ([email protected])
Berndt, W.L.De p. of Resort and Hospitality
ManagementFlorida Gulf Coast Univ.Fort Myers, FL 33965-6565 ([email protected])
Bertrand, A.So ils and Crops Research and
Development CentreAgriculture and Agri-Food Canada2560 Hochelaga Blvd.Québec City, QuébecCanada, G1V 2J3 ([email protected])
Bhowmik, P.C.Dep. of Plant, Soil, and Insect SciencesUniv. of Massachuse sAmherst, MA 01003 ([email protected])
Bigelow, C.A.Dep. of AgronomyPurdue Univ.915 W. State St.West Lafaye e, IN 47907 ([email protected])
Bonos, S.A.Dep. of Plant Biology and PathologyRutgers Univ.Foran Hall59 Dudley Rd.New Brunswick, NJ 08901 ([email protected])
Brandenburg, R.Dep. of EntomologyNorth Carolina State Univ. 3301 Gardner HallBox 7613Raleigh, NC 27695 ([email protected])
Breuninger, J.M.Dow AgroSciences LLC9330 Zionsville RoadIndianapolis, IN 46268 ([email protected])
Brilman, L.Seed Research of Oregon27630 Llewyllyn Rd.Corvallis OR 97333 ([email protected])
Brosnan, J.T.Dep. of Plant SciencesUniv. of Tennessee252 Ellington Plant Sciences Bldg.2431 Joe Johnson Dr.Knoxville, TN 37996 ([email protected])
xvi
Bushman, B.S.US DA-ARS Forage and Range Research
Laboratory695 N 1100 ELogan, UT 84322-6300 ([email protected])
Castonguay, Y.So ils and Crops Research and
Development CentreAgriculture and Agri-Food Canada2560 Hochelaga Blvd.Québec City, QuébecCanada G1V 2J3 ([email protected])
Christians, N.E.Dep. of HorticultureIowa State Univ.133 Horticulture Bldg.Ames, IA 50011 ([email protected])
Cline, V.The Toro Company8111 Lyndale Ave. South Bloomington, MN 55420 ([email protected])
Crow, W.T.Entomology and Nematology Dep.Bldg. 970, Natural Area Dr.Univ. of FloridaGainesville, FL 32611 (wtcr@ufl .edu)
DaCosta, M.Stockbridge School of AgricultureUniv. of Massachuse s17 Stockbridge Hall80 Campus Center WayAmherst, MA 01003 ([email protected])
Dernoeden, P.H.De p. of Plant Science and
Landscape Architecture1112 H.J. Pa erson HallUniv. of Maryland, College Park, MD 20742 ([email protected])
Devi , D.School of Life SciencesUniv. of Nevada, Las Vegas4505 Maryland Pkwy.Las Vegas, NV 89154 (Dale.Devi @unlv.edu)
Turfgrass: Biology, Use, and Management
Dionne, J.Syngenta Crop Protection AGSchwarzwallee 215P.O. Box, CH-4002 Basel, Swi erland ([email protected])
Dockrell, C.A.Te agasc College of Amenity HorticultureNational Botanic GardensGlasnevinDublin, Ireland ([email protected])
Drijber, R.A.Dep. of Agronomy and HorticultureUniv. of Nebraska–LincolnLincoln, NE 68583-0915 ([email protected])
Ebdon, J.S.Stockbridge School of AgricultureUniv. of Massachuse s22B Stockbridge HallAmherst, MA 01003-9246 ([email protected])
Ervin, E.H.De p. of Crop and Soil Environmental
SciencesVi rginia Polytechnic Institute
and State Univ.330 Smyth HallBlacksburg, VA 24061 ([email protected])
Frank, K.W.Dep. of Crop and Soil SciencesMichigan State Univ.584E Plant and Soil SciencesEast Lansing, MI 48824 ([email protected])
Gardner, D.S.Dep. of Horticulture and Crop ScienceOhio State Univ.2021 Coff ey Rd.Columbus, OH 43210-1086 ([email protected])
Gaussoin, R.E.Dep. of Agronomy and HorticultureUniv. of Nebraska–Lincoln202 Keim HallLincoln, NE 68583-0915 ([email protected])
xvii Contributors
Goss, R.M.De p. of Plant and Environmental
SciencesNew Mexico State Univ.P.O. Box 30001Las Cruces, NM 88003-8001 ([email protected])
Guertal, E.A.Dep. of Agronomy and SoilsAuburn Univ.253 Funchess HallAuburn, AL 36849([email protected])
Hanna, W.Dep. of Crop and Soil SciencesUniv. of Georgia115 Coastal WayTifton, GA 31793-0748 ([email protected])
Held, D.W.De p. of Entomology and
Plant PathologyAuburn Univ.301 Funchess HallAuburn, AL 36849 ([email protected])
Henderson, J.De p. of Plant Science and
Landscape ArchitectureUniv. of Connecticut1376 Storrs Road Unit 4067Storrs, CT 06269-4067 ([email protected])
Higginson, F.R.Turfgrass Scientifi c Services Pty Ltd.14 Carolyn Ave.Carlingford NSW 2118Australia([email protected])
Horgan, B.P.Dep. of Horticultural ScienceUniv. of Minnesota254 Alderman Hall1970 Folwell Ave.St. Paul, MN 55108 ([email protected] )
Huang, B.Dep. of Plant Biology and PathologyRutgers Univ.301 Foran Hall59 Dudley Rd.New Brunswick, NJ 08901 ([email protected])
Huff , D.R.Dep. of Plant SciencePennsylvania State Univ. 116 A.S.I. Bldg.College Park, PA 16802 ([email protected])
Jiang, Y.Dep. of AgronomyPurdue Univ.915 West State St.West Lafaye e, IN 47907([email protected])
Johnson, P.G.Dep. of Plants, Soils, and ClimateUtah State Univ.4820 Old Main HillLogan, UT 84322-4820 ([email protected])
Karcher, D.E.Dep. of HorticultureUniv. of Arkansas316 Plant Sciences Bldg.Faye eville, AR 72701 ([email protected])
Kerns, J.P.Dep. of Plant PathologyNorth Carolina State Univ.2510 Thomas HallRaleigh, NC 27695-7616 ([email protected])
Kopec, D.School of Plant SciencesUniv. of Arizona1140 E. South Campus Dr.Tucson, AZ 85721-0036 ([email protected])
Kopp, K.Dep. of Plants, Soils, and ClimateUtah State Univ.4820 Old Main HillLogan, UT 84322-4820([email protected])
xviii Turfgrass: Biology, Use, and Management
Koppenhöfer, A.M.Dep. of EntomologyRutgers Univ.Blake Hall, 93 Lipman Dr.New Brunswick, NJ 08901 ([email protected])
Krum, J.M.300 11th Ave. NorthNashville, TN 37203 ([email protected])
Kruse, J.K.Dep. of Environmental HorticultureUniv. of Florida1541 Fifi eld HallGainesville, FL 32611(jkk@ufl .edu)
Latin, R.Dep. of Botany and Plant PathologyPurdue Univ.Lilly Hall915 W. State St.West Lafaye e, IN 47907 ([email protected])
Leinauer, B.Dep. of Extension Plant SciencesNew Mexico State Univ.N230 Skeen Hall945 CollegeLas Cruces, NM 88003([email protected])
Li, D.Dep. of Plant SciencesNorth Dakota State Univ.Fargo, ND 58108 ([email protected])
Liu, H.Sc hool of Agricultural, Forest,
and Environmental SciencesClemson Univ.Clemson, SC 29634 ([email protected])
McElroy, J.S.Dep. of Agronomy and SoilsAuburn Univ.201 Funchess HallAuburn, AL 36849([email protected])
McGraw, B.A.Golf and Plant SciencesState Univ. of New York–DelhiFarnsworth HallDelhi, NY 13753 ([email protected])
McMaugh, P.E.Turfgrass Scientifi c Services Pty Ltd.14 Carolyn Ave.Carlingford NSW 2118Australia ([email protected])
McNi , A.S.Dep. of Plant SciencePennsylvania State Univ.11 6 Agricultural Science and
Industries BuildingUniversity Park, PA 16802 ([email protected])
Menchyk, N.A.Sc hool of Agricultural, Forest, and
Environmental SciencesClemson Univ.Clemson, SC 29634([email protected])
Meyer, W.A.De p. of Plant Biology and Pathology and
Center for Turfgrass ScienceRutgers Univ.59 Dudley Rd.New Brunswick, NJ 08901([email protected])
Morris, K.Na tional Turfgrass Evaluation ProgramBARC-West, Bldg. 005Room 307Beltsville, MD 20705 ([email protected])
Murphy, J.A.De p. of Plant Biology and
PathologyRutgers Univ.Ne w Jersey Agricultural
Experiment Station59 Dudley Rd.New Brunswick, NJ 08901-8520 ([email protected])
xix
Pa on, A.J.Dep. of AgronomyPurdue Univ.915 W. State St.W. Lafaye e, IN 47907 (ajpa [email protected])
Raymer, P.Dep. of Crop and Soil SciencesUniv. of Georgia1109 Experiment St.Griffi n, GA 30223 ([email protected])
Reicher, Z.J.Dep. of Agronomy and HorticultureUniv. of Nebraska–Lincoln161 Keim HallLincoln, NE 68583 ([email protected])
Richardson, M.D.Dep. of HorticultureUniv. of Arkansas316 Plant Sciences Bldg.Faye eville, AR 72701 ([email protected])
Richmond, D.S.Dep. of EntomologyPurdue Univ.901 West State St.West Lafaye e, IN 47907([email protected])
Rogers, J.N., III Dep. of Crop and Soil SciencesMichigan State Univ.A160 Plant and Soil Science Bldg.East Lansing, MI 48823 ([email protected])
Rossi, F.S.Dep. of HorticultureCornell Univ.134a Plant Science BuildingIthaca, NY 14853([email protected])
Schwar , B.Dep. of Crop and Soil SciencesUniv. of Georgia115 Coastal WayTifton, GA 31793-074([email protected])
Contributors
Serensits, T.J.Dep. of Plant SciencePennsylvania State Univ.11 6 Agricultural Science and
Industries BuildingUniversity Park, PA 16802 ([email protected])
Soldat, D.J.Dep. of Soil ScienceUniv. of Wisconsin–Madison245 King HallMadison, WI 53706 ([email protected])
Sorochan, J.C.Dep. of Plant SciencesUniv. of Tennessee2431 Joe Johnson DriveKnoxville, TN 37996([email protected])
St. John, R.A.Ryan Lawn and Tree9120 BartonOverland Park, KS 66214 ([email protected])
Steinke, K.Dep. of Crop and Soil SciencesMichigan State Univ.1066 Bogue St.East Lansing, MI 48824 ([email protected])
Stier, J.C.Plant Sciences Dep.Univ. of Tennessee–Knoxville Knoxville, TN 37996 ([email protected])
Torres, M.S.De p. of Plant Biology and Pathology
and Center for Turfgrass ScienceRutgers Univ.59 Dudley Rd.New Brunswick, NJ 08901 ([email protected])
Tredway, L.P.Syngenta Crop Protection5804 Enchanted Creek CircleKnightdale, NC 27545 ([email protected])
xx Turfgrass: Biology, Use, and Management
Trenholm, L.E.Environmental Horticulture Dep.Univ. of FloridaP.O. Box 110675Gainesville, FL 32611 (letr@ufl .edu)
Vanini, J.T.New Dimensions Turf 9 Colvin Ave. Buff alo, NY 14216 ([email protected])
Wal , F.C., Jr.Dep. of Crop and Soil SciencesUniv. of Georgia1109 Experiment St.Griffi n, GA 30223 (cwal @uga.edu)
Warnke, S.E.US DA-ARS Floral and Nursery
Plants Research Unit10300 Baltimore Avenue Beltsville, MD 20705-2350 (Sco [email protected])
Watkins, E.Dep. of Horticultural ScienceUniv. of Minnesota338 Alderman Hall1970 Folwell Ave.St. Paul, MN 55108([email protected])
Welterlen, M.S. PBI/Gordon Corporation1217 W. 12th St.Kansas City, MO 64101 ([email protected])
White, J.F., Jr. De p. of Plant Biology and Pathology
and Center for Turfgrass ScienceRutgers Univ.59 Dudley Rd.New Brunswick, NJ 08901 ([email protected])
Williamson, R.C.Dep. of EntomologyUniv. of Wisconsin–Madison246 Russell Laboratories1630 Lindin Dr.Madison, WI 53706 ([email protected])