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riruun 'ldrrri ln:.:airrtornonfirrirxnarurilur.ra druru uacrfioro{f{'[ri druaunr{ruaa t 'i ^ 4 'ilufrhi dnuilsloqlGnriil[!ofiauri #ns cn t:unnui o'lriunntorrnT nrirJurourinaon ru

, j i- d,,riru6u 1ro{nonfirniiUriludru :cnourorHa uu'jr Ho[{rs n-noqluilrsrRyrro{,.ra fln. d, ; "i zl a a s d A,,^l '(Fleshy Fruit) ril u H nr6 u :fi ur' o mi tgrn u l n 16l fi udri ru' orj u ua t an o {1u il : v rR r l o { rro3 o

(eritt) ,iluHoand.rrdofifurJ:yvrru'lrlGun'irrroia rniqurorndruto.rrlf,ndrroiqoonur

fio{rrrun6 (outsrowrh of Seed) uacfirflSnrfilriJuril6onrior.?lo{#uuon6n#unut.r

I

I) ra

t

-

2t

I

:)

?

13

< /' a d AA t(d! [Rru Ho.r ld rlasnflflxl lId tv{f,la. 00u [au.2553) rrJof,nlllljuuun l:[05 [ulelu lnllol,.laJ2l q ?

lnufruqrni rflorir'urni (25s6: 128) ldnrirlir nr:r0SEUrAulnlorflafi 2 gillruu n-ruen':1u

nl rvno! 12 o-{fii,l. nlltoiqtlgtu lgt,lro{Ha g{r?{ (Simple Sigmoid Growth Curve)

nrrri;iy"rel!Iolo{fl4fld.:dx{ 6o r.rnfia'nsflsnr:rnityrer'rlnlnoGrduodrrdr "la , ' d q s ^ - s ia

uatn6nt!6?ut? inl:tnu1u161uon:'ln'l: t0: nilnu lnfllt?!1n unvon:1n1: [!:qilnu In10'tul o c, ,a.i

Hnrvnoair!oHn tnnufl rntYl

2. ntrlo?qttAulflto{flado{d?l @ouble Sigmoid Growth Curve)

nr:ri3;6uIn1o{fl ndo{d?{ 6o runio'nuorsnr:roitgrGrlnlnuGuduodr#r 1a 9 ' A q s - - s j< i - J , q 'rlnunon 1!61uY?{nttlYtu,rJulnluon: tn t:[n: EUtnu lsrYl[:'ullfl [0urnol uclo'tx'].1 [uY']{nnl.t

J - ^ r i< : J a y10.:n1:rn: r9r1l l$t10l?,touu fl1 n: [nu lnuo{flancuogt:]fl{Yl llt u]n10{flalY{ltlulanu0u

oouble slgmold curveSigmold Curv€

,E

,E

RrflrJ:vRou 12 ;iluuunr:[n:qrn'uIntorHa

dut : Physiology rruit . n.d.

9oy9 l,0]ni,0{a:yuyfrl:1rflu'r1o.:[{'ry{,T{ftu r't]yuaflBfusnlr[05 [lJ[gtu tgtlo.tHnuuu14u,tl?t

.ia , a Z ," ?;lnod:rfifinr:roiqrnulnotirr:rorirriuo{lurirlf,rJnrrifi 5-7 ra-.r6on 1u fiHnrrrviv&'c

fl 't5 dl't.:[uo unyfl 1:Itt gulnu [nou1.:: 2n[: ?

- L ' &' a ".inrt otu-t.uo.u.tlv9t,lttottrt 39t{!00uo.ttvuytnutRo?. ! ' !, 4 d i d

fl1t 110tu-lt o.:t.t 1g 9t{([n: u uyn,:10 nu 0,ttyuylfl1[nu?d'llt'l: ftlu uut[flun1fl'lti

6":nrn :snou tz

t4

21 25

au

15

15

lu

11rllJ ,1

Rtfl tyRou 13 t y uynlty{i0tu 110{a on[{ 19

YIU'I :UUs nlnfl ltosflols.2557 : 19

ttt n, h 4 i - . i Afuuloun tufla (oou tau.2552) tnn n1?[nu?nun 1t inn]: [{ ]v[11or1runuR1 10r

nanfin'irnr:dnueirdonaruvna:d'nilarudoilaoonil:yuril llu3 ro{n?ruur?donnriol#fi,'ride

01u?u a tulnu l0 flanotoluoHoolrl 3-4 d]JalH Ha{nonu'ru.iot nl n lnu?flat{1s

A a 4 'd j urqroy

l{ ty[:! flafl]non ruo{ Hnrlfiriuliu? ldl.rt? a ]il: E u 10t 130-160 iu uaco"{uginonA & c ae - J< 'c i hr,15urruoun:yvrl-.:Hd!rfl5 n'ldrlarrJ:su'rru 25-30 iura-roonu ruoun: vrird.: Haurirfrrriur 1d

l{nar too-t zo i'u drri.ll:a16'rnrir:flrrtr:nrilduuuilarldduo{riuanruuandor uovniu{mrvJ id .i t d , .iry{01rl1o.rr'rlyylrnurflu'uJfl?'rlr d1niugr0o1rln 1 n tflu1 ua3fl1:?1tn1 u'ru l.tlyYtf.t0lR1lt

Au '<d I c v d .itasrO*hi6!Harionr5lfi fnu] nr:':rriirrrirtr fiHar{ryfirfirrcot n o nt: rn!rn0?rl{[[da {

lunrn :cnou t: ot'uru'ldornnrnil:cnoil 6o

nafi t :yusu.Jduufi to i'u qilnlflHo6rreiAnnu-ldourfiuhJi,f,fln 2 :rucr nuud l3 ?u q Rlflfl nnrlstri,.lou{oournu-hJj-

'f, - to Lofloyr 3 :rusI nuud t6 ?uf]ruRtytflaoqluln cyl6 liid [[ny:dxtgt

; '.i

. da '--HAfl4 :euyl nu d l9?un Rty{Haoqluln cylo!]n T-{d uny:flslet.i-

'i - e ,.r td l,-HAyr 5 :EUyl auud 22 ?un RlllHnoqlulflOlqlnrJln T{d [[ny:d$19t

j,iofr 6 rsusr Auu? 25 iun nr ,lao{1urnnrd6 rrlrrrit fuflarnlurJ:sryrr;- ri 'flnfr z :cusr 6uu? 28 iununr flnoqjlurnani6 rulrvdrriun nrnluil: srrqi ,- "flnm 8 Hnunlnu lrl

!E

nI

l:!rlat!(a1l:E:tYtO

aiA:tt(ln5iEiti9IiD!

'a

l:6:t1,ail

l5

d1rn?uq n1rlo?qr6ulntoln'r

fi:rov vro,:dr'lvr t2529 : hirJ:rnqraurrirt kinrirrd{so$Irrultr (plant Hormones)

' d, ^ atAa e E e t1 Gr r,o1!n1M{l.t1U tUrA.:'r11n15 ?',t nllldr50ull: uYtlrTd:1{lluroi tull: tJ'lolu 0ulj tn unuHn tun'lu

a - C ,.i e, a Z Z ln ,

n1:d.:Id:11 fi:o uuu.:nttlllauuuunnyll.td::?Ytu1fl1u lu0tultluu 1 Yt{u tlJ:']1lfl1:

J:vrnvrrirnra uioar:ormr:drfluornr:vr'rloun:r d?udr:frrrn:u#fifiqtuallGndru/* -i. d , o ra ^ u Ae, d ' dr /sg0: Iuuf ItuYlglvl lulqo u lliTlYlnnanu E9r'luYr0r0.:n15 tuQn[:ufl?'lluu00: ttiulls

'o d' v tAa .^ , /s . & < nyo s 6u /unnoruud r: fl.tlfr5]v11T!n0ld!!nfl n tu00: t!ufiruu0{ to!n1:ugugunfl YtYtl l?Y'ln r:'r1

I d, d?dltn?Ufl!n1:[Oiqr6UierlO'rn't (Plant Growrh Regulators, PGRS) .r{[UUnq!tO'ldltOua:u

i -g . -Jtnudl:I nlu01nlfln1un'lu tuflY tnun:ifitolfl9,t un1nn1:dnrn:lsfi t9u?nfl::! ]nrflri

: r:our fiurfirr uasnuv (1.il.il. : t-a) nrir.l'i rsof llun'sfi rilufi ifinri'u6 rroc

finr:rirurl#niuotir.rn{xtrxrfufioqj s n{l ldud oon6u 6rnor:afiu lryIyrlnfiu ,ovri6u-'- a /q L ,' !, -uosdlruuu{nl:[r:qlnu t0r10{Tri uonn]n0o: tuunq fian 1 r{ 5 nQUuii?ru ilr!uuuur0n

r n{l fiinr:dnutniu!1n rravuo!fu'jrriluaofllufirn{l1rri drluournnotiinlrdlxrld

odtn{rrt'lrrmrrni:rnun: ldud rL:rfi lua16urouri ii'dluluyr uasryrAlryla

1 2 3.+,l1

ffiF',Trr

\

ilt

U,.--i$l&'.:,,

r\l .(fffi,, ; t

4 F'3t'^

.

L

orfl rjrvRou tl ft fi afi nlrrgclcionr:rfi uriua.i ^u t amlJl : dt'lol nU llflno{. 2557 :28

l6

- 4 t 6et I t t.duflYlD tfl:U!?if, (2549 : 50) tonn12? lgot tIJUY{tlln'l 11,fl1fltUflOn'l:11OlU11O lH n

' -e- , d,nluu5snl:u ritunnltnofla nlt1ulu1u]o1o.l a n1:rQn:?.11o{flo nufl:vvl{flltqnlolHn

b ' , i, " - {: i zs a , tv.t'B{n:EU?unrrsr1.:1lfinrud?uun!n?]!dlfliuT.tdu n1:Yrelr1l1:ofi?ufllJU0: trJulltt alu tn- d, A .i" - -i ..e - l' et F h,

1lr[1lUd.:mdln yrdM:nyrllr4R']un!Hnflaflmln'lull:tJl ungn fl111tft

d,, - ")^ a i ,

tirriunr: [nd1: ri{[n:rr11Ylud']uHfllj10id1:n'luq!n1:[nt qlnu tn10lfln [ 0n?0

o 3 e, .a A .a d A -lunr:toruturnto.r aur-u do{dr'lu:ruvfrHadnixu rnrdnts luioqjlud'r ix oint: tni iUrAulsr

nlr ltl dl: lount {nufla00ury{oq tu:vusflln {[0:gul9lu tgr tJd1u12u t?ifln!1ulfi tt4fuxu:l--iat;ctArtar

lfro{n1no f 1r d10.t d1: Yt! Hnn0nl: Iru {lcxnafii 0n1r 1u1u1u]n10 i rsn aflu: s n0ulultjuJ^

riionn (ohyrn f,idsoin uaudrRr rirr{sr. zss: : sz-s:)

rJyt ln.uD{dl:n?ununl:to30,lAuIfl to.t oirionltrYfl ul uovlytixx 1nt,o.tfl o

- 4 ? ns t 4 ' dlriuflYr! rfllu!?.rfl (2549: 52) [nnn l?n.:n1:1ulurju19t O{ n2lrtjUflArJ',rllnnlruu {n']

r;, t a' I .* + /<uasfl'lfiJu'tu1uloljoflcraaYr :!noulultJulu 0Ha s.:Qnn'lrnu tfru80: tuul,{1n]u tu n 3 suff

6o oonfiu 6urror:afiu uau'lsln'[nfiu ]noiur6nrflurrurirairraoflluddrritylnurnnrr6 d' - .ia

oonryu [r0s0uruolrn0u r1]udr:fr'luq!nlrrntqlau [911o.tvlIvr1r?]nflon1:lIsrurq nrY{

flafl46 uonolnnd!rordr:n?uq!nr:rriqrn'ulntou'vdirLvrrrvreionr:fl?uq!n'ltrtEtUreruleld - zi .ia

lta&1u1n10.:fla[[n ?u { tnrnlt nuY{ud'lt n']uq! n l: [n: qtnu tgr10{v{ixunou dl YllJ,,lan0

nr:rn: ilrgl'uInto{flnd?u rdu r:rfiIuar6u:sori (Bra-ssinosteroids) dl:o1llqunl:lni ryreil]nt o l. ;-" , e alrnYltH0n0nr:YlRru1 [[nylYurllulnl0tH0 lnrln

1. oon6u (Auxins)

-Z ' a dl!1u [noflqn (2550 : 49-52) tnnnl??1 o0nryul udl:n?Uq!n]ttn:qlnU tn1o{nn

, i i, - i, tten rj u u: n fr l qu ririu n u rfr n r o.r c r: fr du u rL ldud IAA n*{ ufl n { lu n ryr il : v n ou t + luiJ

< / - A a,.4 dln.fl. 1880lounrf a nriu fi':qoralri6i,o{oonfrueioyly n-o riludr:n?uq nr:rnirXrn'llnrorn'l.ia - I , i - a :- ,

tflllJlt0Hnnlt,UlJ1t0'l tn tnulJgr'lul1u lYl Hann 0u 0 tuu00 o?u?tylu{00u tnuooflcru.l f, t , ' dr j ie - - d j4

' :i jtnaouYtotnuatuuona{11{41{lduo ruunltrflnouyt luytflytlttFlu? t4:o1l[:un']'l lfln0 ytltuu

ir?r (por-i y) hjinr:t?ouna-rn!odaumlrnlu nr:6nn-:rorrryadrfioldfuar:oonfiu u;r.t6u4qYe, , I - r, zl ^ Cr .in , im tn:u[[fl{!n?1![lJ!1u10 {o0n$uu0u [u0tn'ln 00ncru luud'l:yl [?n0[[d.t!1n [ll-0Qnrld{

rfiolqrunrn riruuirarufihilrifrualifl?'lrlrdu? ro.rDonfiulrnn.jr rryadlSnorfihilrifuual41o-rInr:dnd'urnn'jt (Audesilk et aI ; 2002 : 521) ; (Mader. 1993 : 525) uonnlnfioonauu-,r1ii{1ur4tr< t ? 4 i o, - ^ - ! -ldri: nouauo'rciorryn.irow{-ylnu tu'rnrnru.: r0 urm n{uu!n'rruriluhJld'ir nalanr:?i'r,r'lu

ro.roofl 6utfi nloun:{fi ,{rf{rrynd (ctetana. rlls :'hirJ:rn6raurrir;

t7

Al.l itolo{oonorufl oRlflotqtfl ulgtl,o{tTt!a d, i-

oonryultjud'l:Yt!Hanont:[n:qrnlj tgr10{1r1 tuHnru 1 6]u [Yu flxuq!e4rrred,i^i,

n'l:1u1un?10.:lcr0a nlxq!n1: uon']l0.llcraa nt3q nl:r[1J.:n?uo.tlcra n tuluoluoln: qmog, 't , e

:vfi21.:vr0 1 [rov1roo11r1, n:vriunl:rna:1n uocnl:[n:qt0{t1n Y?u tunltr0:tu tud?u0r1i dl

torx'r i6vrinarionr:roicgtornrrhr nrrqlnr:ro;EX [[avn1: qnjr.rto.rlu ri.:raiu,jiade

n1:00nn0n nrtr nuurllflo0n [11!n1:aa a:?!0{n]tfr?ufllJfl'r:116ru'r10{fla rraE

nrsu?unrroiu 1 fitfiurr?ori:rurru'jroonfiufirirudrn''tylunr:#nrjraft oNe luurqrtilar

tlngnl:ttU {[$0ni,0 {[tn: OYI n1fln]:flU UO0nryUn'][[5n fl0 Indole Acetic Acid (lAA) ryil U

^ .ia v t rv t00ficruT Sf,lr,1:0d51'ttU tn[0.:n1!!::Ui1n n0! 1r,n1l fl{lfl:]yl'l Indole Butyric Acid

(tBA) rros Naphthalene Acetic Acid rNne) durLrlnurqud dr wr#u ld'jrn:;uau nr: eirr 1t, Ao n .i

luvlYo0ncru!d?ua?u tun'l:nlueun:vu?un1t9t'l.t 1 n1: tfoonryu tulJ:1r'lslYllt4lrlsdlrti

daulunr:roSqrnr'lotorvr'r uriluvrrn:.rriudrrLnr:ldoonsulurJilrtufilrnrfiu'[rJovfinrn

" -! - t dc, - / .lunl:0'ru.:n'r:[0: qrnu t9r6?u (:tldqun n]?nu. 2540 :79); (duglr, rfl:u!']rrfl.2549: 15)

oonAufl r!rr0urjloonriiur nrun{l fi r nrunrjrufi io?rudrriru rrnufi u!f 1ri r1d

lumtrnr: rnun: rutrlrtiu6o nrjlrunvrrdu (Naphrhalene) n-.:uanrlunrxrJ:rnou l5 d{[iJu- i 4v a! - o o e,rq e a

f, rtoonfiufi r,UUtif,rrn:rclifiu fr ulrirl rl{lunr:n:sriu: rnn.:ilnrir 1dn:v{ulunr:oon

nonro{dudr:o rirl#durJv:aooflnonilgorniu ilo.rniunr:irlro.:nn'h?nnrorfin nJfiou

rlrflnonrorflAxulrrun l-n1ffinr:finriuh{6'tfiryvrth aur:odrr.irurdrbllurujoru'or.r'r,,rk s a i^1d6 InuoonSunrjuuilillAufifiuxdr:.Jrl{yn'tnr:m!m:'lduri Nee (0, - naphthalene acetic acid)

ar e' A ,i-riludu 6.:ar:nreirfrHa-nrr'tuI:Jnr:rilfiorulfin rdu uuauIufind (plunofi**) In:nda.- -- o. I ra .4 ] o A ' ! . e e(Grc..Plus-) [Y{0tfl-dnlf,Yr (Ferti-Srart*) "X'rflr:[xn 1, tylJnll![{!{uno{dl: NAA [[nnsn{nU

(111U [9f 0df.]4. 2550 : 57)

OHc

RrnrjryRou ts In:laitlto,t Indole Acetic Acid (tAA).i,frrr : :r:our d'uytimr unsnory. tL.rJ.rl. :8

H{o

NI

H

t8

Naphllultnc aclds

c-Nephttrlantroedcrid PNephthrlorcreticecld(o.NAA) (0-NAA)

nrnrjrvRou ro ln:.rairltolar:d.rrn:rtr{n{l Naphthalene Acid

drl , tjt"'tor'l fiuv{mrunvnorv. l.rj.rl. : t t

1.2 t-tu?ou tnu?no{nuooflo t{a o- io y a s t tA!.: r u T o-u fr I rio o n 6 ri! y{-x 14 o r u l fi n I o o l' r-n q rJ r u d { fi fr ttn n gi 1.t fiu

9a[[ns n u v{]l H n1 0 { 0 0 ncuu 9l o1ITO.tU

ri'lqvrd rn:ux r.rd (25a9 : 53)'[doi utcrj t i n r: f; nu rnt: ld Nee ludrJr]s :n

nujrfi :vaiuorrlrdurlu t oo !n.ln. d?urfi rrturnxo{fl aduils:n'[d

u1n (Bhar er al. 1997 : t:so)'ldf,nurHntolnr:ldoonfiuriudu6 ru'jr nr:l#' 1)- t L F ' tracaont: anir{1o.:Hn'[dOOnryU 40-50 xn./d. AqflUOUn UI Dehmdum d.tlfl]rJnl:On A

n$r (ctost et al. 1990: r:s) ldfrnurxu'jr nr:ldoonfiu zo un.ln. rirbi2A-r - -.i0UnlIut Bombai 910tldflfl Qn

q: {io l'rnrin un&nuc e522 : 45) ldfl nu rdr6vrinnxo.tdr:rnieionr:,i-?<+a/dar 6 uurugno nt o.rr.r r; n'uf 1: rG ouvt r ir unauluiln"i il6vrinnsiont:t iuut qoonorn

nonaur;:oirunhhilunonure{lddrunr:tiutirnr:rfio ungdtar0f.i'ldto{nond!Utoirilf,l#e e.aee - A

fl qn:1.r tll drrld:rJn1:[n:q10.:[nd:9r?4YI1 trnlulounyoo.:lfl f, 5u1?1u

fi:rnry nordrln (2529 : 4s) r,ril'ir nr:lfunnuIuf, nd fl nriu}Jfi rionon,.,rr,,o,rd

nronrira-rurulnulfnrunirriu soo rLn-ra. rirl#nonursnJ6uunrs'ldnruiln-.rnr:biar: o a'u

z. 6utuor:aiu lGibberellins)

i,o"nn-u1u;rJto': c,r, rilun{ltorar:n'ruqunl:r03qrn^uIsrfiifion::leion'lgrA , t tdt .---s i,' / s , a I d s c, i,r:lJflufl a{rlnlJ fl.f,. 1890 tnull?ulruqu 6ufluntn l:n{1?3uor4u.r Fun [:nti'r? ?1 ulnlr

-t(Bakane) (dlqvri ra:untld z5l9 ' 2r) lufl n.s. 1926 E. Kurosawa fnnqnuaranflraflrlu ld

l9

rfluofln{rurdu?niul:arorciundrrhrfirir}idundrq.: ucinsrdu:rqtdur(u uavnrolufiqne e e ^ I :i , -nlfln1:fln!1 ltu?'l l:n10{n fl61.[1?n{nn1?[nn!]flnl:n6[Yo:1Tn01'l sutu0[:aal {nQ l:o

I & ^ a jj , ^ - a i.ov-cr{Hn{d'l:lunHu{ylro?l n ruol:anu cr{flltu1l1t tnn01n1:10{ t:Rn.:nn'l? (qlm qua?ol1u1.

at )a ,' 3 t ^ d e hv2533 :478) iUllJ0l:naul udltYlllxd:1.:1,ulo't tO [[As[!05111'l{rUgnfl]!'ltflf,:1i tn

- ' !- ' d, - .i- ared1:fi'tuflufll:ln: qlnu lolorntnQrlurJrJ lnn?l 60 xun lln GAr rlluru9rvlutJlJ trnu!]ni * A - o v . o,ylqn tnu GAr ! lfl:.:fl5'r{nrlrdn{ [uRrlrlj:snou l6 cr{ nuru0[t a auuuYrulYr tunl:xnul tlJ

te { - e a v 4 , t. cyl Aa0{ul'tt u a{$lfln1td:1nu0o n'lun!n1: unnxlo tlcrna fl]5ltu{tcraa ,?u tl,llu0tuo[n:q

d e 9 ^ :un1:rn:furgr1I tgr n1u tufllt{ofl10{r!an rrnv ti,fl1a1un1: nn210.:lttu1{Iun lfl!n'l:nofln

2.1 Haro{6uruo[roauoionr:ro;alfi ulnto.rf,r

^l i ---U1u [nodQ0 (2550 : 8G9l) tnnal?tfl U?nU0Ylr11A10'td'l:tUlU0[:n0uel0Y{Y2'l

I a e | 4 o Y . ct to , ad1:n tuo[:oou el]r,1:on:vqunl:unn? tlovnl:ttu.:lcrnatJo.lalou Yl] tfinlnuunul?,- e-

'A o e i^trutnu?nu0onryu [[nuuu.:n]:lfla:1n t !ry{ol'1flu1'lunro0ncru ttfio.td'1tfl:'lvfin]J[uot:oou

a .i" - o o <lufir 6o rirurorrun flaeiou r!6n fi rir a-r roi tyur^u 1er nruuon luludoutou'ry un&lu:rnt,otod'.i.'Aecr.rt'lflrl n{drlfi:'tu11 tuylr u11 tu[! a600uu drru l1-lutll]fuq{ tu fl?u10 {fl'ltlfl n0uB 10

A v ,. ' ! ,i-!ulu0[:nau lu1lt lJn1:lflaou01uyr.r tuTorj] lla3r001fi1:ll9rui l!xn1:flIlJfltart^tre?r!utuo[:ooulfl nouuruuuuln

..toHOH

CHzcFI3 CoOH

ar :snou lZ In:.:Airltol GA,

dul ,d:lrour ir'um{mruaynary. r.rj.rl. :2I

o

H

H

20

2.2 .t]u2oa lnax.uo{nuouluoltaa

-- ia r tA'rlu2dufr fl nu1HolJo{nutu ol: naunul{t n'luYu9 [nuu?fl qu: s d{nYl

unngilidu6uruor:aaufir,ra'nrntucilunr:1{6o n?:lddrtlounl:6nriuar:o.tuuo't-orsfi, oya , s hi/ q9 "-60{nr:lrifrsuun0uf,u0'r lnunr{ n?uann1:qEUltiut0{d1: [q unsfl?t llld1:nnnonu 3-4 R:{

turfi rerdoaqn. zsso : ss) dri:rurru'irru'oririrruor:nfiuhJldnluojurfu{go,noi,nvrri duilu- t;i,a d 'd d h,ht, a oc,

11U{yr ltJ ![U AA[[OrJrJU',rOlO{flnrnn tU tnUlnr!1U nnu:]n0?1nutu0[:nnun1 t tlolJlulo

" ,l - I r A , s o a 4 o hlilfutu (ai!6lyr! lflru!?'rfl. 2549 :24) cr{donn a0 tnufl'lotululo{1t: [o1 Yl0{01 111 (2529 :

hirJ:rnnrntnr?r) ldoiuru'irinr:l#ar: Ge, otnnruuonuir}ISorfr-urJBltorto.t cR,r,tndu

rirr.ral#nr:turururnro {rryadrfl'rr rirl#rfi lturnto'rnnld

rJ:rIut irrqt (2526 : umflntjo) ldrirnr:finurvrvjr dl:asoru6!luot:aau

fi:yriunarrunlrLrlu 250 un./n. dMtod{fl'l:rilAuuAxo're{ar{rvvriufl:rrSuukirirn'irhildcr:

t-z iuq: fi n n^ 13 qa uo r n olr (2s 42 : 186-199 ) ldn rirar6uariu rmn lu lainr: n 6 n

or< I Ia0{fl0t tfi!nolR 1lt ?'l tun'l: d.tld: ullolu lfi'l:10'!r0o0 n n'l:qnY{u GArn?'l![1u1u 100

!n./a. fidononluo-nrr 200 r,n./n. 1udxfi{lnrnondudncr'rriiudouun,,ulud, olr,roae I ne

Y?UUOXOnOn tfiU',l?1U tn

ixvr o n'er: u: : urri unun u e553 : 4zt -424)'ldrirnr:dnu rn r:no! du o {tqJ

touoojuniuf uu"oo seeatess sionr:riu GA5 flu'jr nr:1d c4 finrunilriu so ln.n. rir}1cl,a

ft n0luululo tlliu Itnv!nlt1loru"tl0tluanu0uYr g0

rfi q:fi vrol6uvrf rras:i rdr! n-n6 (2542 : 4349) fi ldvrnao.:1# cn,, rirHorl:ol

riu{nnla16 rJ:rnn'i Hafiriuar: cA! fl1ur:nralnr:roityrnuln'ldlrnn'jrHafihildriuar:A . s L s 6 d h, .i a

rl=orhornm r rdru or r unsdrun{rrtorHo uasu-rdrrr:onndrururrfinn,rldlrnfiqn fio

rr6orf,o'r t t t nrHndoHaurjorGuurfiurriuHafihildiuarr 6o tss ruf,adoHn

ieia (siswas et al. lgg0 : 152-156) ti}?ar: cernrrunlurlu so un.ra. rildi,r

r{u{ l-+e ru'fi rirh?naturnlnqjdun'jrriunruql

orfi nti ni lauyd'roqa uav:i ra:1n-n6 e54z : 45\'Iffi nurnatorar: Ge,t&Anta d v r

rionr:dnnn rrasnr:mTorsrulnrotr.radf.:fihifinrfinvluriur,rnonrrorJlflo vrl'ir cAr d'lur:04 | Jd t a

' , v j j ! a

rur=lr oir urinr:n-nr.ranrtnarlr{triutolatt frrn-uiu fio zs, so, 75 rrac 100 run.lo. Iouriu

1u:yovn n'rlrnnont ru s riu arlr:nrfiuru rorua1d6fian

s ae a hY< j - q, ;SrnrR:ol nu 'rflt uasn u.(2550: urnnuo) lnflnlllnu2nuflalo{nl: t! GAr Yl

t e - tl I qe -- 4 ecy f, 1,lJflanoqolR'1111o{[[n?!{n:lruIruoti 11 vru?'l fl'l: tfidl: GAr !Rnm1 t?ifl']]lJU1? ul unr{a

fl ?ru?rur uagfrroudoud:r-.:n: rnauunn'jrnr:'hjl{ar: c,q,.

A,tii?1Yt 'l uavrytn (Srivastana and Singh. 1969 : l-6) n1t1{dr: c4 luduini{'l

6nno + drlnrri drun?rrru1rrriu 25 un./a. uau 50 !n./n. rirb?turnto rnalnqjdu/h

rlnof uoln e (Thakur et al. l99l :212\ t6:lu'i lunl:l{dr: ca, rfio:J:-urJ1lJi, " '- o Ji- t .

nunl 'lJo.tt{naun?1n1t1ru GA3 tudntl 50 !n./n. ilU?U 5 R:.: tuauolluI P'nbi uAv Deshi Yll

oe d o 'ltfiflar,1uln tfiruluda

3, Ull lUdlSlUrOUn (Brassinosteroids)AJ

u:'rrt tudtnu:ouo (Brassinosreroids nlo sRs) lttniiu: nlioluri2rfl 1970 Inud' a ,.i n v-

narun?ru0'rn usDA [uudl:fl?uqrnl:[n: Eulnu tn,t o{T1sflQ! tfilJYr ln ruf]1:0aa lnu0nn1fl

oonBu 6rrror:a6u lryIvrlnfi u n:auoil'[,r6n uovrofi Au Inofi or:arrrrnnr:trr

ar:rJ:;noun{ldfiin:;ortro{m'rlrJlufirunruvun uovlnuiHacion'llurvn'unrruu?ldurirt rYi v I I t,lauuan{Ha'ldfr: vdrnrtrr{urlu l0'r2 (Picomolar) uny l0'e (Nanomolar) lutorvfiat:nruql

a , :l n c,inlrrnlq,r6ulnro.rnrnril6u 1 dr!rr0uf,n{Ha'ldfi:ydunrrlrdrr?u l0n (Micromolar)'l '

o ;o a da ,i d a,uon 01nufl'r:nouduo{10 {lrxYuJel0fl1: nQ!u:1fl tufllourouau a'luuuu uaE[!0!n1t tl.r

l. , r a f,-dl:UnUd?U tnd?UfiU.tlO{ SnCrn9n',l:01rnUt luU.t9i',llr utnl{ cl flrnanl:9tou du 0ivt'l {

h, e e d - i dc,n??YlUl tli (n:fluuYl nlr!. 2548 : 34) ; (aod n1?nt uarflolt. 2556 :229)

drar::l:ynoun-ru:nfi aflnld f,oir u:r?Iu1nd (Brassinolide r,ti o Bl) dladnorn

ocoo.:tnel:1o{fir01n Alder tsee (l/nraglrn'rara L.) tlaylo.t Rape plant (Brassica napts L) rtaytlhia i 9 a s q a a d

f,l:n?UflUfl1:[0: tl,[nU t9l1o.tfltn?]t:n Uflnt tn:{fl:1{Ylltlflllfl6lUnU Steroid Hormone [lJf,gl?

o-ruaa{1unry{il:rnur,, lnohiiolurinruila-!'it ur1fl6u (Brassins)

3.1 fl oto{urtaluar6u:oudeisnr:ro?qrfiulntol{ir

r:rfi Iuar6u:oud riluar:nruqlnr:ro! ty16ulntoln'vfi ieiru drfirglunr:

rirri'rgqJriudiohi'tr'uoSqrn'lIer'hlotixrJnn^IouiarrrLalul:olunt:fl'runrnl:ufln,!oonro{Auj r o yaYlnot lnnfll:Unul? (Clouse et al. 1992 : 1377-1383) ; (Zurek and Clouse. 1994 : 16l-170)

9-auavnrrflfr{loyr-Au (wang et al. 1993 : 965-968) lu i) a.fl. tslz Mitchelt and Gregory u'ul.j]

4?d'iu: rAIudr6u: ouriilnnrionr:rfilnonfinn"r n?1!f,tutro1un1:ro3€UrrtuInlo.tu't uasnrru

d d 4 r 1 t i- i orul{u:{r,otlrjan (seed vigor) 0u0-.rr] ilriuiluil rut ld ludlau:ouri uacar:frin:rurfiualio,r

riuu:rfiIuarfiu:oudiluvr'rydrurulln unrfiflaeionrruirgrer'r1o rrav 'orurtorfl'sluHaru dl

2l

22

9 ,A - A 9 | 4 4q4t ,, . varu n?1!grnu?nunlrlflriou61u1o'rd1: nq! xr 1d tudlnu: ounu{!noul,r':tlnn un!:1u'l1u

'irar:n{ldaur:orn6oudruorn:rn}Ju-.!rluroifil'ld n'rlunr:rirnr: mnaouvr-xieiuura - a L y s de h r e I h o

[[ay [u:3 nuvtlvt{nuu9llJ t luu nu{ ttJYtt lurn[0u o i na tfln t: T1.t 1u,t 0 {11:'ld tudlglu:oun

( :r:our t'uvrfvrr uavRolv. u. . . : sg) dlgio!r1uii \.fl.2549 n ernua:fl1flnt

urrinurri'urf,uslnri 6qldnisrurar:ndru r.r:refiu (Brassin-like Substance r,ri o gs) durnioil-flur.ni-n,ng e .ui. -o If,

nornl l0iHaal luYtu{ lrl tnuln:!1u lnulay{tunl [uYrYrln1l nfluonqn :or{nn tu uYl

: - .u A .i 6,, <r,

nu01nrfl t![ul,]s du er0fi1: rn: €ureru tnlo.:r{nii1 tu cu.:dll,1:0[ 111u]nHa10{a1 tullju

nodrrl r Inonnrionr:ninrurxo.!fi rlurartr 1 drudlluriruro{nr:fliruurlo{fl a 60

ri.r rai u n r : 6o sr'r l o l r.ua d a o n l r ?r q o i a.: t o r ru a d o u utr ilS l r ar lra E q gtn 1 fl 1 u'Ifl rua rduJi ee .h ^4au0 unsnl tu (fln{to{adl: dulmflnl lu[r{nn.00u tau.2554)

OH

OH

OH

OH Ho

otflrJrcRou l8 qnt]n:{dit{ Brassinosreroid

dr'l ,rlr"'rorl f,uyrfrr uacnily. u.rl.rj. :57

-- i;3.2 {'n{?oulllot ?nuurlcludlslurouri

--;ey as I dq , r;Itu?nuyl tyu:1d tu dtnu: ou6nulIxfi a luyuo tnuuzno :gf,lnytunnnliY ,t q9 - l, a o.lnu [[nsr 1n t 1u.t1u Har 0 t fl'tt fluy{u Ha n 1nfl 1t ttryl:ur 1fl lufllstut 0unn0{suf{,tu

nlfi drrots uayfuu.i'u niuflnulqt (2552 : umn-auo) d,rrirnr:vrnoolldatr,

lJ51dcrunU , tnunt5flflu1fla10,:d1llJ:',lflcrunon1t[lIlJtJuln nlsrit.ruurryun nuirj,n]:qn ud'lru:'ld,uu on:10.025 xn./a. fl1u1:oryM uhfln unyrulnHa'ld

23

n:qi aiiriu: uasnlr n11elruu ti (zsc2, 3B4z) 'ldrirnr:f,nurnrrx{onto.:, u Ja - lf, u, 4 v 4< a J o te a

nvoo{rnd:1o{l,gu?{ 3ltu{ no vluILllnon llmxluluo:4lluI tnflouun rlnvflul!urnouy ' i e r, a aA e tt

ln'r lru?1 nJo lafl'r:u:1fl [ufllnu:oufi frn:urrir{u 0.05 un./a. rirbinr:.ronlo.ravoo{rnf,r. . aZ u, r o r. . t 4 i - t, ,

uv!?tltu{utn0n tlrYr?luluo: 4 uasltul ttnouu$tlltlu unvm:vnun?rrrti!1u 0.1 !n./0.

rirl#nr:.ronto.rasoo{rnd:rrsri'1.:yr-ufniu16ourdrqdu

qua?::0r inufinurn:af uasruvfu n'u#rnuuqt (2sss : 8-t+) 'lddnuri . . r u / ' orl - .i t etflu?nuHa10.td1:x: tdcrun0qolnllt nlotn'l tuflulno flu?'l nlt t d1:u:ldcru fl?ltJruulu

Agsy1.5 lln./n. dltJl:o[fr'!d1ruhfla fl2run2r{Ho nlrur.rurHa n?ru11u1[LJ6on Itnsn2]rltu11o{d nel e v c q t rr, ' o, - f,

lUoflA tq ${flonRA 0{nUr: dUtJyl nltu (2548: UylnnU0) tS5'lu.l]U?1 n]5 ttdltu:ldcruYl

:gd'uoerrrrrludu 0.01 !n./a.d?unt!fl?r!n{r{ n?'u,14u'lro{fla rirnrinar, rirnu-nur?r! - I y l.h hei d s

u'1fi unruoIn4.1 [[nvn?.l 4u'l10.uuon1 tu t6rnurnu2nu! .A. ' ^ :tl n{n tlagflolg (Peng et al. 20M: 4O7416) ct.t?11fl1:llud1tu:'1flryU Tfl?lU

zt s o y- J i[lrilu 0.5,0.75 [[as 1.0 rJn./o.l nunut 1ru?1 fl1tu:ldcrur?uaanl:rlerflt 0{Ha uncA e a o, A

fl "tu1:0[v{u1l:lrl0l nHan tr!1fl 1u

{Iofrr,fi ieasionr:'liar: nruqunr:ro?qrGllorto'r$il

lrfi rn4oaqa (2s50 : ll) ldnrirafirrlor-ufifinndonr:ldar:n'r!qrrnr:roiqrn'uIn

tory{'i'i'l nr:ldf,rtn?uqunr:rniEUrerulnxo'rx'r {1drio.:finrrluhlodrihr'uxoruil:snr:drir}?nr:ldar:rJ:saunadr6o'lduri

Al. tjoouo'lRtrn

r.r rYuqnrartolf,ri , a .qea I A , . t , .oe

,u11UIn:5!YtUnfl nlrnUT1 t nfl !01stJ0't[lJoluounf lnl{fl ucI{d{fl nfl 1 tfiai,!n?1rJdM: 0 tunt:n0ud1Jo{n0d1:fl 1u!!unttln:fulnu tnl0wllytunnn I {nu

1.2 o]rl{o{n!< deylrYD.lolrF,lnun1:n0!du0,19t0flltfi?ufllJn1:[n:6urnu In10t xr[nnnl{0.1n

< <r e i I A - a 4d , ! rr< c , , ! AyyrrJ01qu0u tnulu0[u0[nt 6u10.t?{tytuo]qu0unynoudu o{'[66 uas: ?n[5?nilniltfrotu-o^ a;<

r0:gunlnY{aT110'lq!1n

.A13 n?-ll!d ulst.uo,lli!dt

riiuHautn1fl fl ?t!dMrfi lunrrf,{[nrry#fl 1: or].t1rlufir nr:ri'lrn: ry#drn?uq!n1:rrityrn'l]nto,:fi rnrolun's narrrcrlr:o1unr:cllurruI:n rirfi iorruclr;:oiinr:noueuo,:eiodr: }i6n.j1fi ud1ilalrl:oi

24

1.4 oig2Jlor'!ol-urvnorn'r niu :rn iirsiu 1l oon Ha ruHn inaciordor6odrrirHncio

n:ru drrr:olun r: q6du dltn ?u rl u n 1: lnirU[6u In ua&fl?rud'un:n1unr:nouauorfi

rr9rfl0r1.1flu

2. UOOI'IOU?nUAltA?U8tJntflOtqrOU lOmUltrl li'

2.1 rfioto.rar:Y 6 a 4 e t d *9l

nr:lf,ar:6orrirfil6.r':-erqJ:rasritor{1d rfi o rrrnar:16u.lri'urn'orir}J1#6'uh, a - - Y

1r. fl1.:yuaflu tljfl'D.r1:o111 tlrv{ano fluo.: lnlHUo n 9nuu nou tafll:no tvl'tn1: maoot

n0urdu0

2.2 n?lutlnn uta{o1:

ar:rfi n16oariurn'orirhll#niun'vrfi n16orfi uueiinarur{l{usixriurirl#, h9.rn ou du o't r19l fl n 1nfl u to

z.s 3rJtorar:or:neuqrnr:roitg16ulntorx'rfi rirrLrldmr':nr:rnun:ig rJuernrirrriu rdu

fr i s, - -'- o or9 a a a t Ioq tu!ljund H{ dl:avalu fi:u n{uu nl: 'rfll:rJl ti6 o.r1r!1:ol1o.: ?nq :cd{n n?lrJd3n?n

iny- - :l a e a . ero e I d,Hnfr'ldir ar:rrrr nrfrornSou ar:do.rT rl{hiurn otirr::nri r iqsuiu ar: ovrfio!fl nr1r

ua s rio r 1{1u dx fi o r n 1 f, L; Eo u,o"'n

2.4 2trnrrltett c 4 9 o11 r i. oc/

n15 ttdl: nU?{SnO{nrU {6r{9u10'1dl: YrU lrJ',r ti n1: tirflr:n{! a101r lyu

rfio1{ar:fioqjlug xo{flr:nsa1u oa:lffinr:finriuar: riluiifi:rnrBr rrayldrnSoru'oh, ,i qy Aa 4 d, ,tfif,vo?n [tJ0 tsdl:Yrtlsdlul:nqori,l1l't t: ]n tn fl?: ti,?f nt::tndlra.!f,u rllxr0lu

c - id3. tJoountnuzRudRlt' r?oaon

r.r qorrgiqe ' a e '. , & ui nr,-nlr tljd'r:lrernutuon 0.:ltnt: ol tqu fl!un10{ f,l: u u dl ?ldlluutrjo lnSu

arrliourirl#rf,olqornrn'hJ1dn3o'hi rdu ar:rovrt^6u 6'11dlurixiiqililg!^q{ iorrllouoo, d <l' ,!]nnv 1 tt4d'l::yntu[:2 luu911]

3.2 rto{i .. , a as a !, llru0t 0tnlld{!r.{an 0n1: It, qlnu tstt0{yr-y 1i.:uiu0 gnusunt o.tud.!lra s

nauuTutoqrer d.rfinarionr:lflntouJrnlun'r niriunr:1#ar:urllfinnr:hilurir,rfirJrnlurflnrn:rsn'rrcinr:tir16urorrr:'hJtilri:urirq 1 to.:rir nr:biar:1urialfiilrn}Llflnoyrirq 9d h rys i , ,'hir.r-r ki:-u a r : I r n n'i lia,r fi rJ r n 1 ufl n

25

:"3.3 f,ltrnuluolntfl

fi n a ri o n r : g n du a r : t o r try a d u a udu,6 u ri r hfl a r : rn ai o u r{r { r"r n dn'r

rlerflei1{niu

s.l rir uacdrdu<, l .i'uludrYrtltnilnunludl:n'lu41ln1:rn:[u[elu tglxo{Y{n unca?u tufl1ta]tau{d'l:

'[rJtfteirusirr 1 ro{ii! luanrflu':ndorLfiirJiuronirrrnrirL?nrrrLrdliutolar:nnrlooa.r.l'a oe , v 9 t ah,c , ,io- it, t'o.:uu rufl1: rxfllletorrfl{rnnflRl u')o#olJ'irosfirlunnni o'lilludNmnra{n1r r1fllr [[os [rn?:o, o c 9d \r,ttifl 1: n{n1nfl ufi qnnfl lvl51uYl 1 nt1lrQnfl 'lr tnuou

35 ax lravoonirou

fi Hacionr:rirfi unfi rtnrlunr:6nriuf,r: tv'ir{nr:6nriuar:r1o'rqfiavrr.rau& a. o]y

ruavruunifl liluo?!n1r r flr:urifirirn::ldii1n6'rldfuna6 uovrl:snu-nar:d1*1utorvfi

oonArnuiilaeionlrlir{1u10{flr:n?uqunrrr0:iUrAulntorfirrrlrfin tdu lunr:rir.:ruei ov h, e I - i - t -iloidl:roflf nu r o tr4fla tudfr9ro.:r.rooflcrrn allulY{o u0nnlnuooncrttuu{lJnan0

A9nl:r!: qtnu tnlo.:lttf, ?t,

3.6 snlfl0firjtvtnfl

fi n a ri o g onr q fi [r fl { n ? ] ! [{u I o r u a r ri r 1 r?i niu f n' l fi rrn n ei r l riu4aada

fl?1lJfl1U1:0 [Un1:fl{tnll& rln&n1: Handl:n?uqunl:[nt€urn1l tnnlu tulcraat{[[nfln]{nulu

rRu.mRlr0ntcouqoml r.{1v

drfnr rur rer:g rufiurirrnun:unsorr4 r: [[r.i.!rrn' n3o rnor.l2 (oou'lnf. 2549)

'ldrirrunturoto{Hor{ryrrnd'lu?uilaeiofi]nn:-lrfiohirilulrnrslulunrrdotrru

Rrulurj:vlyrfl uascir{ :vr qrrnnr:1.: I n-{d

9ogntl{ I 10nl ufi[:0t1u lFltJo':r.rlcHnrnu?

t lt dlru 1fi riru:unarioilanil

I

2

3

4

uoun?l 26

26-29

30-33

34-38

d-,,Ylll"1 : !19:! lUf,Un'llnun: ttns0'1fi1:Ufi{ttn. 2549

26

,Lt. nrrud{ffuqornrnrni{tfl u 3 i'uqornrr,rL^

1.1 IlllllflU (Extra Class)

A I iq, < ,<i v ? h'd:vnou6?ur{ls tu:1.rftIu'rnm I suurio.:1lq R lfln Enn:t9tlljY{uI a ttJ!

. ao -i-. a, cr o aa d ,9t1l.tU lUn: vllJft1l.tu910{ruusr1fi ufl ?[anu0u

1.2 tiufit{tt (Class I)A '- ly a

:gn0un2ultls tu:ltdlulnYl 2 [.tlssuunD{!f]olR1 n n:{n1rlv{uI fl0lJ

eirnfildrf,nrioudru:rim:rlnuhiiHnn:vyrueio: a-nunivr'a'[rJtou6nr"ra Inodun'rfin'rrfi! )- !, L!",

:'lun0fl0 t!lnu 5% 10'tyi1]r ?11{ulJft yNu tlJt ?tJo i9t1fiulJo tl,ut.t1v

1.3 frudo.r (Class II)

d:vno!drunrclur#dru16fi : mrv#ud:rrlrlrsdlr;,,flltir#udqon'jr urifi

narnrnriun:r6o nofieirrfildrf;nr?oudru:rlvr:rlntrhiinan:vmurio::Jn-nuoin-r'hjtor

Hnfl A tnUY{Ur'l?ll911l4U:llJOOfln t!tfllJ lOplo uO{Y{UYl}l?11{fiUO 1l{U tUt?UO.:9l1l4U1O.lIU['tlv

nlndo{aro.!nrunr:r{rnrululd}ido{anr: fildor':reilh{1: rG oueruqorn rn

rua s n'r 1s : rfl rfl a r n l n tr o r n r?o ry

a 1u 16 ou !- q u r uu ij zs s u : rn rirdol o.r r r rv #u fi ras il3 0

"-^,-[n:n A 35-40 uryr/nn. [{rs#u luSorn:a B 30-40 u'l /nn. rrrvd'u 2 nSorn:o c l5-20 u'lyynn.

uos[.:1vn nslyrno{ l5-20 urfl/nn. (dru-nr ru nr: flrn rulu o-rx e-n:v uo.r. oou'lnf. 2557) dr

ofirrn' #asorn uavri# rirl:.rrdu (2556:78) Lil#r?orya'jr laorir'lrJudaurv]:rfiuuhinT :fi

druauHariu 28-30 fla/nn. uay lnHar!'lvi,r,uredrn'jrfi uro:gruril un osrirl#:rorfrdo- | ?.t t iny !,! t e J , - t ' A

nnn{ 50 [:lo5lcruntJo.:1,u1oYr tn1l1fl:r]uyl{un?'r1r[[nflrl1.:fi'lu:1n11u0qna]ntury0rlnayyl