Plant Embrogenesis

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    Plant embryog

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    Process that produces a plant embryofrom a fertilised ovule by asymmetric cell

    division and the differentiation ofundifferentiated cells into tissues.

    Single-celled zygote undergoes a

    programmed pattern of cell division

    resulting in a mature embryo Occurs naturally as a result of sexual

    fertilization and the formation of

    the zygotic embryos.

    Divided up into two phases, the firstinvolves morphogenetic events & second

    phase, or postembryonic development,

    involves the maturation of cells.

    http://en.wikipedia.org/wiki/Zygotehttp://en.wikipedia.org/wiki/Zygotehttp://en.wikipedia.org/wiki/Morphogenesishttp://en.wikipedia.org/wiki/Morphogenesishttp://en.wikipedia.org/wiki/Zygotehttp://en.wikipedia.org/wiki/Zygote
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    seed formation andgermination

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    flower

    Nectar

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    FLOWERING PLANT LIFE

    CYCLE

    Divided into Haploid & Diploid

    generations.

    Haploid generation begins after meiosis

    with spores that undergo mitosis &

    differentiate into either a pollen grain or

    an embryo sac.

    Diploid generation begins after

    fertilisation with the zygote & forms the

    mature plant with vegetative organs.

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    FERTILIZATIONsperm cell (n) + egg cell (n) ZYGOTE (2n)

    EMBRYOTRIPLE FUSION:

    sperm cell (n) + polar nuclei (2n)ENDOSPERM NUCLEUS (3n)

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    TWO FERTILISATION EVENTS OCCUR IN

    FLOWERING PLANTS

    One sperm unites with the egg cell to formzygote & initiate embryogenesis.

    Second unites with a specialised cell withinthe embryo sac to initiate the differentiationof endosperm which is a triploid tissue.

    Endosperm is present during seeddevelopment & provide nutrient to theembryo, the germintaing seedling or both.

    Fertilisation also causes the ovule todevelop into a seed & ovary to differentiateinto fruit.

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    Embryo DevelopmentZygote (within the embryo sac)

    First Division

    Upper cell Lower cell

    Proembryo Basal cell of the Suspensor

    Mass of cells Suspensor Stalk

    (surrounded by endosperm) Suspends embryo into endosperm

    Proembryo, divided into 3 layers

    Protoderm ---> Surface Tissues

    Procambium ---> Vascular Tissues

    Ground Meristem ---> Ground Tissues

    The embr o takes on the sha e of an axis with meristems at both ends

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    PRIMARY ORGAN SYSTEMS IN A MATURE

    EMBRYO

    The AXIS & the COTYLENDON

    Both are composed of 3 tissue layersprotoderm, procambium & groundmeristem.

    These will develop in epidermal, vascular &parenchyma tissue of the young seedling.

    Axis region contain the root & the shootmeristem.

    Cotylendon is terminally differentiatedorgan that accumulates food reserves thatare utilised by the seedling for growth.

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    EMBRYO DEVELOPMENT The first division of the zygote produces a small proembryo cell and

    a larger basal cell that develops the stalked suspensor, which

    elevates the developing Proembryo into the endosperm tissue.

    The embryo meristems are the apical shoot meristem and the apical

    root meristem, from which structures of the shoot system and root

    system will ultimately develop.

    The cotyledons rapidly elongate, and the embryo is divided intoregions, with respect to the cotyledons.

    The region above the attachment of the cotyledons is the epicotyl,

    which contains the apical shoot meristem

    The region below the attachment of the cotyledons is the hypocotyl

    that ends with the rradicle, containing the apical root meristem. Typically the embryonic axis will have to fold, to fit within the embryo

    sac

    Endosperm may or may not be absorbed into the cotyledons. It may

    be consumed completely in the maturation of the embryo, or some

    may remain for germination.

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    EMBRYOGENESIS IN HIGHER PLANT

    THEREFORE SERVES.

    To specify meristem & root-shoot plant

    body pattern.

    To differentiate the primary plant tissue

    types.

    To generate a specialised storage organ

    essential for seed germination.

    To enable the sporophyte to lie dormantuntil the conditions are favourable for

    postembryonic development.

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