1

Literature Cited

Ådahl, E., P. Lundberg, and N. Jonzén. 2006. From climate change to population change: the

need to consider annual life cycles. Global Change Biology 12:1627-1633.

Agee, J.K. 1993. Fire Ecology of Pacific Northwest Forests. Island Press, Washington, D.C.

Ager, A.A., M.A. Finney, B.K. Kerns and H. Maffei. 2007. Modeling wildfire risk to northern

spotted owl (Strix occidentalis caurina) habitat in Central Oregon, USA. Forest Ecology and

Management 246:45–56.

Ager, A.A., N.M Vaillant, M.A. Finney, and H.K. Priesler. 2012. Analyzing wildfire exposure

and source-sink relationships on a fire-prone forest landscape. Forest Ecology and

Management 267: 271-283.

Allen, C.D., M. Savage, D.A. Kalk, K.F. Suckling, T.W. Swetnam, T. Schulke, P.B. Stacey, P.

Morgan, M. Hoffman, and J.T. Klingel. 2002. Ecological restoration of southwestern

ponderosa pine ecosystems: a broad perspective. Ecological Applications 12(5): 1418–1433.

Anderson, D.R., and K.P. Burnham. 1992. Demographic analysis of northern spotted owl

populations. Pages 319–328 in Draft final recovery plan for the northern spotted owl. U.S.

Fish and Wildlife Service, Portland, Oregon.

Anthony, R.G., and F.F. Wagner. 1998. Reanalysis of northern spotted owl habitat use on the

Miller Mountain Study Area. Unpublished report. Submitted to Bureau of Land

Management, Medford District and Forest and Rangeland Ecosystem Science Center,

Biological Resources Division, U.S. Geological Survey.

Anthony, R.G., E.D. Forsman, A.B. Franklin, D.R. Anderson, K.P. Burnham, G.C. White, C.J.

Schwarz, J. Nichols, J.E. Hines, G.S. Olson, S.H. Ackers, S. Andrews, B.L. Biswell, P.C.

Carlson, L.V. Diller, K.M. Dugger, K.E. Fehring, T.L. Fleming, R.P. Gerhardt, S.A. Gremel,

R.J. Gutiérrez, P.J. Happe, D.R. Herter, J.M. Higley, R.B. Horn, L.L. Irwin, P.J. Loschl, J.A.

Reid and S.G. Sovern. 2006. Status and trends in demography of northern spotted owls,

1985–2003. Wildlife Monograph No. 163.

Aubry, K.B., C.B. Halpern, and C.E. Peterson. 2009. Variable-retention harvests in the Pacific

Northwest: A review of short-term findings from the DEMO study. Forest Ecology and

Management 258:398-408.

Baker, S. 2011. Seeking a balance between forestry and biodiversity the role of variable

retention silviculture. Forest & Wood Products Australia Limited PDG167-0910.

Baker, W. L. 2012. Implications of spatially extensive historical data from surveys for restoring

dry forests of Oregon’s eastern Cascades. Ecosphere 3(3):23.

2

Barrowclough, G.F., and R. J. Gutiérrez. 1990. Genetic variation and differentiation in the

spotted owl. Auk 107:737–744.

Barrowclough, G.F., R.J. Gutiérrez and J.G. Groth. 1999. Phylogeography of spotted owl (Strix

occidentalis) populations based on mitochondrial DNA sequences: gene flow, genetic

structure, and a novel biogeographic pattern. Evolution 53:919–931.

Barrowclough, G.F., J.G. Groth and R.J. Gutiérrez. 2005. Genetic structure, introgression and a

narrow hybrid zone between northern and California spotted owls (Strix occidentalis).

Molecular Ecology 14:1109–1120.

Barrows, C. and K. Barrows. 1978. Roost characteristics and behavioral thermoregulation in the

spotted owl. Western Birds 9(1): 1-9.

Bean, M.J. 2002. Overcoming unintended consequences of endangered species regulation.

Idaho Law Review. 38:409-420.

Betts, M.G., J.C. Hagar, J.W. Rivers, J.D. Alexander, K. McGarigal, and B.C. McComb. 2010.

Thresholds in forest bird occurrence as a function of the amount of early-seral broadleaf

forest at landscape scales. Ecological Application 20(8):2116-2130.

Bingham, B.B. and B.R. Noon. 1997. Mitigation of Habitat ''Take": Application to Habitat

Conservation Planning. Conservation Biology 11(1): 127-139.

Blakesley, J.A. 2004. Habitat Associations. Chapter 5 in S.P. Courtney, J.A. Blakesley, R.E.

Bigley, M.L. Cody, J.P. Dumbacher, R.C. Fleischer, A.B. Franklin, J.F. Franklin, R.J.

Gutiérrez, J.M. Marzluff and L. Sztukowski. Scientific evaluation of the status of the

northern spotted owl. Sustainable Ecosystems Institute, Portland, Oregon.

Blakesley, J.A., A.B. Franklin and R.J. Gutiérrez. 1992. Spotted owl roost and nest site

selection in northwestern California. Journal of Wildlife Management 56(2):388-392.

Blate, G.M., L.A. Joyce, J.S. Littell, S.G. McNulty, C.I. Millar, S.C. Moser, R.P. Neilson, K.

O’Halloran and D.L. Peterson. 2009. Adapting to climate change in United States national

forests. Unasylva 231/232. Vol. 60:57–62.

Bockstael, N. E., A. M. Freeman, III, R. J. Kopp, P. R. Portney, and, V. K. Smith. 2000. On

measuring economic values for nature. Environmental Science & Technology 34(8):1384-

1389.

Bond, M.L., R.J. Gutiérrez, A.B. Franklin, W.S. LaHaye, C.A. May and M.E. Seamans. 2002.

Short-term effects of wildfires on spotted owl survival, site fidelity, mate fidelity, and

reproductive success. Wildlife Society Bulletin 30:1022–1028.

3

Boyd, C., T.M. Brooks, S.H.M. Butchart, G.J. Edgar, G.A.B. da Fonseca, F. Hawkins, M.

Hoffmann, W. Sechrest, S.N. Stuart, and P.P. van Dijk. 2008. Spatial scale and the

conservation of threatened species. Conservation Letters 1:37-43.

Brook, A., M. Zint, and R. De Young. 2003. Landowner’s responses to an endangered species

act listing and implications for encouraging conservation. Conservation Biology. 17:1638-

1649.

Buchanan, J.B. 2004. Managing habitat for dispersing northern spotted owls – are the current

management strategies adequate? Wildlife Society Bulletin 32:1333–1345.

Buchanan, J.B., L.L. Irwin, and E.L. McCutchen. 1993. Characteristics of spotted owl nest trees

in the Wenatchee National Forest. Journal of Raptor Research. 27:1-7.

Buchanan, J.B. 2009. Balancing competing habitat management needs for northern spotted owls

and other bird species in dry forest landscapes. Pages 109–117 in: T.D. Rich, C. Arizmendi,

D. Demarest, and C. Thompson (editors), Proceedings of the Fourth International Partners in

Flight Conference: Tundra to Tropics. McAllen, Texas.

Burnham, K.P., D.R. Anderson and G.C. White. 1994. Estimation of vital rates of the northern

spotted owl. Colorado Cooperative Fish and Wildlife Research Unit, Colorado State

University, Ft. Collins, Colorado.

Camp, A.E., C.D. Oliver, P.F. Hessburg and R.L. Everett. 1997. Predicting late-successional fire

refugia from physiography and topography. Forest Ecology and Management 95:63–77.

Carey, A.B. 1985. A summary of the scientific basis for spotted owl management. Pp. 100-114

in R.J. Gutierrez and A.B. Carey, (technical editors), Ecology and management of the spotted

owl in the Pacific Northwest. U.S.D.A. Forest Service General Technical Repport PNW-

185. Pacific Northwest Forest and Range Experiment Station, Portland, Oregon.

Carey, A.B. 1995. Sciurids in Pacific Northwest managed and old-growth forests. Ecological

Applications 5(3):648–661.

Carey, A.B. 2000. Effects of new forest management strategies on squirrel populations.

Ecological Applications 10(1):248–257.

Carey, A.B. 2003. Bioclomplexity and restoration of biodiversity in temperate coniferous forest:

inducing spatial heterogeneity with variable-density thinning. Forestry 76:127–136.

Carey, A.B. 2006. Active and passive forest management for multiple values. Northwest

Naturalist 87(1):18-30.

Carey, A.B., and K.C. Peeler. 1995. Spotted owls: resource and space use in mosaic

landscapes. The Journal of Raptor Research. 29:223-239.

4

Carey, A.B., S.P. Horton and B.L. Biswell. 1992. Northern spotted owls: influence of prey base

and landscape character. Ecological Monographs 62(2):223–250.

Carey, A.B., C.C. Maguire, B.L. Biswell and T.M. Wilson. 1999. Distribution and abundance of

Neotoma in western Oregon and Washington. Northwest Science 73(2):65–80.

Carroll, C. 2010. Role of climatic niche models in focal-species-based conservation planning:

assessing potential effects of climate change on northern spotted owl in the Pacific

Northwest, USA. Biological Conservation. 143:1432-1437.

Carroll, C., D.C. Odion, C.A. Frissell, D.A. Dellasala, B.R. Noon, and R. Noss. 2009.

Conservation implications of coarse-scale versus fine-scale management of forest

ecosystems. Klamath Center for Conservation Research, Orleans, CA. 39 pgs. Available at

www.klamathconservation.org.

Carroll, C., J.R. Dunk and A. Moilanen. 2010. Optimizing resiliency of reserve networks to

climate change: multispecies conservation planning in the Pacific Northwest. Global Change

Biology 16:891-904.

Case, M.J., and D.L. Peterson. 2007. Growth-climate relations of lodgepole pine in the North

Cascades National Park, Washington. Northwest Science 81:62–74.

Chmura, D.J., P.D. Anderson, G.T. Howe, C.A. Harrington, J.E. Halofsky, D.L. Peterson, D.C.

Shaw,and J.B. St. Clair. 2011. Forest response to climate change in the northwestern United

States: ecophysical foundations for adaptive management. Forest Ecology and Management

261: 1121-1142.

Chow, N. 2001. Distribution and habitat associations of northern spotted owls in Marin County,

California. Master’s thesis, Humboldt State University. 56 pages.

Chutter, M.J., I. Blackburn, D. Bonin, J. Buchanan, B. Costanzo, D. Cunnington, A. Harestad, T.

Hayes, D. Heppner, L. Kiss, J. Surgenor, W. Wall, L. Waterhouse and L. Williams. 2004.

Recovery strategy for the northern spotted owl (Strix occidentalis caurina) in British

Columbia. British Columbia Ministry of Environment, Victoria, Canada.

Clark, D.A. 2007. Demographic and habitat selection of northern spotted owls in post-fire

landscapes of southwestern Oregon. Thesis, Oregon State University, Corvallis, Oregon.

Conley, J.L., M.E. Fernandez-Gimenez, G.B. Ruyle, and M. Brunson. 2007. Forest Service

grazing permittee perceptions of the Endangered Species Act in southeastern Arizona.

Rangeland Ecology and Management. 60:136-145.

Collins, L. R.A. Bradstock, E.M. Tasker, and R.J. Whelan. 2012. Impacts of fire regimes,

logging, and topography on hollows in fallen logs in eucalypt forest of south eastern

Australia. Biological Conservation 149: 23-31.

5

Costanza, R., R. d'Arge, R. de Groot, S. Farberk, M. Grasso, B. Hannon, K. Limburg, S. Naeem,

R.V. O'Neill, J. Paruelo, R. G. Raskin, P. Sutton,and M. van den Belt. 1997. The value of the

world's ecosystem services and natural capital. Nature 387: 253-260.

Courtney, S.P., J.A. Blakesley, R.E. Bigley, M.L. Cody, J.P. Dumbacher, R.C. Fleischer, A.B.

Franklin, J.F. Franklin, R.J. Gutiérrez, J.M. Marzluff and L. Sztukowski. 2004. Scientific

evaluation of the status of the northern spotted owl. Sustainable Ecosystems Institute,

Portland, Oregon.

Crouse, D.T., L.A. Mehrhoff, M.J. Parkin, D.R. Elam, and L. Y. Chen. 2002. Endangered

species recovery and the SCB study: a U.S. Fish and Wildlife Service perspective.

Ecological Applications. 12:719-723.

Crozier, M.L., M.E. Seamans, R.J. Gutiérrez, P.J. Loschl, R.B. Horn, S.G. Sovern and E.D.

Forsman. 2006. Does the presence of barred owls suppress the calling behavior of spotted

owls? Condor 108: 760–769.

Davis, L. R.Haugo, and D. Kavanagh. 2012. An ecological context for ―whole system‖

conservation of eastern Washington forests. The Nature Conservancy. Seattle, Washington,

USA.

Davis, R.J., and K.M. Dugger. 2011. Chapter 3: Habitat status and trend. Pp. 21-62 in Davis,

R.J., K.M. Dugger, S. Mohoric, L. Evers, and W.C. Aney. Northwest Forest Plan—the first

15 years (1994-2008): status and trends of northern spotted owl populations and habitats.

U.S.D.A. Forest Service General Technical Report PNW-GTR-850. Pacific Northwest

Research Station, Portland, Oregon.

Davis, R., and J. Lint. 2005. Habitat status and trends. Pages 21–82 in J. Lint, technical

coordinator. Northwest Forest Plan—the first 10 years (1994–2003): status and trends of

northern spotted owl populations and habitat. General Technical Report PNW-GTR-648,

Pacific Northwest Research Station, U.S. Department of Agriculture, Forest Service,

Portland, Oregon.

Dodson, E.K., A.Ares, and K.J. Puettmann. 2012. Early responses to thinning treatments

designed to accelerate late successional forest structure in young coniferous forests of

western Oregon, USA. Canadian Journal of Forest Research 42:345-355.

Drever, C.R., G. Peterson, C. Messier, Y. Bergeron and M. Flannigan. 2006. Can forest

management based on natural disturbances maintain ecological resilience? Canadian Journal

of Forest Research 36:2285–2299.

Dugger, K.M., F. Wagner, R.G. Anthony and G.S. Olson. 2005. The relationship between habitat

characteristics and demographic performance of northern spotted owls in southern Oregon.

Condor 107:863–878.

6

Dugger, K.M., R.G. Anthony and L.S. Andrews. 2011. Transient dynamics of invasive

competition: barred owls, spotted owls, habitat, and the demons of competition present.

Ecological Applications. 21:2459-2468.

Dunk, J. R., B. Woodbridge, D. LaPlante, N. Schumaker, B. Glenn, B.C. White, K. Halupka, S.

Livingston, M. M. Zwartjes and J. Peters. 2012a (March 6, 2012). Modeling and analysis

procedures used to identify and evaluate potential critical habitat networks for the northern

spotted owl. Unpublished report, U.S. Fish and Wildlife Service, Portland, Oregon. 48 pp.

Dunk, J. R., B. Woodbridge, D. LaPlante, N. Schumaker, B. Glenn, B.C. White, K. Halupka, S.

Livingston, M. M. Zwartjes and J. Peters. 2012b (November 21, 2012). Modeling and

analysis procedures used to identify and evaluate potential critical habitat networks for the

northern spotted owl. Unpublished report, U.S. Fish and Wildlife Service, Portland, Oregon.

85 pp.

Elith, J., C.H. Graham, R.P. Anderson, M. Dudik, S. Ferrier, A. Guisan, R.J. Hijmans, F.

Huettmann, J.R. Leathwick, A. Lehmann, J. Li, L.G. Lohmann, B.A. Loiselle, G. Manion, C.

Moritz, M. Nakamura, Y. Nakazawa, J. McC. Overton, A. Townsend Peterson, S.J. Phillips,

K. Richardson, R. Scachetti-Pereira, R.E. Schapire, J. Sobero´n, S. Williams, M. S. Wisz and

N.E. Zimmermann. 2006. Novel methods improve prediction of species’ distributions from

occurrence data. Ecography 29(2):129-151.

Ellis, T.M., and M.G. Betts. 2011. Bird abundance and diversity across a hardwood gradient

within early seral plantation forest. Forest Ecology and Management. 261:1372-1381.

Everett, R.L., R. Schellhaas, D. Keenum, D. Spurbeck and P. Ohlson. 2000. Fire history in the

ponderosa pine/Douglas-fir forests on the east slope of the Washington Cascades. Forest

Ecology and Management 129:207–225.

Fiedler, C. E., P. Friederici, M. Petruncio, C. Denton, and W. D. Hacker. 2007. Managing for

old growth in frequent-fire landscapes. Ecology and Society 12(2):20.

Folliard, L.B., K. Reese, and L.V. Diller. 2000. Landscape characteristics of Northern Spotted

Owl nest sites in northwestern California. Journal of Raptor Research 34: 75–84.

Fontaine, J.B. and P.L. Kennedy. 2012. Avian and small mammal response to fire severity and

fire surrogate treatments in US fire-prone forests: a meta-analysis. Ecological Applications:

22: 1547–1561.

Forest Management Strategy. 2005. A Forest Management Strategy for the Fort Lewis Military

Reservation, Washington 2005. Forestry Program, Environmental and Natural Resources

Division. U.S. Army, Installation Management Agency, Fort Lewis Military Reservation.

Revised May 2005. 95 pp.

7

Forsman, E.D., E.C. Meslow and H.M. Wight. 1984. Distribution and biology of the spotted owl

in Oregon. Wildlife Monographs 87:1–64.

Forsman, E.D., I.A. Otto, S.G. Sovern, M. Taylor, D.W. Hays, H. Allen, S.L. Roberts and D.E.

Seaman. 2001. Spatial and temporal variation in diets of spotted owls in Washington. Journal

of Raptor Research 35:141–150.

Forsman, E.D., R.G. Anthony, J.A. Reid, P.J. Loschl, S.G. Sovern, M. Taylor, B.L. Biswell, A.

Ellingson, E.C. Meslow, G.S. Miller, K.A. Swindle, J.A. Thrailkill, F.F. Wagner and D.E.

Seaman. 2002. Natal and breeding dispersal of northern spotted owls. Wildlife Monographs

149:1–35.

Forsman, E.D., R.G. Anthony, E.C. Meslow and C.J. Zabel. 2004. Diets and foraging behavior

of northern spotted owls in Oregon. Journal of Raptor Research 38:214–230.

Forsman, E.D, T.J. Kaminski, J.C. Lewis, K.J. Maurice, C. Ferland and E. M. Glenn. 2005.

Home range and habitat use of northern spotted owls on the Olympic Peninsula, Washington.

Journal of Raptor Research 39(4):356-377.

Forsman, E.D., R.G. Anthony, K.M. Dugger, E.M. Glenn, A.B. Franklin, G.C. White, C.J.

Schwarz, K.P. Burnham, D.R. Anderson, J.D. Nichols, J.E. Hines, J.B. Lint, R.J. Davis, S.H.

Ackers, L.S. Andrews, B.L. Biswell, P.C. Carlson, L.V. Diller, S.A. Gremel, D.R. Herter,

J.M. Higley, R.B. Horn, J.A. Reid, J. Rockweit, J. Schaberl, T.J. Snetsinger and S.G. Sovern.

2011. Population demography of northern spotted owls: 1985–2008. Studies in Avian

Biology. No. 40. Cooper Ornithological Society.

Frankin, A.B. and R.J. Gutiérrez. 2002. Spotted owls, forest fragmentation, and forest

heterogeneity. Studies in Avian Biology 25: 203-220.

Franklin, A.B., K. P. Burnham, G.C. White, R.G. Anthony, E.D. Forsman, C. Schwarz, J.D.

Nichols and J. Hines. 1999. Range-wide status and trends in northern spotted owl

populations. Colorado Cooperative Fish and Wildlife Research Unit, Colorado State

University, Fort Collins, Colorado.

Franklin, A.B., D.R. Anderson, R.J. Gutiérrez and K.P. Burnham. 2000. Climate, habitat quality,

and fitness in northern spotted owl populations in northwestern California. Ecological

Monographs 70:539–590.

Franklin, J.F., and C.T. Dyrness. 1988. Natural vegetation of Oregon and Washington. Oregon

State University Press, Corvallis, OR. 452 pp.

Franklin, J. F. and D. B. Lindenmayer. 2009. Importance of matrix habitats in maintaining

biological diversity. Proceedings of the National Academy of Sciences 106(2):359-350.

Franklin, J.F., T.A. Spies, R. Van Pelt, A.B. Carey, D.A. Thornburgh, D.R. Berg, D.B.

Lindenmayer, M.E. Harmon, W.S. Keeton, D.C. Shaw, K. Bible and J. Chen. 2002.

8

Disturbances and structural development of natural forest ecosystems with silvicultural

implications, using Douglas-fir forests as an example. Forest Ecology and Management

155:399–423.

Franklin, J.F., R.J. Mitchell, and B.J. Palik. 2007. Natural disturbance and stand development

principles for ecological forestry. General Technical Report NRS-19. U.S.D.A. Forest

Service, Northern Research Station, Newtown Square, Pennsylvania.

Franklin, J.F., M.A. Hemstrom, R. Van Pelt, J.B. Buchanan, and S. Hull. 2008. Types in eastern

Washington: perpetuating and creating old forest structures and functions. Washington State

Department of Natural Resources, Lacey, WA. 105pp.

Fulé, P.Z., J E. Crouse, J.P. Roccaforte, E. L. Kalies. 2012. Do thinning and/or burning

treatments in western USA ponderosa or Jeffrey pine-dominated forests help restore natural

fire behavior?, Forest Ecology and Management 269: 68-81.

Gaines, W.L., R.J. Harrod, J. Dickinson, A.L. Lyons and K. Halupka. 2010. Integration of

Northern spotted owl habitat and fuels treatments in the eastern Cascades, Washington, USA.

Forest Ecology and Management 260:2045–2052.

Ganey, J.L., R.P. Balda and R.M. King. 1993. Metabolic rate and evaporative water loss of

Mexican spotted and great horned owls. Wilson Bulletin 105(4):645-656.

Glenn, E.M. 2009. Local weather, regional climate, and population dynamics of northern

spotted owls in Washington and Oregon. Dissertation. Oregon State University, Corvallis

OR, USA. 299 pp.

Glenn, E.M., M. C. Hansen, and R. G. Anthony. 2004. Spotted owl home range and habitat use

in young forests of western Oregon. Journal of Wildlife Management 68::33-50.

Glenn, E.M., R.G. Anthony, and E.D. Forsman. 2010. Population trends in northern spotted

owls: associations with climate in the Pacific Northwest. Biological Conservation 143: 2543-

2552.

Glenn, E.M., R.G. Anthony, E.D. Forsman, and G.S. Olson. 2011a. Local weather, regional

climate, and annual survival of the northern spotted owl. Condor 113:159-176.

Glenn, E.M., R.G. Anthony, E.D. Forsman, and G.S. Olson. 2011b. Reproduction of northern

spotted owls: the role of local weather and regional climate. Journal of Wildlife Management

75(6):1279-1294.

Gomez, D.M., Anthony, R.G. and Hayes, J.P. 2005. Influence of thinning of Douglas-fir forests

on population parameters and diet of northern flying squirrels: Journal of Wildlife

Management 69:1670–1682.

9

Graham, C.H., J. Elith, R. Hijmans, A. Guisan, A.T. Peterson, B.A. Loiselle and the NCEAS

species Distribution Modeling Group. 2008. The influence of spatial errors in species

occurrence data used in distribution models. Journal of Applied Ecology, 45: 239-247.

Gremel, S. 2005. Factors controlling distribution and demography of northern spotted owls in a

reserved landscape. Thesis, University of Washington, Seattle.

Groves, C.R., L.S. Kutner, D.M. Stoms, M.P. Murray, J.M. Scott, M. Schafale, A.S. Weakley,

and R.L. Pressey. 2000. Owning up to our responsibilities—Who owns lands important

for biodiversity? Pages 275–300 in B.A. Stein, L.S. Kunter, and J.S. Adams, editors,

Precious heritage: The status of biodiversity in the United States. Oxford University

Press, New York, New York. 399 pp.

Gustafsson, L., S.C. Baker, J. Bauhus, W.J. Beese, A. Brodie, J. Kouki, D.B. Lindenmayer, A.

Lõhmus, G. Martínez Pastur, C. Messier, M.Neyland, B. Palik, A. Sverdrup-Thygeson, W. J.

A. Volney, A.Wayne, and..J. F. F ranklin. 2012. Retention forestry to maintain

multifunctional forests: a world perspective. BioScience 62: 633–645.

Gutiérrez, R.J., A.B. Franklin and W.S. LaHaye. 1995. Spotted owl (Strix occidentalis) in A.

Poole and F. Gill (editors), The birds of North America, No. 179. The Academy of Natural

Sciences and The American Ornithologists’ Union, Washington, D.C.

Hagar, J.C. 2007. Wildlife species associated with non-coniferous vegetation in Pacific

Northwest conifer forests: A review. Forest Ecology and Management 246:108-122.

Haig, S.M., T.D. Mullins, E.D. Forsman, P.W. Trail, and L. Wennerberg. 2004. Genetic

identification of spotted owls, barred owls, and their hybrids: legal implications of hybrid

identity. Conservation Biology 18:1347-1357

Halofsky, J.E., D. C. Donato, D. E. Hibbs, J. L. Campbell, M. Donaghy Cannon, J. B. Fontaine,

J. R. Thompson, R. G. Anthony, B. T. Bormann, L. J. Kayes, B. E. Law, D. L. Peterson, and

T. A. Spies. 2011. Mixed-severity fire regimes: lessons and hypotheses from the Klamath-

Siskiyou Ecoregion. Ecosphere 2(4):40.

Hamm, K.A., and L.V. Diller. 2009. Forest management effects on abundance of woodrats in

northern California. Northwestern Naturalist. 90:97-106.

Hanson, C.T., D.C. Odion, D.A. DellaSala and W.L. Baker. 2009. Overestimation of fire risk in

the Northern Spotted Owl Recovery Plan. Conservation Biology 23:1314-1319.

Healey, S.P., W.B. Cohen, T.A. Spies, M. Moeur, D. Pflugmacher, M.G. Whitleyand M. Lefsky.

2008. The relative impact of harvest and fire upon landscapelevel dynamics of older forests:

lessons from the Northwest Forest Plan. Ecosystems 11: 1106-1119.

10

Hershey, K.T., E.C. Meslow and F.L. Ramsey. 1998. Characterisics of forests at spotted owl

nest sites in the Pacific Northwest. Journal Wildlife Management 62(4):1398-1410.

Herter, D.R., L.L. Hicks, H.C. Stabins, J.J. Millspaugh, A.J. Stabins, and L.D. Melampy. 2002.

Roost site characteristics of northern spotted owls in the nonbreeding season in central

Washington. Forest Science. 48:437-444.

Herter, D.R., L.L. Hicks, H.C. Stabins, J.J. Millspaugh, A.J. Stabins and L.D. Melampy. 2002.

Roost site characteristics of northern spotted owls in the nonbreeding season in central

Washington. Forest Science 48:437–446.

Hessburg, P.F., and J.K. Agee. 2003. An environmental narrative of Inland Northwest United

States forests, 1800–2000. Forest Ecology and Management 178:23–59.

Hessburg, P.F., B.G. Smith, R.B. Salter, R.D. Ottmar and E. Alvarado. 2000a. Recent changes

(1930s–1990s) in spatial patterns of interior northwest forests, USA. Forest Ecology and

Management 136:53–83.

Hessburg, P.F., R.B. Salter, M.B. Richmond, and B.G. Smith. 2000b. Ecological subregions of

the Interior Columbia Basin, USA. Applied Vegetation Science. 3:163-180.

Hessburg, P.F., J.K. Agee and J.F. Franklin. 2005. Dry forests and wildland fires of the inland

Northwest USA: Contrasting the landscape ecology of the pre-settlement and modern eras.

Forest Ecology and Management 211:117–139.

Hessburg, P.F., R.B. Salter and K.M. James. 2007. Re-examining fire severity relations in pre-

management era mixed conifer forests: inferences from landscape patterns of forest structure.

Landscape Ecology 22:5–24.

Heyerdahl, E.K., D. McKenzie, L.D. Daniels, A.E. Hessl, J.S. Littell and N.J. Mantua. 2008.

Climate drivers of regionally synchronous fires in the inland Northwest (1651-1900).

International Journal of Wildland Fire 17:40-49.

Hicks, L.L., D.R. Herter, and R. Early. 2003. Clines in life history characteristics of the spotted

owl in Washington. Northwestern Naturalist 84:57-67.

Hobbs, R.J., D.N. Cole, L. Yung, E.S. Zavaleta, G.H Aplet, F.S. Chapin III, P.B. Landres, D.J.

Parsons, N.L. Stephensen, P.S. White, D.M. Graber, E.S. Higgs, C.I. Millar, J.M. Randall,

K.A. Tonnessen, and S. Woodley. 2010. Guiding concepts for Park and wilderness

stewardship in an era of global environmental change. Frontiers of Ecology and the

Environment 8(9):483–490.

Holloway, G.L., and W.P. Smith. 2011. A meta-analysis of forest age and structure effects on

northern flying squirrel densities. The Journal of Wildlife Management. 75:668-674.

11

Hunter, J.E., R.J. Gutiérrez and A.B. Franklin. 1995. Habitat configuration around spotted owl

nest sites in northwestern California. Condor 97:684-693.

Industrial Economic, Inc. (IEc), ―Economic Analysis of Critical Habitat Designation for the

Northern Spotted Owl, Final Report,‖ prepared for the U.S. Fish and Wildlife Service,

November XX, 2012.

INRMP, 2007. Final Integrated Natural Resources Management Plan, 2007-2011. Department

of Defense (Army). Fort Lewis, Washington. June 2007.

Irwin, L.L., D.F. Rock and G.P. Miller. 2000. Stand structures used by northern spotted owls in

managed forests. Journal of Raptor Research 34:175–186.

Irwin, L. L., T. L. Fleming, and J. Beebe. 2004. Are spotted owl populations sustainable in fire-

prone forests? Journal of Sustainable Forestry 18:1–28.

Irwin, L.L., L.A. Clark, D.C. Rock, and S.L. Rock. 2007a. Modeling foraging habitat of

California spotted owls. The Journal of Wildlife Management. 71:1183-1191.

Irwin, L.L., M. Stephens, D. Rock, and S. Rock. 2007b. Spotted owl habitat selection and home

range in California coastal forests: draft summary report. National Council for Air and

Stream Improvement. Corvallis, OR.

Irwin L. L., D.F. Rock and S. C. Rock. 2012. Habitat selection by northern spotted owls in

mixed-coniferous forests. Journal of Wildlife Management. 76:200-213.

James, S.M. 2002. Bridging the gap between private landowners and conservationists.

Conservation Biology. 16:269-271.

Johnson, K.N., and J.F. Franklin. 2009. Restoration of Federal forests in the Pacific Northwest:

strategies and management implications. Unpublished manuscript. August 15, 2009. 120 pp.

Available online at

<http://www.cof.orst.edu/cof/fs/PDFs/RestorationOfFederalForestsInThePacificNorthwest.p

df>. Accessed 3 March 2011.

Johnson, K.N., and J.F. Franklin. 2012. Southwest Oregon secretarial pilot projects on BLM

lands: our experience so far and broader considerations for long-term plans. Februray 15,

2012. Available on line at http://www.blm.gov/or/news/files/pilot-report-feb2012.pdf .

Kane, V.R., R.F. Gersonde, J.A. Lutz, R.J. McGaughey, J.D. Bakker, and J.F. Franklin. 2011.

Patch dynamics and the development of structural and spatial heterogeneity in Pacific

Northwest forests. Canadian Journal of Forest Research 41: 2276-2291.

Karl, T.R., J.M. Melillo, and T.C. Peterson. 2009. Global Climate Change Impacts in the

United States. Cambridge University Press. Cambridge, UK.

12

Kelly, E.G., E.D. Forsman and R.G. Anthony. 2003. Are barred owls displacing spotted owls?

Condor 105:45–53.

Kennedy, R.S.H., and M.C. Wimberly. 2009. Historical fire and vegetation dynamics in dry

forests of the interior Pacific Northwest, USA, and relationships to northern spotted owl

(Strix occidentalis caurina) habitat conservation. Forest Ecology and Management.

258:554-566.

Kerr, A. 2012. Ecologically appropriate thinning in the Northwest Forest Plan area: a policy

and technical analysis. Conservation, Northwest, GEOS Institute, Klamath-Siskiyou

Wildlands Center, and Oregon Wild. 91 pp.

King, G. 1993. Habitat Characteristics of Northern Spotted Owls in Eastern Washington.

Master’s thesis; Univ. of California, Berkeley. 81 pp.

Knight, R.L. 1998. Ecosystem management and conservation biology. Landscape and Urban

Planning 40:41-45.

Krebs, C.J. 1972. Ecology. Harper and Row. New York, NY. 694 pp.

Kuchler, A.W. 1977. The Map of the Natural Vegetation of California. University of Kansas,

Lawrence.

Kuuluvainen, T. and R. Grenfell 2012. Natural disturbance emulation in boreal forest

ecosystem management – theories, strategies, and a comparision with conventional even-

aged management. Canadian Journal of Forest Research 42: 1185-1203.

LaHaye, W.S., and R.J. Gutiérrez. 1999. Nest sites and nesting habitat of the northern spotted

owl in northwestern California. Condor 101:324-330.

LaHaye, W.S., R.J. Gutiérrez and J.R. Dunk. 2001. Natal dispersal of the spotted owl in southern

California: Dispersal profile of an insular population. Condor 103:691–700.

Lamberson, R.H., R. McKelvey, B.R. Noon and C. Voss. 1992. A Dynamic Analysis of

Northern Spotted Owl Viability in a Fragmented Forest Landscape. Conservation Biology

6(4):505-512.

Lamberson, R.H., B.R. Noon, C. Voss, and K.S. McKelvey. 1994. Reserve design for territorial

species: the effects of patch size and spacing on the viability of the northern spotted owl.

Conservation Biology. 8:185-195.

Latta, G., H. Temesgen, D. Adams and T. Barrett. 2010. Analysis of potential impacts of climate

change on forests of the United States Pacific Northwest. Forest Ecology and Management

259: 720–729.

13

Lehmkuhl, J.F., L.E. Gould, E. Cazares and D.R. Hosford. 2004. Truffle abundance and

mycophagy by northern flying squirrels in eastern Washington forests. Forest Ecology and

Management 200:49–65.

Lehmkuhl, J.F., K.D. Kistler, J.S. Begley and J. Boulanger. 2006. Demography of northern

flying squirrels informs ecosystem management of western interior forests. Ecological

Applications 16:584–600.

Lindenmayer, D., R.J. Hobbs, R. Montague-Drake, J. Alexandra, A. Bennett, M. Burgman, P.

Cale, A. Calhoun, V. Cramer, P. Cullen, D. Driscoll, L. Fahrig, J. Fischer, J. Franklin, Y.

Haila, M. Hunter, P. Gibbons, S. Lake, G. Luck, C. MacGregor, S. McIntyre, R.M. Nally, A.

Manning, J. Miller, H. Mooney, R. Noss, H. Possingham, D. Saunders, F. Schmiegelow, M.

Scott, D. Simberloff, T. Sisk, G. Tabor, B. Walker, J. Wiens, J. Woinarski and E. Zavaleta.

2008. A checklist for ecological management of landscapes for conservation. Ecology

Letters 11:78-91.

Lindenmayer, D.B., M.L. Hunter, P.J. Burton, and P. Gibbons. 2009. Effects of logging on fire

regimes in moist forests. Conservation Letters. 2:271-277.

Lindenmayer, D.B., J.F. Franklin, A. Lo˜ hmus, S.C. Baker, J. Bauhus, W. Beese, A. Brodie, B.

Kiehl, J. Kouki, G. Mart´ınez Pastur, C. Messier, M. Neyland, B. Palik, A. Sverdrup-

Thygeson, J. Volney, A. Wayne, & L. Gustafsson. 2012. A major shift to the retention

approach for forestry can help resolve some global forest sustainability issues. Conservation

Letters 0 (2012) 1–11.

Littell, J.S., D. McKenzie, D.L. Peterson and A.L. Westerling. 2009. Climate and wildfire area

burned in western U.S. ecoprovinces, 1916-2003. Ecological Applications 19:1003–1021.

Littell, J.S., E.E. Oneil, D. McKenzie, J.A. Hicke, J.A. Lutz, R.A. Norheim and M.M. Elsner.

2010. Forest ecosystems, disturbance, and climate change in Washington State, USA.

Climate Change 102: 129-158.

Loehle, C.L., L. Irwin, J. Beebe, and T. Fleming. 2011. Factors influencing the distribution of

northern spotted owls in the eastern Cascades, Washington. Northwestern Naturalist,

92(1):19-36.

Long, J.N. 2009. Emulating natural disturbance regimes as a basis for forest management: A

North American view. Forest Ecology and Management 257:1868–1873.

Lubowski, R.N., M. Vesterby, S. Bucholtz, A. Baez, and M.J. Roberts. 2006. Major uses of

land in the United States, 2002. U.S. Department of Agriculture, Economic Research

Service, Economic Information Bulletin Number 14. Washington, D.C.

Luoma, D. L., J. M. Trappe, A.W. Claridge, K.M. Jacobs and E. Cazares. 2003. Relationships

among fungi and small mammals in forested ecosystems. Pages 343–373 in C.J. Zabel and

14

R.G. Anthony (editors), Mammal Community Dynamics in Western Coniferous Forests:

Management and Conservation. Cambridge University Press, Cambridge, United Kingdom.

MacDonald, T.L., F.J. Bryan, R.M. Manly, and L.V. Diller. 2006. Discrete-choice modeling in

wildlife studies exemplified by northern spotted owl nighttime habitat selection. The Journal

of Wildlife Management, 70(2): 375-383.

Main, M.B., F.M. Roka, and R.F. Noss. 1999. Evaluating costs of conservation. Conservation

Biology. 13:1262-1272.

Marlon, J.R., Bartlein, P.J., Long, C., Gavin, D.G., Anderson, R.S., Briles, C. Brown, K.J.,

Colombaroli, D., Hallett, D.J., Power, M.J., Scharf, E.A., Walsh, M.K. A long-term

perspective on wildfires in the western U.S. (2012) Proceedings of the National Academy of

Sciences. Published online before print February 14, 2012, doi: 10.1073/pnas.1112839109.

McNab, W.H., and P.E. Avers. 1994a. Ecological subregions of the United States: Chapter 25.

Publication WO-WSA-5. USDA Forest Service, Washington, D.C. Downloaded December

21, 2011 from http://www.fs.fed.us/land/pubs/ecoregions/ch25.html.

McNab, W.H., and P.E. Avers. 1994b. Ecological subregions of the United States: Chapter 32.

Publication WO-WSA-5. USDA Forest Service, Washington, D.C. Downloaded December

21, 2011from http://www.fs.fed.us/land/pubs/ecoregions/ch32.html.

McNab, W.H., and P.E. Avers. 1994c. Ecological subregions of the United States: Chapter 33.

Publication WO-WSA-5. USDA Forest Service, Washington, D.C. Downloaded December

22, 2011 from http://www.fs.fed.us/land/pubs/ecoregions/ch33.html

McNab, W.H., D.T. Cleland, J.A. Freeouf, J.E. Keys, Jr., G.J. Nowacki, and C.A. Carpenter

(comps). 2005. Description of ecological subregions: sections of the conterminous United

States. U.S. Department of Agriculture, Forest Service, Washington, D.C.

Meiman, S., R. Anthony, E. Glenn, T. Bayless, A. Ellinson, M.C. Hansen and C. Smith. 2003.

Effects of commercial thinning on home-range and habitat-use patterns of a male northern

spotted owl: a case study. Wildlife Society Bulletin 31:1254–1262.

Messier, M.S., J.P.A. Shatford, and D.E. Hibbs. 2012. Fire exclusion effects on riparian forest

dynamics in southwestern Oregon. Forest Ecology and Management 264: 60-71.

Meyer, J.S., L.L. Irwin and M.S. Boyce. 1998. Influence of habitat abundance and fragmentation

on northern spotted owls in western Oregon. Wildlife Monographs 139:1–51.

Miller, G.S. 1989. Dispersal of juvenile spotted owls in western Oregon. Thesis, Oregon State

University, Corvallis.

Miller, G.S., R.J. Small, and E.C. Meslow. 1997. Habitat selection by spotted owls during natal

dispersal in western Oregon. Journal of Wildlife Management. 61:140-150.

15

Miller, J. D., H. D. Safford, M. A. Crimmins, and A. E. Thode. 2009. Quantitative evidence for

increasing forest fire severity in the Sierra Nevada and southern Cascade Mountains,

California and Nevada, USA. Ecosystems 12:16–32.

Miller, J.D, C.N. Skinner, H.D. Safford, E.E. Knapp, and C.M. Ramirez. 2012. Trends and

causes of serverity, size, and number of firese in northwestern California, USA. Ecological

Applications 22: 184-203.

Mills, S.L., R.J. Fredrickson, and B.B. Moorhead. 1993. Characteristics of old-growth forests

associated with northern spotted owls in Olympic National Park. Journal of Wildlife

Management 57(2): 315-321.

Mitchell, S.R., M.E. Harmon and K.E.B. O’Connell. 2009. Forest fuel reduction alters fire

severity and long-term carbon storage in three Pacific Northwest ecosystems. Ecological

Applications 19:643–655.

Moilanen A., and H. Kujala. 2008. Zonation: software for spatial conservation prioritization.

User Manual v2.0. Metapopulation Research Group, University of Helsinki, FI. Available at

<http://www.helsinki.fi/bioscience/consplan/software/Zonation/index.html>. Accessed 3

March 2011.

Moilanen, A., A.M.A. Franco, R.E. Early, R. Fox, B. Wintle, and C.D. Thomas. 2005.

Prioritizing multiple-use landscapes for conservation: methods for large multi-species

planning problems. Proceedings of The Royal Society B. 272:1885-1891.

Mori, A. S. 2011. Ecosystem management based on natural disturbances: hierarchical context

and non-equilibrium paradigm. Journal of Applied Ecology 48:280–292.

Moritz, M. A., M.A. Parisien, E. Batllori, M. A. Krawchuk, J. Van Dorn, D. J. Ganz, and K.

Hayhoe. 2012. Climate change and disruptions to global fire activity. Ecosphere 3(6):49.

http://dx.doi.org/10.1890/ES11-00345.1

North, M.P., J.F. Franklin, A.B. Carey, E.D. Forsman, and T. Hamer. 1999. Forest stand

structure of the northern spotted owl's foraging habitat. Forest Science 45(4):520-527.

North, M., and W. Keeton. 2008. Emulating natural disturbance regimes: an emerging approach

for sustainable forest management. Pages 341–372 in R.Lafortezza, J. Chen, G. Sanesi and T.

Crow (editors), Landscape ecology:Sustainable management of forest landscapes. Springer-

Verlag Press, The Netherlands.

Noss, R.F., J.F. Franklin, W.L. Baker, T. Schoennagel, and P.B. Moyle. 2006. Managing fire-

prone forests in the western United States. Frontiers in Ecology and the Environment. 4:481-

487.

16

ODF (Oregon Department of Forestry). 2010a. Northwest Oregon State Forests Management

Plan. Revised Plan April 2010. Oregon Department of Forestry. Salem, Oregon.

ODF (Oregon Department of Forestry). 2010b. Southwest Oregon State Forests Management

Plan. Revised Plan April 2010. Oregon Department of Forestry. Salem, Oregon.

ODF (Oregon Department of Forestry). 2011. Elliott State Forest Management Plan. Final Plan

Draft. Oregon Department of Forestry. Salem, Oregon.

Olson, G.S., E.M. Glenn, R.G. Anthony, E.D. Forsman, J.A. Reid, P.J. Loschl and W.J. Ripple.

2004. Modeling demographic performance of northern spotted owls relative to forest habitat

in Oregon. Journal of Wildlife Management 68:1039–1053.

Olson, G.S., R.G. Anthony, E.D. Forsman, S.H. Ackers, P.J. Loschl, J.A. Reid, K.M Dugger,

E.M. Glenn and W.J. Ripple. 2005. Modeling of site occupancy dynamics for northern

spotted owls, with emphasis on the effects of barred owls. Journal of Wildlife Management

69:918–932.

Perry, D.A., P.F. Hessburg, C.N. Skinner, T.A. Spies, S.L. Stephens, A.H. Taylor, J.F. Franklin,

B. McComb and G. Riegel. 2011. The ecology of mixed severity fire regimes in

Washington, Oregon, and Northern California. Forest Ecology and Management 262: 703-

717.

Phillips S.J., and M. Dudik. 2008. Modeling of species distributions with Maxent: new

extensions and a comprehensive evaluation. Ecography 31:161–175.

Phillips, S.J., R.P. Anderson and R.E. Schapire. 2006. Maximum entropy modeling of species

geographic distributions. Ecological Modeling 190:231–259.

Phillips, S.J., M. Dudik, J. Elith, C.H. Graham, C.H., A. Lehmann, J. Leathwick and S. Ferrier.

2009. Sample selection bias and presence-only distribution models: implications for

background and pseudo-absence data. Ecological Applications 19(1):181-197.

Pious, M. 1995. Home range and habitat use of spotted owls in managed redwood/douglas-fir

forests, California. Report to the California Dept. of Fish and Game. Louisiana-Pacific Corp.

Calpella, CA. 49 pp.

Prather, J.W., R.F. Noss, and T.D. Sisk. 2008. Real versus perceived conflicts between

restoration of ponderosa pine forests and conservation of the Mexican spotted owl. Forest

Policy and Economics. 10:140-150.

Raes, N. and H. ter Steege. 2007. A null-model for significance testing of presence-only

species distribution models. Ecography 30: 727-736.

Ricklefs, R.E. 1979. Ecology. Chiron Press, New York, NY, USA. 966 pp.

17

Rieman, B.E., P.F. Hessburg, C. Luce, and M.R. Dare. 2010. Wildfire and management of

forests and native fishes: conflict or opportunity for convergent solutions? BioScience.

60:460-468.

Reinhardt, E.D., R.E. Keane, D.E. Calkin and J.D. Cohen. 2008. Objectives and considerations

for wildland fuel treatment in forested ecosystems of the interior western United States.

Forest Ecology and Management 256:1997-2006.

Ripple, W.J., P.D. Lattin, K.T. Hershey, F.F. Wagner, and E.C. Meslow. 1997. Landscape

composition and pattern around northern spotted owl nest sites in southwest Oregon. Journal

of Wildlife Management. 61:151-158

Roberts, S.L., J.W. van Wagtendonk, A.K. Miles and D.A. Kelt. 2011. Effects of fire on spotted

owl site occupancy in a late-successional forest. Biological Conservation 144:610–619.

Roloff, G.J., S.P Mealey, and J.D. Bailey. 2012. Comparative hazard assessment for protected

species in a fire-prone landscape. Forest Ecology and Management 277: 1-10.

Rosenberg, D. K., and R.G. Anthony. 1992. Characteristics of northern flying squirrel

populations in young second- and old-growth forests in western Oregon. Canadian Journal of

Zoology 70:161–166.

Rosenberg, D.K., and K.S. McKelvey. 1999. Estimation of habitat selection for centralplace

foraging animals. Journal of Wildlife Management 63:1028–1038.

Rosenberg, D.K., C.J. Zabel, B.R. Noon, and E.C. Meslow. 1994. Northern spotted owls:

influence of prey base—a comment. Ecology. 75:1512-1515.

Rosenvald, R., and A. Lohmus. 2008. For what, when, and where is green-tree retention better

than clear-cutting? A review of the biodiversity aspects. Forest Ecology and Management

255:1-15.

Safford, H.D., J.T. Stevens, K. Merriam, M.D. Meyer, and A. M. Latimer. 2012. Fuel treatment

effectivenss in northern California yellow pine and mixed conifer forests. Forest Ecology

and Management 274: 17-28.

Sakai, H.F., and B.R. Noon. 1993. Dusky-footed woodrat abundance in different-aged forests in

northwestern California. Journal of Wildlife Management 57:373–382.

Sakai, H.F., and B.R. Noon. 1997. Between-habitat movement of dusky-footed woodrats and

vulnerability to predation. Journal of Wildlife Management. 61:343-350.

Schumaker, N.H. 2008. HexSim (Version 1.3.6.9). U.S. Environmental Protection Agency,

Environmental Research Laboratory, Corvallis, Oregon. Available online at

<http://www.epa.gov/hexsim>. Accessed on 3 March 2011.

18

Schumaker, N. H. 2011. HexSim. Version 2.1. U.S. Environmental Protection Agency, Corvallis,

Oregon. Available online at <http://www.epa.gov/hexsim>. Accessed on 3 March 2011.

Schoennagel T, and C.R. Nelson.2011. Restoration relevance of National Fire Plan treatments

across the western US. Frontiers in Ecology and the Environment 9:271–277.

Seymour, R.S., and M.L. Hunter, Jr. 1999. Principles of ecological forestry. Pages 22-61 in M.L.

Hunter, Jr. (editor), Maintaining Biodiversity in Forest Ecosystems. Cambridge University

Press, Cambridge, United Kingdom.

Sierra Legal Defence [sic] Fund and Western Canada Wilderness Committee. 2005. In defence

[sic] of Canada’s spotted owl. Sierra Legal Defence Fund, Vancouver, British Columbia,

Canada.

Singleton, P.H., J.F. Lehmkuhl, W.L. Gaines, and S.A. Graham. 2010. Barred owl space use and

habitat selection in the eastern Cascades, Washington. Journal of Wildlife Management

74:285–294.

Sisco, C.L. 1990. Seasonal home range and habitat ecology of spotted owls in northwestern

California. Thesis, Humboldt State University, Arcata, California.

Sisk, T.D., M. Savage, D.A. Falk, C.D. Allen, E. Muldavin, and P. McCarthy. 2005. A

landscape perspective for forest restoration. Journal of Forestry. 103:319-320.

Skinner, C.N. 1995. Change in spatial characteristics of forest openings in the Klamath

Mountains of northwestern California, USA. Landscape Ecology. 10:219-228.

Skinner, C.N., and A.H. Taylor. 2006. Southern Cascades bioregion. Pages 195–224 in N.G.

Sugihara, J.W. van Wagtendonk, K.E. Shaffer, J. Fites-Kaufman and A.E. Thode (editors),

Fire in California’s Ecosystems. University of California Press, Berkeley.

Skinner, C.N., A.H. Taylor and J.K. Agee. 2006. Klamath Mountains bioregion. Pages 170–194

in N.G. Sugihara, J.W. van Wagtendonk, K.E. Shaffer, J. Fites-Kaufman and A.E. Thode

(editors), Fire in California’s Ecosystems. University of California Press, Berkeley.

Smith, F.A. 1997. Neotoma cinerea. Mammalian Species 564:1–8.

Solis, D.M. 1983. Summer habitat ecology of spotted owls in Northwestern California. Thesis,

Humboldt State University, Arcata, California.

Solis, D.M., and R.J. Gutiérrez. 1990. Summer habitat ecology of northern spotted owls in

northwestern California. Condor 92:739–748.

Spies, T.A., M.A. Hemstrom, A. Youngblood, and S. Hummel. 2006. Conserving old-growth

forest diversity in disturbance-prone landscapes. Conservation Biology. 20:351-362.

19

Spies, T.A., K.N. Johnson, K.M. Burnett, J.L. Ohmann, B.C. McComb, G.H. Reeves, P.

Bettinger, J.D. Kline, and B. Garber-Yonts. 2007a. Cumulative ecological and

socioeconomic effects of forest policies in coastal Oregon. Ecological Applications 17:5–

17.

Spies, T.A., B.C. McComb, R.S.H. Kennedy, M.T. McGrath, K. Olsen, and R.J. Pabst. 2007b.

Potential effects of forest policies on terrestrial biodiversity in a multi-ownership province.

Ecological Applications 17(1):48-65.

Spies, T.A., T.W. Geisen, F.J. Swanson, J.F. Franklin, D. Lach and K.N. Johnson. 2010. Climate

change adaptation strategies for federal forests of the Pacific Northwest, USA: ecological,

policy, and socio-economic perspectives. Landscape Ecology 25:1185–1199.

Stalberg, D., K.E. Fehring, L.Y. Pomara, N. Nur, D.B. Adams, D. Hatch, G.R. Geupel, and S.

Allen. 2009. Modeling nest-site occurrence for the northern spotted owl at its southern limit

in central California. Landscape and Urban Planning 90:76-85.

Stenseth, N.C., A. Mysterud, G. Ottersen, J. W. Hurrel, K-S Chan, and M. Lima. 2002.

Ecological effects of climate fluctuations. Science 297:1292-1296.

Stephens, S.L., J.J. Moghaddas, C. Ediminster, C.E. Fiedler, S. Hasse, M.Harrington, J.E.

Keeley, J.D. McIver, K. Metlen, C.N. Skinner, and A.Youngblood. 2009. Fire treatment

effects on vegetation structure, fuels, and potential fire severity in western U.S. forests.

Ecological Applications 19: 305-320.

Stephens, S.L., R.E.J. Boerner, J.J. Moghaddas, E.E.Y. Moghaddas, B.M. Collins, C.B. Dow, C.

Edminster, C.E. Fiedler, D.L. Fry, B. R. Hartsough, J.E. Keeley, E.E. Knapp, J.D. McIver,

C.N. Skinner, and A. Youngblood. 2012a. Fuel treatment impacts on estimated wildfire

carbon loss from forests in Montana, Oregon, California, and Arizona. Ecosphere 3(5):38.

http://dx.doi.org/10.1890/ES11-00289.1.

Stephens, S.L., J.D. McIver, R.E.J. Boerner, C.J. Fettig, J.B. Fontaine, B.R. Hartsough, P.

Kennedy, and D.W. Schwilk. 2012b. Effects of forest fuel reduction treatments in the United

States. BioScience 62:549-560.

Swanson, F.J., Kratz T.K., Caine N., and Woodmansee RG. 1988. Landform effects on

ecosystem patterns and processes. BioScience 38: 92–98.

Swanson, M.E, J.F. Franklin, R.L Beschta, C.M. Crisafulli, D.A. DellaSala, R.L Hutto, D.B.

Lindenmayer and F.J. Swanson. 2011. The forgotten stage of forest succession: early-

successional ecosystems on forest sites. Frontiers in Ecology and the Environment. doi:

10.1890/090157.

Swindle, K.A., W.J. Ripple, E.C. Meslow and D. Schafer. 1999. Old forest distribution around

spotted owl nests in the central Cascade Mountains, Oregon. Journal of Wildlife

Management 63:1212–1221.

20

Syphard, A.D., J.E. Keeley, and T.J. Brennan. 2011. Comparing the role of fuel breaks across

southern California national forests. Forest Ecology and Management 261: 2038-2048.

Thomas, J.W., E.D. Forsman, J.B. Lint, E.C. Meslow, B.R. Noon and J. Verner. 1990. A

conservation strategy for the northern spotted owl. Interagency Scientific Committee to

Address the Conservation of the Northern Spotted Owl. U.S. Forest Service, U.S. Bureau of

Land Management, U.S. Fish and Wildlife Service, and U.S. National Park Service, Portland,

Oregon.

Thomas, J.W., J.F. Franklin, J. Gordon, and K.N. Johnson. 2006. The Northwest Forest Plan;

origins, components, implementation experience, and suggestions for change. Conservation

Biology. 20:277-287.

Thome, D.M., C.J. Zabel and L.V. Diller. 1999. Forest stand characteristics and

reproduction of northern spotted owls in managed north-coastal California forests. Journal of

Wildlife Management 63(1):44-59.

Thompson, J.R., S.L. Duncan, and K.N. Johnson. 2009. Is There Potential for the Historical

Range of Variability to Guide Conservation Given the Social Range of Variability? Ecology

and Society 14(1):18. [online] URL: http://www.ecologyandsociety.org/vol14/iss1/art18/

Tidwell, T. 2011. Statement before the Committee on Energy and Natural Resources; U.S.

Senate. June 14, 2011

USDA (U.S. Department of Agriculture). 2010. The Okanogan-Wenatchee National Forest

Restoration Strategy: adaptive ecosystem management to restore landscape resiliency. U.S.

Forest Service, Pacific Northwest Region. September 2010. 99 pp.

USDA (U.S. Department of Agriculture), and USDI (U.S. Department of the Interior). 1994a.

Record of decision for amendments to Forest Service and Bureau of Land Management

planning documents within the range of the northern spotted owl: standards and guidelines

for management of habitat for late-successional and old-growth forest related species within

the range of the northern spotted owl. U.S. Forest Service and U.S. Bureau of Land

Management, Portland, Oregon.

USDI (U.S. Department of Interior). 1992. Draft final recovery plan for the northern spotted owl.

U.S. Fish and Wildlife Service, Portland, Oregon.

USFWS (U.S. Fish and Wildlife Service). 2004. Northern spotted owl: Five Year Review

Summary and Evaluation. U.S. Fish and Wildlife Service, Portland, Oregon.

USFWS (U.S. Fish and Wildlife Service). 2011a. Revised recovery plan for the northern spotted

owl (Strix occidentalis caurina). U.S. Fish and Wildlife Service, Portland, Oregon. Xvi +

258 pp.

21

USFWS (U.S. Fish and Wildlife Service) 2011b. Biological opinion on Pilot Joe Project,

Medford District, Bureau of Land Management, Oregon. August 10, 2011. 77pp.

USFWS (U.S. Fish and Wildlife Service) and NMFS (National Marine Fisheries Service). 1998.

Endangered Species Consultation Handbook: Procedures for Conducting Consultation and

Conference Activities Under Section 7 of the Endangered Species Act. Washington, D.C.

Van Gelder, J.J. 2003. Variation in four-note location calls of male spotted owls (Strix

occidentalis). Thesis, Humboldt State University, Arcata, California.

Walther, G. R., E. Post, P. Convoy, A. Menzel, C. Parmesan, T. J. C. Beebee, J. Fromentin, O.

Hoegh-Guldberg, and F. Bairlein. 2002. Ecological responses to recent climate change.

Nature 416: 389-395.

Ward, J.W. Jr., R.J. Gutiérrez and B.R. Noon. 1998. Habitat selection by northern spotted owls:

the consequences of prey selection and distribution. Condor 100:79–92.

Waters, J.R., K.S. McKelvey, C.J. Zabel and W.W. Oliver. 1994. The effects of thinning and

broadcast burning on sporocarp production of hypogeous fungi. Canadian Journal of Forest

Research 24:1516–1522.

Weathers, W.W., P.J. Hodum and J.A. Blakesley. 2001. Thermal ecology and ecological

energetics of the California spotted owl. Condor 103:678–690.

Westerling, A.L., M.G. Turner, E.A.H. Smithwick, W.H. Romme, and M. G. Ryan. 2011.

Continued warming could transform Greater Yellowstone fire regimes by mid-21st century

Proceedings of the National Academy of Science 108:13165-13170.

Wiens, J. D. 2012. Competitive interactions and resource partitioning between northern spotted

owls and barred owls in western Oregon. PhD dissertation, Oregon State University,

Corvallis, OR.

Wiens, J. D., R. G. Anthony, and E. D. Forsman. 2011. Barred owl occupancy surveys within the

range of the northern spotted owl. Journal of Wildlife Management 75:531-538.

Wilcove, D.S., and L.Y. Chen. 1998. Management costs for Endangered Species. Conservation

Biology. 12:1405-1407.

Willems, E.P., and R. A. Hill. 2009. A critical assessment of two species distribution models: a

case study of the vervet monkey ( Cercopithecus aethiops ). Journal of Biogeography 36:

2300-2312.

Williams, M.A. and W.L. Baker. 2012. Spatially extensive reconstructions show variable-

severity fires and heterogeneous structure in historical western United States dry forests.

Global Ecology and Biogeography 21:1042-1052.

22

Wilson, T. M. 2010. Limiting factors for northern flying squirrels (Glaucomys sabrinus) in the

Pacific Northwest: A spatio-temporal analysis. Ph.D Dissertation. Union Institute and

University, Cincinnati, Ohio, USA.

Wisz, M.S., R.J. Hijmans, J. Lis, A.T. Peterson, C.H. Graham and A. Guisan. 2008. Effects of

sample size on the performance of species distribution models. Diversity and Distributions

14:763-773.

Wimberly, M.C., T.A. Spies and E. Nonaka. 2004. Using criteria based on natural fire regime to

evaluate forest management in the Oregon Coast Range of the United States. Pages 146–157

in A.H. Perera, L.J. Buse, M.G. Weber (editors), Emulating Natural Forest Landscape

Disturbances: Concepts and Applications. New York: Columbia University Press.

Zabel, C.J., K.M. McKelvey and J.P. Ward, Jr. 1995. Influence of primary prey on home-range

size and habitat-use patterns of northern spotted owls (Strix occidentalis caurina). Canadian

Journal of Zoology 73:433–439.

Zabel, C.J., J.R. Dunk H.B. Stauffer, L.M. Roberts, B.S. Mulder and A. Wright. 2003. Northern

spotted owl habitat models for research and management application in California (USA).

Ecological Applications 13:1027–1040.

In Litt. References

Steucke, Paul. 2008. Letter dated December 10, 2008 from Mr. Steucke to Ken Berg. Paul

Steucke, Chief, Environmental and Natural Resources Division, Public Works, Joint Base

Lewis-McChord.

USFWS (U.S. Fish and Wildlife Service). 2004. Application of the ―Destruction or Adverse

Modification‖ Standard under Section 7(a)(2) of the Endangered Species Act. Memo from

Acting Director Marshall Jones to Regional Directors. Dated December 9, 2004.