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    Implications of Changes to the Mekong River on Fish and Fishers

    Livelihoods from Hydropower: A Case in the Tonle Sap Cambodia

    Mak Sithirith

    Fisheries Action Coalition Team (FACT)

    Paper prepared for the workshop on Mekong Environment and Livelihood:

    The Changing situation and Trans-boundary Implications

    Cantho City, Vietnam

    3-4 February 2010

    CAMBODIA covers an area of 181,035 km2. It borders Vietnam in the east, Laos in

    the northeast, and Thailand in the north and west. The Gulf of Thailand borders

    Cambodia to the south. The estimated 2008 population of Cambodia was 13.4 million

    in 24 provinces, consisting of 185 districts, 1,621 communes and more than 14,073

    villages1

    (MoP, 2004 & 2006; RGC, 2006; NIS, 2008). Approximately four-fifths of

    the Cambodian population is rural (World Bank 2004). This population is centred

    primarily around the Mekong River and the Tonle Sap Lake (Lamberts, 2001). Of the

    total number of villages in Cambodia, about 1158 villages are located in the Tonle

    Sap floodplain and of these villages; about 170 are floating villages (Keskinen, 2003;

    MoP, 1999). The Tonle Sap Lake plays a vital role in Cambodian economy andlivelihoods for millions of Cambodians. Many people are directly dependent on

    resources from the Lake.

    The Tonle Sap and the Mekong River

    The Tonle Sap was formed about 5500-6,000 years ago (Penny, 2002; Penny

    et al., 2005; Tsukawaki, 1997). The Tonle Sap Basin is located in the northwest part

    of Cambodia between approximately latitudes 102 15' to 105 50' E and longitudes

    11 40' to 14 28' N that covers an area of about 80,000 km, including the Tonle Sap

    Lake, which constitutes about 44% of the total land area of the country (Wright et al.,

    2004). About 95% of the Tonle Sap Basin lies within Cambodia while about 5 percent

    (about 5,000 km) lies in Thailand (CNMC &NEDECO, 1998; ADB, 2004).

    The Tonle Sap Basin extends over 44% of Cambodias total land area, and is

    home to 32% of Cambodias population, or about 3.6 million people, covering 8

    1 www.nis.gov.kh/index.php/statistics/surveys/census2008/provincial-population-totals.

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    provinces in Cambodia2. It constitutes a critical part of the lower Mekong basin: it

    supplies 6% of the average annual flow of the Mekong but, more significantly, makes

    up 16% of its dry season flow (thereby helping to control salinity intrusion and

    conserve mangrove in the Mekong delta) (ADB, 2004)3.

    The average annual inflow into the Tonle Sap lake is estimated at about 79km

    (Kummu et al., 2008); about 30% of the flow originates from 13 tributaries located in

    the lakes's watershed (Kummu et al., 2008; Sithirith, 2007)4. Due to its distinct

    hydrology,from the Mekong mainstream, any long term decrease in mainstream

    Mekong discharge due to upstream Chinese hydro development will result in a 40-

    50% increase in the contribution of local catchment runoff as a percentage of the total

    water supply to Tonle Sap Lake (ADB, FAO, DoF, 2003).5

    The Tonle Sap River connects the Tonle Sap Lake with the Mekong River. It

    flows from the Southeastern end of the Tonle Sap Lake and meets the Mekong River

    at Chatomouk confluence, north of Phnom Penh, after which the river immediately

    splits into the smaller Bassac River and the larger Mekong River (Kummu et al.,

    2008). The Tonle Sap River has total length of 120 km extending from Phnom Penh

    to Kampong Chhnang province (CNMC, 2004; Keskinen, 2003).

    In the wet season (May to December), flooding in the Mekong River causes

    the Tonle Sap River to change its direction of flow northwestward into the Tonle Sap

    Lake causing a dramatic rise in water level. About 52% of inflow into the lake is from

    the Tonle Sap River (Kummu et al., 2008). In the dry season, the water reverses its

    flow to the Mekong River. Thus, the Tonle Sap River plays a vital role in sustaining

    the Tonle Sap Lake. The inflow and outflow moves fish, nutrients and aquatic

    productivity to and from the lake, making it unique in the world (Van Zalinge et al.,

    2000; Degen and Thouk, 2000).

    The Tonle Sap Lake is an integral part of the Mekong River and it is the

    largest freshwater body in Southeast Asia. The seasonal variations in the lake's area

    2 The Tonle Sap Basin covers 8 provinces in Cambodia: Battambang, Siem Reap, Pursat, KampongThom, Kampong Chhang, Banteay Meanchey, Preah Vihar and Oddar Meanchey (ADB, 2004).3 ADB (2005a). Tonle Sap and its fisheries: Future solutions now. The Tonle Sap Initiative; Asian

    Development Bank, Manila, Philippines.4 Some 13 major tributaries contribute a high volume of water to the Lake including the Stung Sen,

    Stung Sangke, Stung Chinit, Stung Chikreng, Stung Stung, Stung MonkulBorei Rivers, all of them

    located in the watershed areas of the Tonle Sap Basin (Sithirith, 2007).5

    (REF?) suggests that the resident dry season stocks of some species in the Great Lake may representseparate stocks from those which are carried into the Lake by the annual back-flooding of the Mekong

    River.

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    and depth are remarkable. The lake is usually illustrated in the maps in its smallest,

    dry season form. The length of the lake varies from approximately 160km long6

    and

    35km wide during the dry season7, to 250km long and almost 100km wide during the

    peak of the flooded seasons (Matsui et al, 2005; Kummu and Sarkkula, 2008)8.

    The Tonle Sap Lake has an exceptional water regime. The lake covers an area

    varying between 250,000 -300,000 ha in the dry season with an average depth of less

    than two meters9, and it increases to 1.0-1.3 million ha

    10in the rainy season with a

    maximum depth of often more than 10 m (CNMC and NEDECO, 1998; ADB, FAO

    and DoF, 2003; MRC, 200311

    ). The water level in the Tonle Sap Lake varies from

    1.00-1.44 m in the dry season to 8.00-9.5m in the wet season, but some years

    increases to 11m (Kummu et al., 2008; Nikula, 2004; ADB, 2005a).

    During the wet season, the volume of the lake increases from about 1.3 km 3

    during the dry season up to 79-83 km3, depending on the flood intensity. The bottom

    of the lake lies approximately 0.50.7 m above the mean sea level in Hatien datum

    (Kummu et al., 2008).

    Table 1. The size of the Tonle Sap Lake by the dry and wet seasonSeason Lake area

    (Km2)

    Water level

    (m)

    Lake area

    (Km2)

    Water

    level(m)

    Lake area

    (Km2)

    Water

    level (m)

    Dry season

    lake area

    2,300 1.44 2,500 1.2 2,500

    3,000

    1-2

    Wet season

    lake area

    13,260 9.17 15,000 9 10,000-

    16,000

    811

    Source Kummu et al., 2008 Nikula, 2004 ADB, 2005

    Tuk tonle (river water) is active in the wet season from May to October.

    During May and June, Tuk tonle increases only gradually. In July and August it

    increases noticeably when the Mekong River rises and flows into the Tonle Sap Lake

    due to the water level rise in the Mekong resulting from heavy rains and snow melt on

    6 Heinonen U. unpublished thesis(ref? not clear) (2004) estimates that the length of the lake is 140 km.

    The dry season covers an area of 2,590 km2 and in rainy season 10,400 km2. Heinonen, U. (2004)

    Integrated and socially just water resource management in the Lower Mekong River Region and

    Cambodia: How to control water related rural push. Helsinki University of Technology. Department ofCivil and Environmental Engineering, Laboratory of Water Resources, Finland.7 Somony, T. and Schmidt, U. 2004. Aquatic Resource Management: Tonle Sap Great Lake,Cambodia. Somony and Schmidt, 2004 estimates that length of the Lake is about 100 km long and 35

    wide8 (Matsui, S. et al., 2005. Tonle Sap: Experience and Lessons Learned Brief.9 Somony, T. and Schmidt, U. 2004. Aquatic Resource Management: Tonle Sap Great Lake,

    Cambodia. Department of Fisheries. In this paper, it is estimated that the water level in the dry season

    is estimated at 0.8-1m and in rainy season 10-12m.10

    Nikula (2004) states that the size of the lake increases from 2,700 km2

    to 9,000 - 16,000 km2

    (MRCS/MRC/WUP-FIN, 2003).11 MRC, 2003. State of the Basin Report. Mekong River Commission.

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    the Tibetan Plateau (Lambert, 2001). The flow of water from the Mekong to the Tonle

    Sap results in rising water. The average annual inflow varies, but it is estimated at

    about 79-83 km (See Table 2). It comes from four sources; the reverse flow from

    Mekong River via the Tonle Sap River; the tributaries around the Lake; the rainfall;

    and the overland flow from the Mekong. It is estimated that about 40-45 km3

    of water

    comes from the Mekong River and 4 km from the overland flow. These two sources

    account for 57 percent of the water volume in the Tonle Sap Lake. Approximately 24-

    25 km3

    or 30 percent of water comes from the Tonle Sap tributaries, and rainfall

    provides 10-14 km3

    or the remaining 13 percent (Kummu et al., 2008; Matsui et al.,

    2005; CNMC, 2004). The annual average outflow of the outflow the Tonle Sap Lake

    is estimated at about 78.6 km. About 69 km (88% of outflow) from the Tonle Sap

    Lake returns to the Mekong River via the Tonle Sap River (Kummu et al., 2008;

    Matsui et al, 2005; CNMC, 2004).

    Table 2. The Water Volume of the Tonle Sap

    Water Source

    Storing capacity of

    TSL estimated

    by Kumu et al., 2008(km3)

    Storing capacity of

    TSL estimated

    By Matsui et al (km3)

    Storing capacity of

    TSL estimated

    by CNMC (km3)

    Estimate

    (km3)

    Mekong 40.64 45 45 40-45

    Tributary 24.7 24 24.3 24-25

    Overland flow 3.98 n/a n/a 4

    Precipitation 10.36 14 13.9 10-14

    Total 79.68 83 83.2 79-83

    Source: Kumu et al., 2008; Matsui et al, 2005; CNMC, 2004.

    The Tonle Sap as a Heart of the Mekong Region

    The lake is unique due to the natural phenomenon of reverse water flow. One

    half of the annual pulse in which the lake fills from the Mekong River during the wet

    season (May to October) and empties back during the dry season.12 This is the flood

    pulse of the Tonle Sap. Anders Poulsen calls the Tonle Sap Lake the pulsating heart

    of the Mekong, and goes on to say the flood pulse is what keeps the heart beating. If

    the heart stops, the system dies (Poulsen in Nikkula, 2005)13

    .The entire ecosystem

    would be adversely transformed, In theis case, fisheries would collapse, indigenous

    knowledge would be subverted, the poor would go hungry, livelihoods would be

    12 In the wet season, the surface area of the Lake increases from 250,000-300,000 ha to approximately

    1.0-1.6 million ha, with depth increasing correspondingly from 12 m amsl to 911 m amsl (CNMC

    and Nedeco, 1998), and storage capacity reaching a maximum of 80 million cubic meters (Sopharith,

    1998). It absorbs 20 per cent of the Mekong River's floodwaters and serves as a flood regulator (MRC,

    2004; ADB, 2002). The drop of the water level in the Mekong in the dry season creates the reverseflow from the Lake into the Mekong.13 Jussi Nikkula, The Lake and its People. MSc Thesis, Helsinki University of Technology, 2005.

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    disrupted, the communities would be unravelled, and the national economy would

    suffer. Tonle Sap is an critically important area for the Mekong Region that requires

    effective protection (Nikula, 2005)14

    .

    The Tonle Sap a Natural Reservoir for Flood Reduction in the Mekong Region

    The Mekong has one flood pulse a year (Lamberts 2001). During May and

    June when the Southwest monsoon arrives, the water level in the Mekong River

    increases only gradually, but does not increase noticeably until July and August when

    the Mekong River rises. Other streams originating in two chains of mountain, the

    Phnom Dangreks and Phnom Kravagn (Cardamom chain), also flow into this large

    reservoir. During September and October, water in the Mekong River is at its

    maximum level in Kratie province (Nikula, 2005). Annually, the Mekong River

    releases an estimate of 475 km to the China Sea (Kummu et al., 2008). The Tonle

    Sap accepts about 10 percent of the Mekong volume, thus reducing floods in the

    Mekong Delta15

    (Kummu et al., 2008; Matsui et al, 2005; CNMC, 2004). The

    flooded area in the Mekong Delta in Vietnam is thus maintained between 1.2 and 1.4

    million hectares16

    . However, under the high flood, the flooded areas in the Mekong

    Delta in Vietnam may increase to 1.9 million hectares (Anh Tuan et al., 2007)17.

    The Tonle SapReducing Saltwater Intrusion into the Mekong Delta

    Apart from absorbing water from the Mekong River in the wet season, the

    Tonle Sap Lake also releases water from the Lake to the Mekong River in the dry

    season downstream to the Mekong Delta and the South China Sea. It subsides slowly

    until January and February and then continues to gradually recede during March and

    April. Out-flow water from the floodplain and the lake increases the flow of water in

    the lower Mekong River, improving conditions in the Mekong estuary by reducing

    saltwater intrusion during the dry period. About 2.1 million hectares of the Mekong

    Delta are affected by the salinity during the dry season (Anh Tuan et al., 2007). The

    water released from this storage can also be used to irrigate the dry season crops in the

    Mekong delta (MRC 1998). In this sense, the Tonle Sap Lake is considered by many

    14 Jussi Nikkula, The Lake and its People. MSc Thesis, Helsinki University of Technology, 2005.15 Kummu et al (2008) estimates that the annual inflow of the Tonle Sap is estimated at about 79 km.

    About 57% of water in the Tonle Sap originates from the Mekong main stem.16

    The Mekong Delta in Vietnam covers an area of 39,200 km (Kakonen, 2008).17 Thanh Be et al., 2007. Challenges to Sustainable Development in the Mekong Delta: Regional and

    National Policy Issues and Research Needs.

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    Khmers to be the backbone of their struggling nations agricultural system

    (Longstreath 2001).

    The Tonle Sap as Fisheries Space in the Mekong River

    The flow of water from the Mekong River to the Tonle Sap Lake during the

    flood season and the release of water from the Tonle Sap Lake back to the Mekong

    River during the dry season is not only important in terms of reducing floods and

    preventing saltwater intrusion, but also critical for fish migration and productivity in

    both the Mekong and Tonle Sap Lake.

    The inflow of water from the Mekong to the Tonle Sap Lake during the wet

    season induces the fish migration from the river into the lake. The water flowing into

    the Tonle Sap Lake inundates the flooded forest and the fish migrating into the lake

    use the flooded forest as habitat where they breed, lay eggs and grow. In the dry

    season, the water recedes and fish migrate back to the river moving both upstream and

    downstream. According to Van Zalienge et al., (2000), fish from the Tonle Sap

    migrate as far as Laos and the Mekong Delta. Thus, fish from the Tonle Sap Lake are

    distributed all over the Mekong and because of this the Mekong River is high in fish

    biodiversity and the Tonle Sap is the main source of fish for the Region.

    According to Rainboth (1996), some 1,200 fish species have been identified18

    in the Mekong River Basin (Rainboth, 1996). However, Baran (2005) suggests that

    the fish species in the Mekong River Basin ranges between 758 and 1500 species. Of

    these, 500 species are reported to have resided in the freshwater bodies in Cambodia

    (Van Zalinge and Thouk, 1998). Among these species, about 280 fish species reside

    in the Tonle Sap Lake (Wright et al., 2004).

    These fish species classify into two main groups19

    black fish and white fish.

    Black fish in the Tonle Sap spend most time on the floodplain whereas white fish

    spend most of their lives in the main river channels. Many of white species

    including species of catfish and river carps spend the dry season in the deep pools

    upstream of the main floodplain areas, for examples in the Mekong and large

    18 According to the latest figures, the Mekong is now no.2 river for fish biodiversity with 781 sp.

    Identified, the Amazon has 1217 sp. (Baran ref) Rainboth's figures come from an extrapolation of the

    potential number of fish based on the basin area. The Tonle Sap has 197 sp. (both fresh and saltwater

    sp) Making it the 4th most diverse lake, after the 3 Great Lakes of the African Rift lakes.19 According to some research, they have a third category called grey fish, these are sp with

    intermediate migrations that migrate from floodplains to Mekong tributaries, such as between the TonleSap and its local rivers

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    tributaries in Kratie and Stung Treng Provinces. In the wet season, the adult fish

    species spawn, so million of fish eggs and fry are carried downstream by currents and

    swept into the floodplain areas that are being inundated , where nutrient conditions are

    ideal for rapid juvenile growth (Deap et al., 2003).

    Fish numbers and size increase rapidly on flooded areas, but when water

    levels begin to fall, fish are forced to migrate to deeper water. White fish move back

    to river channels and the Tonle Sap Lake, and blackfish moves to floodplain water

    bodies. Many white fish then migrate down the Tonle Sap River to the Mekong River,

    and then most move upstream and stay in the deep pools for the dry season (Deap et

    al., 2003). Among the migratory species, about 23 species migrate down to the

    Mekong Delta (Thanh Be et al., 2007). Thus, the Tonle Sap is a key breeding and

    production in the Mekong River. This production supports millions of people in the

    Lower Mekong Countries.

    The Impacts of Hydropower on Fisheries and Livelihoods in the Tonle Sap

    The Tonle Sap is under pressure. One of these pressures comes from increased

    development of hydropower projects in the Mekong River Basin. Since the 1950s,

    large and small dams have been built in the Lower Mekong (FACT & EJF, 2001). By

    2007, 82 major hydropower dams are in operation in the six countries in the Mekong

    Region (See Table 3). Between 1965 and 2005, 23 major hydropower dams were built

    in the Mekong River Basin, including mainstream dams in China (Manwan dam and

    Dachaoshan dams), and some hydropower dams on the tributaries in the Lower

    Mekong Region (CNMC & NEDECO, 1998; MRC, 2005).

    Table 3. Major Hydropower Dams in the Mekong River BasinExisting large Project

    (operating and Under Construction)

    Potential Large Project

    (Committed and Planned and Identified)

    Number MW Number MWCambodia 2 222 33 8,009

    Laos PDR 11 1779 32 5788

    Vietnam 30 5,910 65 11,160

    Thailand 11 744 0 0

    Myanmar 21 1,506 15 7,852

    China 5 21,150 34 83,360

    Total 82 31,311 149 116,170

    Source: Peter King, Jeremy Bird, Lawrence Haas, 200720

    20Peter King, Jeremy Bird, Lawrence Haas. 2007. The Current Status of Environmental Criteria for

    Hydropower Development in the Mekong Region: A Literature Compilation.

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    The dams in the Mekong River Basin since 1965 have an estimated accumulative

    active storage of 15,328 million cubic meters (mcm) (See Table.4). The long term

    trend of discharge of the Mekong River suggests a decrease in discharge of around

    10-12% since the 1960s (CNMC & NEDECO, 1998; CNMC, 2004).

    Table 4. Major water resource development projects in the Mekong basin

    Year No. of

    Projects

    Power characteristic Irrigation potential (ha) Active storage

    (mcm)MW GWh/year Wet season Dry season

    1965-1975 9 257 1,266 209000 189000 10012

    1975-1995 6 1681 8330 53000 35000 1058

    1996-2005 8 3240 17597 0 0 4148

    Grant Total

    1965-2005 23 5178 27,193 275,000 227,000 15,328

    Source: CNMC & NEDECO, 1998; CNMC, 2004

    Dams have two main impacts on the Tonle Sap; first, the dam and weir

    construction for hydroelectric power results in an increased water level in the dry

    season, increased turbidity and nutrient reduction for fish (Kummu et al., 2008);

    Second, the dams will contribute to the reduction of the wet season flow.

    Due to the considerable variety and ambiguity of different development plans

    for the Mekong River Basin, there is a concern about the impacts on the Tonle Sap.

    Kummu et al., (2008) assess the impacts of flow alteration on the Tonle Sap Lake.

    This assessment is based on the existing cumulative impact 2004 assessment (CIA)

    conducted by the MRC with support from World Bank, the ADB; the wider CIA

    within the Nam Thuon 2 environmental impact assessment study, and Adamson

    (2001) analyzing the downstream hydrological impacts of the Chinese cascade of

    dams (See Kummu et al., 2008 for these references). Under the CIA of MRC, Kummu

    et al., (2008) selected the high development scenario in which wet-season water level

    would be reduced by 0.36m and the dry season water level would be increased by

    0.15m; whereas under the ADB CIA, the 20 years scenario is selected, in which wet-

    season water levels would be reduced by 0.54m and the dry-season water level would

    be increased by 0.60m. However, under the CIA conducted by Adamson, the scenario

    of 20 percent regulation would result in a dry season water level increased by 0.30m.

    Based on these CIAs, Kummu et al., (2008) estimate that the lake areas

    corresponding to water level of 1.44m amsl is 2,300km. Rises of 0.15m, 0.3m, and

    0.6, representing each analyzed CIA, would result in a permanent lake area of

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    2,700km, 3000km, and 3,200km respectively. Thus, the permanent lake area would

    increase between 400 and 1,000km (17%-40%).

    Kummu et al., (2008) further illustrates that the predicted dry-season water-

    level rise of 0.15-0.60m would mean permanent inundation of large areas of flooded

    forest located around the lake. This suggests that if the dry season water level would

    rise 0.6m, 80.4 km2

    of the total area of the flooded forest (197.2 km2) will be

    inundated. That would mean that 41% of the present flooded forest area would be lost.

    With a dry-season water level rise by 0.15m, 0.30m or 0.60m, about 6%, 31% or 83%

    respectively of the total area of 149km of the Ramsar core site would be inundated.

    The reduced wet season water flow could cause less water to inundate the

    flooded forest and floodplain thus reducing lake fish productivity (Sopha and Van

    Zalienge, 2000). Kummu et al., (2008) estimate that due to hydropower dams,

    floodplain area would decrease between 7% and 16% (Kummu et al., 2008). This

    would have direct impact on the ecosystem productivity with a smaller area resulting

    is less nutrient-energy transfer between terrrestrial and aquatic systems, lowering

    productivity (Kummu et al., 2008; Sopha and Van Zalienge, 2000).

    The changes in dry season water flows from the upstream dams will alter a

    major ecological trigger of fish migration. Dams will physically block fish migration,

    and change water temperatures. Turbidity associated with dam may also negatively

    affect the fisheries (Kummu et al., 2008). The increased risk of extreme flood events,

    either by dam disaster or through a sudden release of water would seriously impact

    fishing communities. At present there are no plans to coordinate water release from

    various planned and existing dams in the region (Baker, 1999).

    Food Insecurity for Fishers in the Tonle Sap

    About 80.5 % of the 2008 Cambodian population of 13.4 million is rural with

    a heavy dependence on natural resources. About 10-11 percent of the total population

    lives in the Tonle Sap floodplain21 (NIS, 200822; CNMC & Nedeco, 1998; Keskinen,

    2003).

    In 2008, about 1,388,555 people lived in the Tonle Saps floodplain (between

    Highway no.5 and no.6) (See Table 5). Of these, about 892,722 people live in floating

    21

    about 1.18-1.20 million people live in the Tonle Sap floodplain (CNMC & Nedeco, 1998; Keskinen,2003)22 http://www.nis.gov.kh/index.php/statistics/surveys/census2008/provisional-population-totals

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    and stand-stilt villages in the Tonle Sap. This suggests that abut 64 percent of the

    Tonle Sap population is engaged in fishing as a primary occupation. Furthermore, at

    the national level, about 10 percent of Cambodian population is engaged in fishing in

    the Tonle Sap (See Table 5).

    Table 5. Fishing population in the Tonle Sap and national population

    Type of population

    No. of

    Population

    Percentage of fishing population against the Tonle

    Sap population an national population (%)

    Fishing population in the Tonle

    Sap 892722a

    Population of the Tonle Sap 1,388,555a 64.29

    National Population 13,388,910b 10.37

    Source: a) Field Notes, 2007 and 2008; b) NIS, 200823.

    For generations, the Tonle Sap has been a primary source of food for

    Cambodians. Many people have depended on the Lake as a space of dependence.

    Access to the Lakes resources provides a safety-net and food security for the

    Cambodian people. Fish consumption24

    per capita is estimated at abut 54.2 kg/person

    (Thouk, 2009).

    The impact of hydropower dams on fisheries can lead to increased food

    insecurity for Cambodians, especially fishers in the Tonle Sap. Household fish catch

    has declined from 10kg/day to 5kg/day. The trend shows further future decrease due

    to an increasing fishing population.

    Conclusion

    Upstream Mekong development threatens the Tonle Saps fisheries and

    livelihoods. These effects of hydropower development on the Tonle Sap will extend

    to other parts of the Mekong as Tonle Sap is a heart of the Mekong. We have

    shown how through the flood pulse, the Tonle Sap absorbs the waterthe blood of

    the Mekonginto the heart and then it releases it into the Mekong. If the heart

    stops, the system dies (Poulsen, cited in Nikula, 2004).

    23 http://www.nis.gov.kh/index.php/statistics/surveys/census2008/provisional-population-totals24 The Average Cambodian fish consumption is between 30-35 kg/yr which is the highest FW fish

    consumption in the world. With Cambodia getting 49.9 % of its protein from fw fish; the world averageis 5.9% (Baran, E. WFC MRC SEA Baseline Assessment, Phnom Penh, Jan, 27, 2010)

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