Lesson Overview 20.2 Prokaryotes (2)

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    Lesson Overview20.2 Prokaryotes

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    THINK ABOUT IT

    Imagine living all your life as a member of what you believe is the

    only family on your street. Then, one morning, you open the front

    door and discover houses and neighbors all around you.

    Where did all the people come from? What if the answer turned

    out to be that they had always been thereyou just hadnt seenthem? How would your view of the world change?

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    THINK ABOUT IT

    When the microscope was first invented, we humans had just such a

    shock. Far from being alone, we share every corner of our world with

    microorganisms. Even a seemingly clean toothbrush contains a film

    of bacteria on its bristles!

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    Classifying Prokaryotes

    How are prokaryotes classified?

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    Classifying Prokaryotes

    How are prokaryotes classified?

    Prokaryotes are classified as Bacteria or Archaeatwo of the three

    domains of life.

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    Classifying Prokaryotes

    The smallest and most abundant

    microorganisms on Earth are

    prokaryotesunicellular organisms

    that lack a nucleus.

    Prokaryotes have DNA, like all othercells, but their DNA is not found in a

    membrane-bound nuclear envelope

    as it is in eukaryotes. Prokaryote

    DNA is located in the cytoplasm.

    A bacterium such as E. coli has the

    basic structure typical of most

    prokaryotes.

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    Classifying Prokaryotes

    Recently, biologists have divided prokaryotes into two very distinct groups:

    Bacteria and Archaea.

    These groups are very different from each other; therefore, biologists now

    consider each group of prokaryotes as a separate domain. Eukaryotes are

    the third domain.

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    Bacteria

    The larger of the two domains of

    prokaryotes is the Bacteria.

    Bacteria include a wide range of

    organisms with lifestyles so different

    that biologists do not agree exactlyhow many phyla are needed to

    classify this group.

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    Bacteria

    Bacteria live almost everywherein

    fresh water, in salt water, on land,

    and on and within the bodies of

    humans and other eukaryotes.

    Escherichia coli, a typical bacteriumthat lives in human intestines, is

    shown.

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    Bacteria

    Bacteria are usually surrounded by

    a cell wall that protects the cell from

    injury and determines its shape.

    The cell walls of bacteria contain

    peptidoglycana polymer of sugarsand amino acids that surrounds the

    cell membrane.

    Some bacteria, such as E. coli,

    have a second membrane outsidethe peptidoglycan wall that makes

    the cell especially resistant to

    damage.

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    Bacteria

    In addition, some prokaryotes have

    flagella that they use for

    movement, or pili, which in E. coli

    serve mainly to anchor the

    bacterium to a surface or to other

    bacteria.

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    Archaea

    Under a microscope, archaea look very similar to bacteria. Both are

    equally small, lack nuclei, and have cell walls, but there are important

    differences.

    The walls of archaea lack peptidoglycan, and their membranes contain

    different lipids.

    The DNA sequences of key archaea genes are more like those of

    eukaryotes than those of bacteria.

    Based on these observations, scientists have concluded that archaeaand eukaryotes are related more closely to each other than to bacteria.

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    Archaea

    Many archaea live in extremely harsh environments.

    One group of archaea produce methane gas and live in

    environments with little or no oxygen, such as thick mud and the

    digestive tracts of animals.

    Other archaea live in extremely salty environments, such as Utahs

    Great Salt Lake, or in hot springs where temperatures approach the

    boiling point of water.

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    Structure and Function

    How do prokaryotes vary in their structure and function?

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    Structure and Function

    How do prokaryotes vary in their structure and function?

    Prokaryotes vary in their size and shape, in the way they move, and in the

    way they obtain and release energy.

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    Size, Shape, and Movement

    Prokaryotes range in size from 1 to 5 micrometers, making them much

    smaller than most eukaryotic cells. Prokaryotes come in a variety of

    shapes.

    Rod-shaped prokaryotes are called bacilli.

    Spherical prokaryotes are called cocci.

    Spiral and corkscrew-shaped prokaryotes are called spirilla.

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    Size, Shape, and Movement

    Prokaryotes can also be distinguished by whether they move and how

    they move.

    Some prokaryotes do not move at all. Others are propelled by flagella.

    Some glide slowly along a layer of slimelike material they secrete.

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    Nutrition and Metabolism

    Prokaryotes need a supply of chemical energy, which they store in the

    form of fuel molecules such as sugars.

    Energy is released from these fuel molecules during cellular respiration,

    fermentation, or both.

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    Nutrition and Metabolism

    Prokaryotes vary in the ways they obtain energy and the ways they

    release it.

    Looking at the two tables on the following slides, notice that some

    species are able to change their method of energy capture or release

    depending on the conditions of their environment.

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    Nutrition and Metabolism: Energy

    Capture

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    Nutrition and Metabolism: Energy

    Release

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    Growth, Reproduction, and

    RecombinationWhen a prokaryote has grown so that it has nearly doubled in size, it

    replicates its DNA and divides in half, producing two identical cells.

    This type of reproduction is known as binary fission.

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    Growth, Reproduction, and

    Recombination

    Because binary fission does not involve the exchange or

    recombination of genetic information, it is an asexual form of

    reproduction.

    When conditions are favorable, prokaryotes can grow and divide atastonishing ratessome as often as once every 20 minutes!

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    Growth, Reproduction, and

    Recombination

    When growth conditions become unfavorable, many prokaryotic

    cells form an endosporea thick internal wall that encloses the

    DNA and a portion of the cytoplasm.

    Endospores can remain dormant for months or even years.

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    Growth, Reproduction, and

    Recombination

    The ability to form endospores makes it possible for some

    prokaryotes to survive very harsh conditions. The bacterium Bacillus

    anthracis, which causes the disease anthrax, is one such bacterium.

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    Mutation

    Mutations are one of the main ways prokaryotes evolve.

    Mutations are random changes in DNA that occur in all organisms.

    In prokaryotes, mutations are inherited by daughter cells produced by

    binary fission.

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    Conjugation

    Many prokaryotes exchange genetic information by a process called

    conjugation.

    During conjugation, a hollow bridge forms between two bacterial

    cells, and genetic material, usually in the form of a plasmid, moves

    from one cell to the other.

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    Conjugation

    Many plasmids carry genes that enable bacteria to survive in new

    environments or to resist antibiotics that might otherwise prove fatal.

    This transfer of genetic information increases genetic diversity in

    populations of prokaryotes.

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    The Importance of Prokaryotes

    What roles do prokaryotes play in the living world?

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    The Importance of Prokaryotes

    What roles do prokaryotes play in the living world?

    Prokaryotes are essential in maintaining every aspect of the ecological

    balance of the living world. In addition, some species have specific uses in

    human industry.

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    Decomposers

    Bacteria called actinomycetes are present in soil and in rotting plant

    material such as fallen logs, where they decompose complex organic

    molecules into simpler molecules.

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    Decomposers

    By decomposing dead organisms, prokaryotes, supply raw materials

    and thus help to maintain equilibrium in the environment.

    Bacterial decomposers are also essential to industrial sewage

    treatment, helping to produce purified water and chemicals that can be

    used as fertilizers.

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    Producers

    Cyanobacteria in the genusAnabaena form filamentous chains in ponds

    and other aquatic environments, where they perform photosynthesis.

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    Producers

    Photosynthetic prokaryotes are among the most important producers on

    the planet.

    Food chains everywhere are dependent upon prokaryotes as producers

    of food and biomass.

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    Nitrogen Fixers

    All organisms need nitrogen to make proteins and other molecules.

    Nitrogen gas (N2) makes up 80 percent of Earths atmosphere, but only

    a few kinds of organismsall of them prokaryotescan convert N2into

    useful forms.

    The process of nitrogen fixation converts nitrogen gas into ammonia

    (NH3). Ammonia can then be converted to nitrates that plants use, or

    attached to amino acids that all organisms use.

    Nitrogen-fixing bacteria and archaea provide 90 percent of the nitrogenused by other organisms.

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    Nitrogen Fixers

    Some plants have symbiotic relationships with nitrogen-fixing

    prokaryotes.

    The bacterium Rhizobium grows in nodules, or knobs, on the roots of

    legume plants such as soybean.

    The Rhizobium bacteria within these nodules convert nitrogen in the air

    into the nitrogen compounds essential for plant growth.

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    Nitrogen Fixers

    The Rhizobium bacteria often live symbiotically within nodules attached

    to roots of legumes, such as clover, where they convert atmospheric

    nitrogen into a form that is useable by plants.

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    Human Uses of Prokaryotes

    Prokaryotes, especially bacteria, are used in the production of a wide

    variety of foods and other commercial products.

    Yogurt is produced by the bacterium Lactobacillus.

    Some bacteria can digest petroleum and remove human-made waste

    products and poisons from water.

    Other bacteria are used to synthesize drugs and chemicals through the

    techniques of genetic engineering.

    Bacteria and archaea adapted to extreme environments may be a rich

    source of heat-stable enzymes that can be used in medicine, food

    production, and industrial chemistry.