*iÃÌÊ* > Ì Û>Ã Ûi iÃÃ ÃÃiÃÃ i Ì - VRO
Transcript of *iÃÌÊ* > Ì Û>Ã Ûi iÃÃ ÃÃiÃÃ i Ì - VRO
AuthorsJohn E.R. Weiss B. Sc. (Hons.)Linda J. Iaconis B. App. Sc.
Contributing StaffJim Backholer B. Agr. Sc. (Hons.)Les BouldCarmen Sporle
� The State of Victoria, Department of Natural Resources and Environment, 2002.
DisclaimerThe recommendations in this document are intended only to assist the reader.
The information contained in this publication is based on knowledge available, andunderstanding at the time of writing. However, due to continual advances inknowledge, readers are reminded of the need to ensure that the information they relyupon is current.
This publication may be of assistance to you but the staff at the Keith Turnbull ResearchInstitute, Department of Natural Resources and Environment, does not guarantee thatthe publication is without flaw of any kind or is wholly appropriate for your particularpurposes and therefore disclaims all liability for any error, loss or other consequencewhich may arise from you relying on any information in this publication.
Pest Plant Invasiveness Assessment
Prepared by the Keith Turnbull Research Institute, 2002. 2
Contents
Contents .............................................................................................................................2
List of Figures....................................................................................................................3
List of Tables .....................................................................................................................3
Acronyms...........................................................................................................................4
1. Project Background ........................................................................................ 51.1 Weed Management Within Parks Victoria .........................................................51.2 The Pest Plant Assessment System ....................................................................51.3 Project Objectives................................................................................................6
2. Method, Results and Discussion of thePest Plant Invasiveness Assessment ............................................................. 7
2.1 Invasiveness Potential of Pest Plants...................................................................7 2.1.1 Introduction .......................................................................................................7 2.1.2 Invasiveness Criteria .........................................................................................7 2.1.3 Results and Discussion of the Invasiveness Index ..........................................122.2 Present and Potential Distribution of Pest Plants ..............................................15 2.2.1 Introduction .....................................................................................................15 2.2.2 Present Distribution.........................................................................................16 2.2.3 Potential Distribution ......................................................................................17 2.2.4 Results and Discussion of the Present and Potential Distribution
Maps ..................................................................................................................20
3. References ..................................................................................................... 23
Appendix 1: Invasive Potential Criteria and Intensity Ratings ......................................24
Appendix 2: Table of Risk Ratings, National Status and InvasivenessIndices......................................................................................................29
Appendix 3: Examples of Present and Potential Distribution Maps ..............................35
Pest Plant Invasiveness Assessment
Prepared by the Keith Turnbull Research Institute, 2002. 3
List of Figures
Figure 1: Group weightings of invasiveness. ...................................................................9
Figure 2: Criterion weightings of invasiveness. ...............................................................9
Figure 3: Individual criterion total weightings expressed in groups ..............................10
Figure 4: Frequency graph of invasiveness indices........................................................13
Figure 5: Distribution of Spartina anglica in the United States of America.....................16
Figure 6: Distribution of Spartina anglica (left) and S. x. townsendii(right) in Victoria. ...................................................................................17
Figure 7: Potential distribution of Spartina anglica (left) andS. x. townsendii (right) in Australia, according to climatic parameters. .....17
Figure 8: Potential distribution of Spartina anglica (left) andS. x. townsendii (right) in Victoria, according to climatic parameters. ....... .18
Figure 9: Dialogue box from CLIMATE® showing the climatic parameters used interrestrial weed modeling. The eight rainfall parameters are not includedwhen modeling the potential climatic range of aquatic weeds .....................18
Figure 10: Potential distribution of Spartina anglica (left) andS. x. townsendii (right) in Victoria, according to climatic parameters,susceptible land uses and BVTs. ..............................................................19
Figure 11: Potential distribution of Spartina anglica and S. x. townsendiiin Eastern Victoria, according to climatic parameters, susceptible landuses and BVTs. ........................................................................................19
Figure 12: Invasion graph indicating stages of expansion of a newspecies into a habitat. ..............................................................................21
Figure 13: Total cost of plant invasions. Costs of early expenditure(Area A) and the resulting benefit (Area B). ..........................................22
List of Tables
Table 1: Group and criteria weightings for determining invasive potential ........................... 8
Table 2: Summary of the biological data used to determine the intensityrankings for each criterion and the overall score ofSpartina anglica and S. x. townsendii. .......................................................11
Table 3: Assessed species, grouped according to their invasiveness indices ....................14
Pest Plant Invasiveness Assessment
Prepared by the Keith Turnbull Research Institute, 2002. 4
Acronyms
AHP Analytical Hierarchal ProcessARI Arthur Rylah InstituteBVTs Broad Vegetation TypesCMA Catchment Management AuthorityEVCs Ecological Vegetation ClassesFIS Flora Information System (Viridans 1999)GIS Geographic Information SystemKTRI Keith Turnbull Research InstituteIPMS Integrated Pest Management System (previously known as PMIS)PMIS Pest Management Information SystemPV Parks Victoria
Pest Plant Invasiveness Assessment
Prepared by the Keith Turnbull Research Institute, 2002. 5
1. Project Background
1.1 Weed Management Within Parks VictoriaParks Victoria (PV) is responsible for the management of 16 percent of the State andhas a critical role as the custodian of natural values and for the development of highquality and innovative environmental management in the State’s parks and reserves.PV is required to conduct its management to achieve the objectives of a range oftreaties, conventions, Acts, regulations and policies, including:
� ensuring that threats to all indigenous flora and fauna occurring in the parks andreserves system are managed, in accordance with the Flora and Fauna GuaranteeAct (1988); and
� the eradication or control of exotic flora and fauna in the parks and reserves is inline with the National Parks Act (1975) and the Catchment and Land Protection Act(1994).
In 2000/2001, PV’s Environmental Management Program included 402 pest plantmanagement projects across the State accounting for 28 percent of PV’s natural valuesmanagement budget. To ensure PV directs its resources to weed species of highest riskpotential to particular environmental values, an assessment of the invasive potential ofpest plant species is required.
1.2 The Pest Plant Assessment SystemIn Victoria, there are over 1200 plant species reported in literature as weeds(Ross 2000). It has been estimated that only approximately ten percent of naturalisedplant species become weeds of significant economic and ecological impact (Williamsonand Fitter 1995). It is therefore unrealistic and unnecessary to expect that all weeds canand should be controlled.
To make informed decisions about the best way to control weeds on public land, it isnecessary that the relative importance and potential impact of each weed be determined.It is essential that this is done prior to the allocation of priority works or funding.Decisions based on limited factual data and emotional reactions, will almost certainlyresult in unnecessary expenditure of resources and damage to the environment throughinappropriate use of control measures.
The Pest Plant Assessment System, developed for Victoria by the KTRI, is a riskassessment process that allows for the prioritisation of weeds based on a combinationof the following factors:
(1) Assessing the plant’s invasiveness;
(2) Comparing the plant’s present and potential distribution; and
(3) Determining the impacts of the plant on social, environmental and agriculturalvalues.
Pest Plant Invasiveness Assessment
Prepared by the Keith Turnbull Research Institute, 2002. 6
1.3 Project ObjectivesThe objective of this project is to determine the invasive and current and potentialdistribution of 112 pest plant species using the method already established by KTRI(factors (1) and (2) above). The species were selected by PV on the basis of:
� risk rating for environmental weeds (Carr et al 1992);
� pest plant species new to a district, park or area of the park;
� number of hectares per pest plant species proposed to be controlled in2001/02; and
� number of parks in which each pest plant species occurs.The selected species are listed alphabetically, with some updates to the nomenclatureused in the report, at the start of Appendix 3.
This report documents:
1. The method, results and discussion of the invasiveness score and current andpotential distribution for the 112 environmental weeds selected by PV (Section 2);
2. Tabulated data for the 112 weed species including environmental risk rating(according to Carr et al 1992), national status, and invasiveness score (Appendix 2);and
3. Present and potential distribution maps for the 112 weeds (Appendix 3), which havealso been provided in digital format.
The data compiled in this report on the invasive potential of pest plants will assist withthe development of a pest plant risk assessment method. This method will be consistentwith Victoria’s Pest Plant Assessment System, incorporating additional measures ofimpact on environmental, social and economic values, and is being developed under aseparate, but related project.
Pest Plant Invasiveness Assessment
Prepared by the Keith Turnbull Research Institute, 2002. 7
2. Method, Results and Discussion of the Pest PlantInvasiveness Assessment
2.1 Invasiveness Potential of Pest Plants2.1.1 IntroductionMany researchers have focused on the relative invasiveness of species as an indicator ofpotential spread rate. Invasiveness can be defined as the ability to establish, reproduce,and disperse within an ecosystem. Plant propagules arrive at a new site with certaininherent characteristics that previously enabled their successful survival and continuedreproduction throughout their evolutionary history. There is no single suite ofcharacteristics which make a plant invasive, rather there are several predisposing factorsthat act either alone or together to increase the chance of a plant becoming invasive.
Many researchers have also agreed that the following biological attributes of a plantspecies are associated with invasiveness.
� Ecological status; a generalist or specialist plant.Most common and noxious weeds in southern Australia are generalist andopportunistic rather than requiring specific niches or special habitat requirements.
� ‘Weedy’ phenology and biology; such as competitive growth, seed dispersalmechanisms, seed dormancy and propagule production.Major weeds can have attributes such as high seed production, rapid vegetativespread, long-lived seeds, staggered germination, competitive growth and long-distance seed dispersal. However, there is no defined group of ecological andbiological attributes that can be used to identify all major weeds. Differentattributes may be important for different plant families and different ecosystems.
� Wide native range.Within a genus the more important weeds may have a wider native range.
� Taxonomic position; members of generally ‘weedy’ plant families.Certain plant families such as Poaceae (grasses), Asteraceae (eg. daisies, thistles),Iridaceae (irises) and Brassicaceae (eg. mustards, turnips) are noted for havingmany ‘weedy’ species.
� Effective modes of reproduction and genetic variation.Plant species that vegetatively reproduce or self-pollinate have the potential tostart new populations from a single, isolated plant. However, high levels ofinbreeding in self-pollinators may limit their adaptability compared to cross-pollinators.
2.1.2 Invasiveness CriteriaCriteria for a generic model to assess the potential invasiveness of weeds weredetermined at workshops by national participants at the Arthur Rylah Institute (ARI),in June 1998 (Table 1). A working group at the Keith Turnbull Research Institute(KTRI) then used an expert system, relying on multi-criteria analysis/analyticalhierarchical process (AHP) (Saaty 1995), to develop a decision tree that allows forgroups and criteria to be weighted according to importance (Table 1). Basically, theAHP is a method of breaking down a complex unstructured situation into its componentparts; arranging these parts into a hierarchical order; and assigning numerical values tosubjective judgements to determine which variables have the highest priority and shouldbe acted upon to influence the outcome of the situation. AHP also facilitates effective
Pest Plant Invasiveness Assessment
Prepared by the Keith Turnbull Research Institute, 2002. 8
decisions on complex issues by simplifying and expending the intuitive decision makingprocess.
Table 1: Group and criteria weightings for determining invasive potential.
GROUP CRITERIA GROUP & CRITERIAWEIGHTINGS
Establishment 0.5Germination requirements? 0.085Establishment requirements? 0.671Disturbance requirements? 0.244
Growth/competitive ability 0.096Life form? 0.06Allelopathic properties? 0.09Tolerates herbivory pressure? 0.472Normal growth rate? 0.192Stress tolerances? 0.185
Reproduction 0.119Reproductive system? 0.047Propagule production? 0.46Seed longevity? 0.256Reproductive period? 0.101Time to reproductive maturity? 0.136
Dispersal 0.284Number of mechanisms? 0.333How far do propagules disperse? 0.667
Comparing the major groups (i.e. establishment, growth/competitive ability,reproduction and dispersal), the working group indicated that the ability of a plant toestablish in an ecosystem was by far the most important indicator of its invasiveness,followed by the plant’s ability to disperse, then its reproduction strategy, and finally itsgrowth/competitive ability. Some of the reasons (and comments made during theweighting process) for this ranking are mentioned below:� “It doesn’t matter how many seeds/propagules the plant produces, if they can not
establish themselves they are not going to be invasive.”� “The ability to disperse great distances, no matter how they reproduce, gives the
plant a better chance of finding a suitable location for it to establish.”� “The more locations it establishes in, (i.e. multi loci), the quicker the plant will
invade an area.”
These group weightings can be expressed graphically as shown below in Figure 1.
Pest Plant Invasiveness Assessment
Prepared by the Keith Turnbull Research Institute, 2002. 9
Figure 1: Group weightings of invasiveness.
Within each group, the individual criterion (Appendix 1) were compared and weightedagainst each other. For instance, within the dispersal group, the working group decidedthat the distance propagules disperse was twice as important as the mechanisms fordispersal. The results of the intra-group criteria weightings are illustrated in Figure 2.
Figure 2: Criterion weightings of invasiveness.
Within the invasiveness hierarchy, the weightings of individual criterion are multipliedby the groups weight (eg. distance of dispersal x dispersal � 0.667 x 0.284 = 0.189).The total weightings for individual criterion are illustrated in Figure 3.
0 0 . 1 0 . 2 0 . 3 0 . 4 0 . 5 0 . 6
1
GR
OU
PS
G R O U P W E I G H T I N G S
D I S P E R S A L
R E P R O D U C T I O N
G R O W T H / C O M P E T I T I V E
E S T A B L I S H M E N T
0 .0 00 0.10 0 0.200 0 .3 00 0 .4 00 0.50 0 0.600 0.7 00 0 .8 00
Gro
ups
W e igh t
D is ta nce o f d ispe rsa lN u m be r o f d isp ersa l m echa n ism s
R e pro duc tive ag erep rodu c tive p eriodS ee d lon ge vityN u m be r o f p ro pag u lesR e pro duc tive m o de
S tre ss to le ran ceN o rm al g row th ra teTo lera te s h erb ivoryA lle lo pa th ic p rope rtiesL ife fo rm
H o w m uch d is turb an ceE s ta b lishm e nt re qu irem e ntsG erm inatio n requ ire m en ts
D is pe rs al
R ep ro d u c tio n
G ro w th &C o m pe titive Ab ility
E s ta b lis h m en t
Pest Plant Invasiveness Assessment
Prepared by the Keith Turnbull Research Institute, 2002. 10
Figure 3: Individual criterion total weightings expressed in groups.
The invasive hierarchy and weightings indicate that the most important invasivebiological features of a plant is its’ ability to establish and disperse widely. The plant’sability to withstand herbivore pressure, number of propagules produced and seedlongevity are of lesser importance. Other biological features such as its life formand reproductive strategies are of much less importance. This was acknowledgedduring the workshops at ARI, where serious weeds from a wide range of life forms(grasses, shrubs, trees, climbers etc) and reproductive systems (vegetative, asexual,self pollinating, cross pollinating etc) were identified.
The criteria are assigned intensity ratings or definitions of ‘high’, ‘medium-high’,‘medium’, ‘medium-low’ and ‘low’ (Appendix 1), which are used to score each species.The scores for each criterion and weightings are then tallied and calculated to producea final ‘invasiveness index’ for each species. An example of this process is shownbelow; a summary of biological data was collated to determine the ‘invasiveness index’of Spartina anglica and S. x townsendii (Table 2). The overall score is only relative toscores of other plants run through the same process, but can be used to rank species asto their potential invasiveness or rate of spread.
The information to rate each criterion is sourced from databases, journal articles, flora’sof the world (books or articles describing the species of a particular country or region),online information, and any other sources. There is much available informationon some species (eg. declared noxious species), and very scant information for others(eg. grasses and new and emerging weed species). Where there is an information ‘gap’for a particular criterion, a ‘medium’ (M) ranking is given to indicate ‘unknown’.Although the invasiveness assessments are undertaken using the best availableinformation, they are only as accurate as the information that is used. Therefore, as webecome more informed about a species, reassessment may be necessary.
0 . 0 0 0 0 .1 0 0 0 .2 0 0 0 . 3 0 0 0 .4 0 0 0 .5 0 0 0 .6 0 0 0 . 7 0 0 0 .8 0 0
Gro
ups
W e ig h t
D is ta n c e o f d is p e r s a lN u m b e r o f d is p e r s a l m e c h a n is m s
R e p r o d u c t iv e a g eR e p r o d u c t iv e p e r io dS e e d lo n g e v i t yN u m b e r o f p r o p a g u le sR e p r o d u c t iv e m o d e
S t r e s s t o le r a n c eN o r m a l g r o w th r a teT o le r a te s h e r b iv o r yA l le lo p a t h ic p r o p e r t ie sL i f e f o r m
H o w m u c h d is t u r b a n c eE s t a b l i s h m e n t r e q u i r e m e n tsG e r m in a t io n r e q u i r e m e n ts
D i s p e r s a l
R e p r o d u c t i o n
G r o w t h &C o m p e t i t i v e A b i l i t y
E s t a b l i s h m e n t
Pest
Pla
nt In
vasi
vene
ss A
sses
smen
t
Pre
pare
d by
the
Kei
th T
urnb
ull R
esea
rch
Inst
itute
, 200
2.
11
Tab
le 2
: Su
mm
ary
of th
e bi
olog
ical
dat
a us
ed to
det
erm
ine
the
inte
nsity
rank
ings
for e
ach
crite
rion
and
the
over
all i
nvas
iven
ess
inde
x of
Spar
tina
angl
ica
and
S. x
tow
nsen
dii.
Com
men
ts re
late
to in
tens
ity ra
king
s sho
wn
in A
ppen
dix
1 an
d ap
ply
to b
oth
spec
ies u
nles
s oth
erw
ise
stat
ed.
QU
ESTI
ON
CO
MM
ENTS
INTE
NSI
TY R
AN
KIN
GEs
tabl
ishm
ent
S. a
nglic
aS.
x to
wnse
ndii
Ger
min
atio
nre
quire
men
ts?
Onl
y re
quire
s wat
er (a
n aq
uatic
env
ironm
ent)
for v
eget
ativ
e pr
opag
ules
to g
erm
inat
e.H
H
Esta
blis
hmen
tre
quire
men
ts?
Occ
urs
in ‘
open
’ ar
eas
whe
re l
ight
and
spa
ce a
re n
ot l
imiti
ng (
eg.
saltm
arsh
and
man
grov
eco
mm
uniti
es).
ML
ML
Dis
turb
ance
requ
irem
ents
?A
ble
to in
vade
exi
stin
g sa
ltmar
sh a
nd m
angr
ove
com
mun
ities
to fo
rm e
xten
sive
mea
dow
s.H
H
Gro
wth
/com
petit
ive
abili
tyLi
fe fo
rm?
Aqu
atic
gra
ss.
HH
Alle
lopa
thic
pro
perti
es?
Non
e de
scrib
ed o
r kno
wn.
LL
Tole
rate
s her
bivo
rypr
essu
re?
Pala
tabl
e to
stoc
k. C
ould
pro
duce
veg
etat
ive
prop
agul
es u
nder
mod
erat
e he
rbiv
ory.
MH
MH
Nor
mal
gro
wth
rate
?Fo
rms e
xten
sive
col
onie
s in
a re
lativ
ely
shor
t per
iod
of ti
me
that
exc
lude
s oth
er p
lant
s.H
HSt
ress
tole
ranc
es?
Tole
rant
of s
alin
ity, w
ater
logg
ing
and
fire.
MH
MH
Rep
rodu
ctio
nR
epro
duct
ive
syst
em?
Bot
h sp
ecie
s are
cap
able
of v
eget
ativ
e re
prod
uctio
n, h
owev
er o
nly
S. a
nglic
a is
abl
e to
pro
duce
viab
le se
eds.
HM
H
Prop
agul
e pr
oduc
tion?
S.x
tow
nsen
dii
does
not
pro
duce
see
ds.
S. a
nglic
a is
a p
rolif
ic s
eede
r w
ith i
nflo
resc
ence
sex
ceed
ing
90 p
er sq
uare
met
re in
mat
ure
and
trans
ition
al z
ones
.M
L
Seed
long
evity
?S.
ang
lica
seed
s ge
rmin
ate
with
in th
e ye
ar a
nd n
o se
ed b
ank
exis
ts.
S. x
tow
nsen
dii d
oes
not
prod
uce
seed
s.M
M
Rep
rodu
ctiv
e pe
riod?
Form
s se
lf -s
usta
inin
g ex
tens
ive
mea
dow
s (i.
e. d
ense
mon
ocul
ture
s th
at c
an b
e te
ns o
rhu
ndre
ds o
f hec
tare
s in
area
).H
H
Tim
e to
repr
oduc
tive
mat
urity
?Fo
rms
exte
nsiv
e m
eado
ws
over
a re
lativ
ely
shor
t per
iod,
Fas
t gro
win
g gr
ass
spec
ies.
Rhi
zom
esse
nd u
p ne
w sh
oots
that
qui
ckly
gro
w in
to a
larg
e cl
ump
(ass
umed
to b
e 1-
2 ye
ars)
.H
H
Dis
pers
alN
umbe
r of m
echa
nism
s?D
istri
bute
d w
idel
y by
wat
er-b
orne
dis
pers
al,
seed
s al
so c
arrie
d by
win
d. F
ragm
ents
of
the
rhiz
ome
may
bre
ak lo
ose
and
drift
in c
urre
nts.
MH
MH
How
far d
o th
eydi
sper
se?
Sea
curr
ents
like
ly to
dis
pers
e so
me
prop
agul
es >
1 ki
lom
etre
how
ever
in e
stua
ries
mos
t lik
ely
to b
e di
sper
sed
betw
een
200-
1000
met
res.
MH
MH
INV
ASI
VEN
ESS
IND
EX (M
axim
um=1
, Min
imum
=0)
0.61
0.58
Pest Plant Invasiveness Assessment
Prepared by the Keith Turnbull Research Institute, 2002. 12
2.1.3 Results and Discussion of the Invasiveness IndexThe invasiveness scores, as well as risk rating according to Carr et al (1992) and national status, for all112 species are given in Appendix 2. The closer the invasiveness index to the maximum score of one,the more invasive a species is. To give the score some perspective, a comparison can be made with the‘non-weedy’ plant Red or Chinese Hibiscus (Hibiscus rosa-sinensis). A common plant grownthroughout southern Australia, it scored 0.43 but only because it has such a high chance of dispersal bydeliberate transport of cuttings/seeds greater than one kilometre. When humans were excluded fromdispersal, the invasiveness score dropped to 0.31.Spartina anglica scored 0.61 and S. x townsendii 0.58. In relation to the other assessed weeds, bothSpartina species scored less than the mean invasive index of the 112 species assessed, which was 0.69.S. anglica scored an equivalent rank to plants such as Angled onion (Allium triquetrum), Slender thistle(Carduus tenuiflorus), and Stinkwort (Dittrichia graveolens), while S. x townsendii scored equivalent toArrowhead (Sagittaria graminea). Many species are considerably more invasive than Spartina, such asPampas grass (Cortaderia selloana) with a score of 0.95, White willow (Salix alba) with a score of0.93, and Grey sallow and Crack willow (Salix babylonica and S. cinerea respectively) which bothscored 0.92 (Appendix 2).
As demonstrated above, an expert system, such as the Pest Plant Assessment System, provides aranking of plants from most invasive to least. The invasiveness indices are not meant to be anabsolute number relating to distance, but to give a relative indication of rate of spread of a plant in itspreferred habitat. To make the invasiveness scores more useful, one can group the resulting indicesof the 112 assessed species into categories (eg. ‘very high’, ‘high’, ‘medium’ invasiveness, etc).Such a process can assist land managers to determine which species, or groups of species, to targetmanagement efforts and resources at for maximum gain. There are several methods of breaking thescore range of zero to one to form ‘invasiveness categories’, with some more relevant than others.
One method is to group the species, ordered from lowest to highest invasiveness index, according topercentile rankings. For instance, using 25 percentiles to break the species into four groups (i.e. indicesbetween the 0-25, 25-50, 50-75 and 75-100 percentiles respectively). Such an approach results in aneven number of species in each category, however did not well reflect the range or ‘natural groupings’of indices when applied to the 112 assessed species. As the majority of assessed species have indices inthe mid range (i.e. between the 25-75 percentiles), species at the lower and higher ends of this midrange were dispersed into the lower and high groups (ie. 0-25 and 75-100 percentiles respectively) toallow for an even distribution of the scores. This elevated the cut-off score for the lowest group andreduced the cut-off score for the highest group to accommodate the extra species falling either side ofthis mid group. Furthermore, if subsequent species are assessed and grouped with the 112 species, the25 percentiles are likely to change. Therefore, a method which arbitrarily fixed the ‘cut off’ points isrequired.
A frequency distribution graph of the scores of all 200 species that have been assessed to date byKTRI was produced (Figure 4). Evaluation of the figure shows indications of some natural breaks inthe data. These ‘natural groupings’ produced the five groups used to categorise the 112 speciesconsidered in this report (Table 3): moderately invasive (species with scores <0.5), moderately-highly invasive (species with scores between 0.5-0.59), highly invasive (species with scores between0.6-0.79), very highly invasive (species with scores between 0.8-0.89), and extremely invasive(species with scores >0.9). As expected, few species occur in the moderately and extremely invasivecategories, while the vast majority (73 of the 112 assessed species) occur in the middle highlyinvasive category (Table 3).
Pest
Pla
nt In
vasi
vene
ss A
sses
smen
t
Pre
pare
d by
the
Kei
th T
urnb
ull R
esea
rch
Inst
itute
, 200
2.
13
Figu
re 4
: Fr
eque
ncy
grap
h of
inva
sive
ness
indi
ces.
024681012
0.40.4
20.4
40.4
60.4
80.50.5
20.5
40.5
60.5
8
0.60.6
20.6
40.6
60.6
80.70.7
20.7
40.7
60.7
80.80.8
20.8
40.8
60.8
80.90.9
20.9
40.9
60.9
8
1
Scor
e
Frequency
Pest
Pla
nt In
vasi
vene
ss A
sses
smen
t
Pre
pare
d by
the
Kei
th T
urnb
ull R
esea
rch
Inst
itute
, 200
2.
14
Tab
le 3
: A
sses
sed
spec
ies,
grou
ped
acco
rdin
g to
thei
r inv
asiv
enes
s ind
ices
.M
oder
atel
y In
vasiv
eSc
ore
(<0.
5)M
oder
atel
y/H
ighl
y In
vasiv
eSc
ore
(0.5
-0.5
9)H
ighl
y In
vasiv
eSc
ore
(0.6
-0.7
9)H
ighl
y In
vasiv
eSc
ore
(0.6
-0.7
9)V
ery
Hig
hly
Inva
sive
Scor
e (0
.8-0
.89)
Extr
emel
y In
vasiv
eSc
ore
(>0.
9)C
enta
urea
cal
citr
apa
Asp
hode
lus
fistu
losu
sA
caci
a ba
ileya
naLi
gust
rum
vul
gare
Aila
nthu
s al
tissi
ma
Cor
tade
ria
sello
ana
Mel
iant
hus
com
osus
Car
tham
us la
natu
sA
cer
pseu
dopl
atan
usLy
cium
fero
ciss
imum
Chr
ysan
them
oide
s m
onili
fera
Salix
alb
aN
asse
lla h
yalin
aC
irsi
um a
carn
aA
gros
tis c
apill
aris
Mar
rubi
um v
ulga
reC
oton
east
er g
lauc
ophy
llus
Salix
cin
erea
Scol
ymus
his
pani
cus
Dip
sacu
s fu
llonu
mA
llium
triq
uest
rum
Myr
ioph
yllu
m a
quat
icum
Cop
rosm
a re
pens
Salix
bab
ylon
ica
Sola
num
linn
aean
umD
iplo
taxi
s te
nuifo
liaA
loe
sapo
nari
aM
ysip
hyllu
m a
spar
agoi
des
Dip
ogon
lign
osus
Salix
x r
uben
sV
erba
scum
thap
sus
Em
ex a
ustr
alis
Alte
rnan
ther
a ph
iloxe
roid
esN
asse
lla n
eesi
ana
Hyp
eric
um a
ndro
saem
umW
atso
nia
mer
iana
Foe
nicu
lum
vul
gare
Ant
hoxa
nthu
m o
dora
tum
Nas
sella
tric
hoto
ma
Leyc
este
ria
form
osa
Hyp
eric
um te
trap
teru
mB
riza
max
ima
Opu
ntia
ficu
s-in
dica
Loni
cera
japo
nica
Junc
us a
cutu
sC
ardu
us p
ycno
ceph
alus
Opu
ntia
rob
usta
Rha
mnu
s al
ater
nus
Ono
pord
um a
cant
hium
Car
duus
tenu
iflor
usO
punt
ia s
tric
taSo
llya
hete
roph
ylla
Ono
pord
um a
caul
onC
estr
um p
arqu
iO
xalis
pes
-cap
rae
Ule
x eu
ropa
eus
Res
eda
lute
ola
Cir
sium
arv
ense
Oxy
lobi
um la
nceo
latu
mSa
gitta
ria
gram
inea
Cir
sium
vul
gare
Pas
palu
m d
ilata
tum
Spar
tina
x to
wns
endi
iC
onvo
lvul
us a
rven
sis
Pen
nise
tum
cla
ndes
tinum
Trad
esca
ntia
alb
iflor
aC
rata
egus
mon
ogyn
aP
hala
ris
aqua
tica
Xan
thiu
m s
pino
sum
Cus
cuta
cam
pest
ris
Phy
salis
vis
cosa
Cyn
ara
card
uncu
lus
Pin
us p
inas
ter
Cyt
isus
sco
pari
usP
inus
rad
iata
Dat
ura
stra
mon
ium
Pitt
ospo
rum
und
ulat
umD
elai
rea
odor
ata
Pol
ygal
a m
yrtif
olia
Ditt
rich
ia g
rave
olen
sP
opul
us a
lba
Ech
ium
pla
ntag
ineu
mP
runu
s lu
sita
nica
Ech
ium
vul
gare
Ros
a ru
bigi
nosa
Ehr
hart
a er
ecta
Rub
us fr
utic
osus
agg
.E
ragr
ostis
cur
vula
Salp
ichr
oa o
riga
nifo
liaE
rica
lusi
tani
caSe
neci
o ja
coba
eaG
alen
ia p
ubes
cens
Sene
cio
pter
opho
rus
Gen
ista
lini
folia
Sily
bum
mar
ianu
mG
enis
ta m
onsp
essu
lana
Sola
num
ela
egni
foliu
mH
eder
a he
lixSp
artin
a an
glic
aH
iera
cium
aur
antia
cum
Spor
obol
us in
dicu
sH
omer
ia fl
acci
daTr
ibul
us te
rres
tris
Hyp
eric
um p
erfo
ratu
mV
erba
scum
vir
gatu
mIl
ex a
quifo
lium
Vin
ca m
ajor
Junc
us e
ffusu
sV
ulpi
a br
omoi
des
Lava
ndul
a st
oech
asX
anth
ium
str
umar
ium
Leuc
anth
emum
vul
gare
Pest Plant Invasiveness Assessment
Prepared by the Keith Turnbull Research Institute, 2002. 15
2.2 Present and Potential Distribution of Pest Plants2.2.1 IntroductionCurrent and potential distributions are another major component required in the decisionsupport system and AHP, to predict the status of a weed. The greater the potentialdistribution of a weed, the greater the potential impact and management costs.To ensure the most cost-effective use of resources, invasive species that have thegreatest potential range should be targeted. Prioritisation is also important, as it isunrealistic to expect that all weeds can be controlled with limited available resources.Knowledge of potential distribution is furthermore important for devising managementprograms. Land managers can be alerted to the risk of weed invasion and measures canbe enforced to prevent the introduction of weed propagules into new areas. Lowpriority can be given to areas where the weed might fail to persist, or be of littleeconomic, environmental or social importance.
Two of the major factors influencing weed distribution are climate and land use.Weed species are typically more invasive in regions that are climatically similar to theirnative environment. Climate limits distribution according to how temperature andmoisture stresses affect the weed's life cycle. Different land uses (eg. cropping,perennial pasture and forestry) have different disturbance regimes that favour differentgroups of weeds. Having determined the climatic preferences of a weed it is necessaryto overlay these on a map of the weed’s associated land use in Victoria. The areas ofthe state that are potentially at risk from this weed can then be identified.
Pest Plant Invasiveness Assessment
Prepared by the Keith Turnbull Research Institute, 2002. 16
2.2.2 Present DistributionInformation on the weed’s present distribution, both overseas and in Australia,is collected from databases, journal articles, flora’s of the world (books or articlesdescribing the species of a particular country or region), online information, and any othersources. Spartina is used herein as an example to highlight the variety of sources andprocess used to determine a weed’s present. The information was sourced from books,conference proceedings, online databases, and a CD database.
Spartina anglica and S. x townsendii were first collected at Lymington Hampshire,United Kingdom in 1892, and then in the Isle of Wight in 1893 (Hubbard 1974). By 1900it occurred in scattered patches from Chichester Harbour to Poole Harbour (Hubbard1974). By 1907 many thousands of tidal mud-flats from Sussex to East Dorset wereinfested (Hubbard 1974). It also appeared on the north coast of France (Hubbard 1974),and has spread through marshes in Germany, Denmark, the Netherlands, New Zealand,China, and the west coast states of the United States of America (Figure 4)(Wu et al 1999).
Figure 5: Distribution of Spartina anglica in the United States of America.(From USDA Plant Database http://plants.usda.gov/plantproj/plants/index.html).
Spartina is widely distributed in New Zealand, with S. anglica occurring from latitudes35025'S to near 470S (Shaw and Gosling 1996).
In Australia, Spartina appears to be restricted to Tasmania, Victoria and South Australia(Bridgewater 1996). In Tasmania it is widespread in estuaries, river mouths andespecially along the north coast (Wells 1996). The worst affected areas of Tasmania arePort Sorell and the Tamar River (Wells 1996). In Victoria, Spartina is found along theGippsland coast in Corner Inlet, Waratah Bay and Andersons Inlet, on the eastern side ofWesternport Bay and near the heads of Port Phillip Bay (Williamson 1996) (Figure 5).
Pest Plant Invasiveness Assessment
Prepared by the Keith Turnbull Research Institute, 2002. 17
Figure 6: Distribution of Spartina anglica (left) and S. x townsendii (right) in Victoria.(Shaded squares indicate quadrats in which one or more plants of a particular specieshave been recorded. The species does not necessarily occupy the entire shaded area).(From Wild Plants of Victoria, Viridans).
2.2.3 Potential DistributionInformation on Australian and overseas distributions were imported into a climatematching program, CLIMATE®, to predict potential distribution in Australia. Using thelocalities where a species occurs overseas and within Australia, the potential climaticrange of any species can be overlaid upon Australia's climatic regions. The maps belowillustrate the climatic regions most suitable for Spartina in Australia and Victoria(Figures 6 and 7).
Figure 7: Potential distribution of Spartina anglica (left) and S. x townsendii (right)and in Australia, according to climatic parameters. (Areas in red indicate a 80%+ matchwith the preferred climate of the plant species, dark green 70%, light green 60% andyellow 50%).
Pest Plant Invasiveness Assessment
Prepared by the Keith Turnbull Research Institute, 2002. 18
Figure 8: Potential distribution of Spartina anglica (left) and S. x townsendii (right) inVictoria, according to climatic parameters. (Areas in red indicate a 80%+ match withthe preferred climate of the plant species, yellow 70%, orange 60% and green 50%).
The 16 climatic parameters that are used to determine potential distribution can begrouped into temperature or rainfall parameters (Figure 8).
Aquatic weeds are modeled for potential climatic range slightly differently thanterrestrial species. Rainfall is obviously not a major criterion for determining thepotential range of aquatic species, especially submergents, although it may play a keyrole in triggering certain biological properties (eg. freshwater flood events appear tostimulate flowering in Spartina) (Strong pers. comm.). Thus rainfall parameters areexcluded when predicting the climatic range. Water temperature is generally moremoderate and has fewer fluctuations than air temperature and provides a more accurateprediction for modeling aquatic species. However, the required data is usuallyunknown. Therefore, modeling the climatic range of aquatic species has included eightair temperature parameters that provide at least some indication of potential range. Theprocess is, consequently, more uncertain and likely to overestimate the species’ actualpotential range.
Figure 9: Dialogue box from CLIMATE® showing the climatic parameters used interrestrial weed modeling. The eight rainfall parameters are not included whenmodeling the potential climatic range of aquatic weeds.
Pest Plant Invasiveness Assessment
Prepared by the Keith Turnbull Research Institute, 2002. 19
The climatic overlays are then used to determine the potential range of the plant speciesby linking or intersecting them with susceptible land uses and broad vegetation types(BVTs) or wetlands using the ArcView Geographic Information System (GIS) program.This refines the potential distribution maps produced using the climate matchingprogram, as plants are limited by other factors, such as disturbance regimes associatedwith land uses. The resulting maps (Figures 9 and 10) illustrate the potential range ofSpartina in Victoria.
Figure 10: Potential distribution of Spartina anglica (left) and S. x townsendii (right)in Victoria, according to climatic parameters, susceptible land uses and BVTs.
Areas in red indicate a very high probability that Spartina could establish inwatercourses and wetlands within this region, yellow a high, and orange a mediumprobability of establishment.
Figure 11: Potential distribution of Spartina anglica and S. x townsendii in EasternVictoria, according to climatic parameters, susceptible land uses and BVTs.
Pest Plant Invasiveness Assessment
Prepared by the Keith Turnbull Research Institute, 2002. 20
The potential distribution maps are estimates and are only as reliable as the data they arebased on. As more records are collected on where the plants occur, the predictions willbecome more accurate. It is expected, consequently, that there are potential distributionmaps that do not yet fully represent existing or potential distribution. Also, for somespecies there may be insufficient data to undertake potential distribution mapping.For instance, only a small number of current distribution records were found forSalix x rubens; too few to produce a meaningful potential distribution map. For thesespecies, a provisional coarse climate map is produced but the next stage of matchingonto susceptible land types is not completed.
2.2.4 Results and Discussion of the Present and Potential Distribution MapsPresent and potential distribution maps for each of the 112 species assessed are given inAppendix 3. These have also been provided to PV in digital format. As 74 of thespecies had already been assessed by KTRI, these maps include CatchmentManagement Authority (CMA) boundaries. The 38 species assessed as part of thisproject have been mapped using PV district boundaries.
The present distribution of weeds is generally underrepresented in the databasesused (i.e. herbarium records, Wild Plants of Victoria (Viridans 1999), and to a lesserextent IPMS/PMIS), with the exception of priority weeds such as Serrated tussock(Nassella trichotoma). Conversely, the modeled potential distribution of weeds is likelyto be overestimated. This occurs as the broad scale (i.e. 1:250,000) of the statewidedatabases used, merges minor differences into the larger BVTs or land uses for eachgrid. Microhabitats within a vegetation or land use type maybe unsuitable for theparticular weed species, and microhabitats outside the identified susceptible land use orvegetation type may be suitable but not recognised (eg. roadsides, small riparian orvegetation corridors). More detailed map layers, such as the soon to be introducedEcological Vegetation Classes (EVCs), will produce better quality predictions.
The many weeds recorded as occurring along roadsides presents another majorlimitation when predicting potential distribution. Victoria has over 170,000 kilometresof roads, however to include all these roads within the image would not be suitable, as itwould be too cluttered and meaningless. Thus, some potential distribution images maynot include the occurrence of weeds within a region, if they only occur along roadsides.For example, Horehound (Marrubium vulgare) can occur along roadsides withincropping regions, but is unable to withstand cultivation. Similarly, some riparian weedsmay occur along small rivers, streams and water channels, but are not included inthe riparian or riverine vegetation classes of the BVT GIS layer, as they are too smallor scattered to be detected, and so don’t appear on the predicted potentialdistribution maps.
The above limitations highlight the need for suitable actions to be undertaken.Where information on a weeds present distribution is known but not recorded, recordsneed to be updated to ensure management and monitoring are effectively undertaken.An accurate comparison of a weeds present distribution with its potential distributionallows managers to make decisions on the course of actions to take. The ratio of presentand potential distribution provides an indication as to what stage the weed is at.Another way of expressing this is the relative position of the species on its invasiongraph (Figure 11).
Pest Plant Invasiveness Assessment
Prepared by the Keith Turnbull Research Institute, 2002. 21
Figure 12: Invasion graph indicating stages of expansion of a new species into ahabitat. (Adapted from Groves (1992) and Hobbs (1991)).
Weeds that have reached or nearly reached the full limits of their distribution, are not amajor concern in terms of potential spread and impacts. Whereas, weeds currentlyoccupying a small area of their potential range, or in the ‘lag or sleeper’ phase, shouldbecome a management priority. Indeed, a powerful weapon against weed invasionsarising from existing infestations is early intervention (Figure 12). Early interventionnot only achieves better in government/land manager investment, but also reduces costsof control and impact on the triple bottom line (social, environmental and agriculturalvalues).
Pest Plant Invasiveness Assessment
Prepared by the Keith Turnbull Research Institute, 2002. 22
Figure 13: Total cost of plant invasions. Costs of early expenditure (Area A) and theresulting benefit (Area B) (Adapted from Hobbs and Humphries (1995)).
Two parts of the Pest Plant Invasiveness Assessment process have been finalised, theinvasiveness assessments and the current and potential distributions within Victoria.The results of this project will provide a valuable tool for determining the risks that pestplants pose to environmental values and for prioritising weed management activitieswithin PV.
Time
Early
Late
A
B
Intervention
Pest Plant Invasiveness Assessment
Prepared by the Keith Turnbull Research Institute, 2002. 23
3. References:Bridgewater, P. 1996. Opening Address. In: Proceedings of the Australasian Conference
on Spartina Control. Department of Conservation and Natural Resources, Victoria.
Carr, G.W., Yugovic, J.V. and Robinson, K.E. 1992. Environmental Weed Invasions inVictoria: Conservation and Management Implications. Department of Conservation andNatural Resources and Ecological Horticulture, Victoria.
Groves, R.H. 1992. Weed ecology and spread. In: Proceedings of the First InternationalWeed Control Congress (Volume 1). Weed Science Society of Victoria, Melbourne.
Hobbs, R.J. 1991. Disturbance a precursor to weed invasion in native vegetation. PlantProtection Quarterly 6(3): 99-104.
Hobbs, R.J. and Humphries, S.E. 1995. An Integrated Approach to the Ecology andManagement of Plant Invasions. Conservation Biology 9(4): 761-770.
Hubbard, C.E. 1974. Grasses: A Guide to Their Structure, Identification, Uses, andDistribution in the British Isles (Revised Edition). Penguin Books, United Kingdom.
Ross, J.H. 2000. A Census of the Vascular Plants of Victoria. Royal Botanic Gardens,Victoria.
Saaty, T. 1995. Decision Making for Leaders: The Analytical Hierarchy Process forDecisions in a Complex World. RWS Publications, Pittsburgh.
Shaw, W.B. and Gosling, D.S. 1996. Spartina Control in New Zealand: An Overview. In:Proceedings of the Australasian Conference on Spartina Control. Department ofConservation and Natural Resources, Victoria.
Strong, D.R. 2002. Professor, Evolutionary and Ecology Department. University ofCalifornia, Davis, United States of America.
USDA PLANTS Profile (Online Database). 2000. Spartina anglica C.E. Hubbard. USDANatural Resources Conservation Service. Available: http://plants.usda.gov/plants/index.html(31 May 2000).
Viridans. 1999. Wild Plants of Victoria. Viridans Biological Databases, Victoria.
Wells, A. 1996. Rice Grass in Tasmania: An Overview. In: Proceedings of theAustralasian Conference on Spartina Control. Department of Conservation and NaturalResources, Victoria.
Williamson, M. and Fitter, A. 1995. The varying success of invaders. Ecology 77:11661-1666.
Williamson, R. 1996. Spartina in Victoria: An Overview. In: Proceedings of theAustralasian Conference on Spartina Control. Department of Conservation and NaturalResources, Victoria.
Wu, M., Hacker, S., Ayres, D. and Strong, D.R. 1999. Potential of Prokelisia spp. AsBiological Control Agents of English Cordgrass, Spartina anglica. Biological Control 16:267-273.
Pest Plant Invasiveness Assessment
Prepared by the Keith Turnbull Research Institute, 2002. 24
Appendix 1:
Invasive Potential Criteriaand Intensity Ratings
Pest
Pla
nt In
vasi
vene
ss A
sses
smen
t
Pre
pare
d by
the
Kei
th T
urnb
ull R
esea
rch
Inst
itute
, 200
2.
25
Cri
teri
aR
ank
Inte
nsity
Rat
ing
Wei
ght
Est
ablis
hmen
tG
erm
inat
ion
/pro
pagu
leH
Opp
ortu
nist
ic g
erm
inat
or, c
an g
erm
inat
e or
strik
e/ se
t roo
t at a
ny ti
me
whe
neve
r wat
er is
ava
ilabl
e.1.
00re
quire
men
ts?
MH
Req
uire
s nat
ural
seas
onal
dis
turb
ance
s suc
h as
seas
onal
rain
fall,
sprin
g/su
mm
er te
mpe
ratu
res f
orge
rmin
atio
n.0.
67
MU
nkno
wn.
0.50
ML
Req
uire
s uns
easo
nal o
r unc
omm
on n
atur
al e
vent
s for
ger
min
atio
n (e
g. fl
oodi
ng, f
ire).
0.33
LR
equi
res s
peci
fic e
nviro
nmen
tal f
acto
rs th
at a
re n
ot p
art o
f an
annu
al c
ycle
to g
erm
inat
e (e
g. sp
ecifi
cte
mpe
ratu
res,
flood
s, fir
e O
R h
uman
cau
sed
dist
urba
nce
such
as p
loug
hing
).0
Seed
ling/
pro
pagu
leH
Can
est
ablis
h w
ithou
t add
ition
al fa
ctor
s.1.
00es
tabl
ishm
ent
MH
Can
est
ablis
h un
der m
oder
ate
cano
py/li
tter c
over
0.67
requ
irem
ents
MU
nkno
wn.
0.50
(i.e.
ligh
t, w
ater
,nu
trien
ts)?
ML
Req
uire
s mor
e sp
ecifi
c re
quire
men
ts to
est
ablis
h (e
g. o
pen
spac
e or
bar
e gr
ound
with
acc
ess t
o lig
htan
d di
rect
rain
fall)
.0.
33
LR
equi
res a
dditi
onal
and
ver
y sp
ecifi
c fa
ctor
s suc
h as
nut
rient
s and
wat
er th
at a
re d
elib
erat
ely
adde
d or
high
ly e
utro
phic
con
ditio
ns.
0
How
muc
h di
stur
banc
eH
Esta
blis
hes i
n he
alth
y an
d un
dist
urbe
d na
tura
l eco
syst
ems (
eg. m
alle
e, a
lpin
e, h
eath
land
s).
1.00
is re
quire
d fo
r see
dlin
ges
tabl
ishm
ent t
o oc
cur?
MH
Esta
blis
hes i
n re
lativ
ely
inta
ct o
r onl
y m
inor
dis
turb
ed n
atur
al e
cosy
stem
s (eg
. wet
land
s, rip
aria
n,riv
erin
e, g
rass
land
s, op
en w
oodl
ands
); in
vig
orou
sly
grow
ing
crop
s OR
in w
ell e
stab
lishe
d pa
stur
es.
0.67
MU
nkno
wn.
0.50
ML
Esta
blis
hes i
n hi
ghly
dis
turb
ed n
atur
al e
cosy
stem
s (eg
. roa
dsid
es, w
ildlif
e co
rrid
ors,
or a
reas
whi
chha
ve a
gre
ater
impa
ct b
y hu
man
s suc
h as
tour
ist a
reas
or c
amps
ites)
OR
in o
verg
raze
d pa
stur
es/p
oorly
grow
ing
or p
atch
y cr
ops.
0.33
LM
ajor
dis
turb
ance
requ
ired
with
littl
e O
R n
o co
mpe
titio
n fr
om o
ther
pla
nt sp
ecie
s.0
Gro
wth
/ Com
petit
ive
Abi
lity
Life
form
?H
Aqu
atic
(sub
mer
ged,
em
erge
nt, f
loat
ing)
and
sem
i aqu
atic
.1.
00M
HG
rass
, leg
umin
ous p
lant
.0.
67M
Unk
now
n.0.
50M
LG
eoph
yte,
clim
ber o
r cre
eper
.0.
33L
Oth
er.
0
Pest
Pla
nt In
vasi
vene
ss A
sses
smen
t
Pre
pare
d by
the
Kei
th T
urnb
ull R
esea
rch
Inst
itute
, 200
2.
26
Cri
teri
aR
ank
Inte
nsity
Rat
ing
Wei
ght
Alle
lopa
thic
pro
perti
es?
HM
ajor
alle
lopa
thic
pro
perti
es in
hibi
ting
all o
ther
pla
nts.
1.00
MH
Alle
lopa
thic
pro
perti
es se
rious
ly a
ffec
ting
som
e pl
ants
.0.
67M
Unk
now
n.0.
50M
LM
inor
pro
perti
es.
0.33
LN
one.
0
Abi
lity
to to
lera
tehe
rbiv
ory
pres
sure
HFa
vour
ed b
y he
avy
graz
ing
pres
sure
as n
ot e
aten
by
anim
als/
inse
cts a
nd n
ot u
nder
a b
ioco
ntro
lpr
ogra
m in
Aus
tralia
/New
Zea
land
.1.
00
and
prod
uce
prop
agul
es?
MH
Con
sum
ed b
ut n
on-p
refe
rred
or c
onsu
med
but
reco
vers
qui
ckly
; cap
able
of f
low
erin
g /s
eed
prod
uctio
n un
der m
oder
ate
herb
ivor
y pr
essu
re (w
here
mod
erat
e =
norm
al; n
ot o
vers
tock
ing
or h
eavy
graz
ing)
.
0.67
MU
nkno
wn.
0.50
ML
Con
sum
ed a
nd re
cove
rs sl
owly
. R
epro
duct
ion
stro
ngly
inhi
bite
d by
her
bivo
ry b
ut st
ill c
apab
le o
fve
geta
tive
prop
agul
e pr
oduc
tion
(by
rhiz
omes
or t
uber
s); w
eed
may
still
per
sist
.0.
33
LPr
efer
red
food
of h
erbi
vore
s. E
limin
ated
by
mod
erat
e he
rbiv
ory
or re
prod
uctio
n en
tirel
y pr
even
ted.
0
Nor
mal
gro
wth
rate
?H
Rap
id g
row
th ra
te th
at w
ill e
xcee
d m
ost o
ther
spec
ies o
f the
sam
e lif
e fo
rm.
1.00
MH
Mod
erat
ely
rapi
d gr
owth
that
will
equ
al c
ompe
titiv
e sp
ecie
s of t
he sa
me
life
form
.0.
67M
Unk
now
n.0.
50M
LM
axim
um g
row
th ra
te le
ss th
an m
any
spec
ies o
f the
sam
e lif
e fo
rm.
0.33
LSl
ow g
row
th, w
ill b
e ex
ceed
ed b
y m
any
othe
r spe
cies
.0
Stre
ss to
lera
nce
ofes
tabl
ishe
d pl
ants
toH
Hig
hly
resi
stan
t to
at le
ast t
wo
of (d
roug
ht, f
rost
, wat
erlo
ggin
g, fi
re a
nd sa
linity
) not
susc
eptib
le to
mor
e th
an o
ne (c
anno
t be
drou
ght o
r wat
erlo
ggin
g).
1.00
fros
t, dr
ough
t, w
ater
logg
ing,
salin
ity, f
ire?
MH
Hig
hly
tole
rant
of a
t lea
st tw
o of
(dr
ough
t, fr
ost,
wat
erlo
ggin
g, fi
re a
nd sa
linity
) and
may
be to
lera
ntof
ano
ther
. Su
scep
tible
to a
t lea
st o
ne.
0.67
MU
nkno
wn.
0.50
ML
Tole
rant
to a
t lea
st tw
o an
d su
scep
tible
to a
t lea
st o
ne.
0.33
LM
aybe
tole
rant
of o
ne st
ress
, sus
cept
ible
to a
t lea
st tw
o.0
Pest
Pla
nt In
vasi
vene
ss A
sses
smen
t
Pre
pare
d by
the
Kei
th T
urnb
ull R
esea
rch
Inst
itute
, 200
2.
27
Cri
teri
aR
ank
Inte
nsity
Rat
ing
Wei
ght
Rep
rodu
ctio
n
Rep
rodu
ctiv
e sy
stem
?H
Bot
h ve
geta
tive
and
sexu
al re
prod
uctio
n.1.
00M
HV
eget
ativ
e re
prod
uctio
n.0.
67M
Unk
now
n.0.
50M
LSe
xual
(sel
f and
cro
ss p
ollin
atio
n).
0.33
LSe
xual
but
eith
er c
ross
or s
elf p
ollin
atio
n.0
Num
ber o
fH
Abo
ve 2
000.
1.00
prop
agul
es p
rodu
ced
MH
1000
-200
0.0.
67pe
r flo
wer
ing
even
t?M
Unk
now
n.0.
50M
L50
-100
0.0.
33L
Less
than
50.
0
Prop
agul
e lo
ngev
ity?
HG
reat
er th
an 2
5% o
f see
ds c
an su
rviv
e ov
er 2
0 ye
ars i
n th
e so
il.1.
00M
HG
reat
er th
an 2
5% o
f see
ds su
rviv
e 10
-20
year
s in
the
soil,
OR
low
er v
iabi
lity
but s
urvi
ves o
ver 2
0ye
ars.
0.67
MU
nkno
wn.
0.50
ML
Gre
ater
than
25%
of s
eeds
surv
ive
5-10
yea
rs in
the
soil,
OR
low
er v
iabi
lity
but s
urvi
ve 1
0-20
yea
rs.
0.33
LG
reat
er th
an 2
5% o
f see
ds su
rviv
e 5
year
s, O
R v
eget
ativ
ely
repr
oduc
es.
0R
epro
duct
ive
perio
d?H
Mat
ure
plan
t pro
duce
s via
ble
prop
agul
es fo
r 10
year
s or m
ore,
OR
spec
ies f
orm
s sel
f-su
stai
ning
mon
ocul
ture
s.1.
00
MH
Mat
ure
plan
t pro
duce
s via
ble
prop
agul
es fo
r 3–1
0 ye
ars.
0.67
MU
nkno
wn.
0.50
ML
Mat
ure
plan
t pro
duce
s via
ble
prop
agul
es fo
r onl
y 1–
2 ye
ars.
0.33
LM
atur
e pl
ant p
rodu
ces v
iabl
e pr
opag
ules
for o
nly
1 ye
ar.
0
Tim
e to
reac
hH
Rea
ches
mat
urity
and
pro
duce
s via
ble
prop
agul
es in
und
er a
yea
r.1.
00re
prod
uctiv
e m
atur
ity?
MH
Prod
uces
pro
pagu
les b
etw
een
1-2
year
s afte
r ger
min
atio
n.0.
67M
Unk
now
n.0.
50M
L2-
5 ye
ars.
0.33
LG
reat
er th
an 5
yea
rs to
reac
h se
xual
mat
urity
.0
Pest
Pla
nt In
vasi
vene
ss A
sses
smen
t
Pre
pare
d by
the
Kei
th T
urnb
ull R
esea
rch
Inst
itute
, 200
2.
28
Cri
teri
aR
ank
Inte
nsity
Rat
ing
Wei
ght
Dis
pers
alN
umbe
r of d
ispe
rsal
mec
hani
sms?
HV
ery
light
, win
d di
sper
sed
seed
s, O
R b
ird d
ispe
rsed
seed
s OR
has
edi
ble
frui
t tha
t is r
eadi
ly e
aten
by
high
ly m
obile
ani
mal
s.1.
00
MH
Prop
agul
es sp
read
by
win
d, w
ater
, ani
mal
s (no
t bird
s), l
ight
veh
icul
ar tr
affic
.0.
67M
Unk
now
n.0.
50M
LPr
opag
ules
spre
ad b
y at
tach
ing
to h
uman
s or a
nim
als.
0.33
LPr
opag
ules
mai
nly
spre
ad b
y gr
avity
.0
Prob
abili
ty (o
r cha
nce)
HV
ery
likel
y th
at so
me
prop
agul
es w
ill d
ispe
rse
grea
ter o
ne k
ilom
etre
.1.
00th
at p
ropa
gule
s will
MH
Few
pro
pagu
les w
ill d
ispe
rse
grea
ter t
han
one
kilo
met
re b
ut m
any
will
reac
h 20
0-10
00 m
etre
s.0.
67di
sper
se to
a d
ista
nce
MU
nkno
wn.
0.50
grea
ter t
han
one
kilo
met
re?
ML
Ver
y fe
w to
non
e w
ill d
ispe
rse
to o
ne k
ilom
etre
, mos
t 20-
200
met
res.
0.33
LV
ery
unlik
ely
to d
ispe
rse
grea
ter t
han
200
met
res,
mos
t les
s tha
n 20
met
res.
0
Pest Plant Invasiveness Assessment
Prepared by the Keith Turnbull Research Institute, 2002. 29
Appendix 2:
Table of Risk Ratings, National Statusand Invasiveness Indices
Pest
Pla
nt In
vasi
vene
ss A
sses
smen
t
Pre
pare
d by
the
Kei
th T
urnb
ull R
esea
rch
Inst
itute
, 200
2.
30
Scie
ntifi
c N
ame
Com
mon
Nam
eR
isk
Rat
ing
Nat
iona
l Sta
tus
Inva
sive
Inde
xA
caci
a ba
ileya
naC
oota
mun
dra
wat
tleV
Non
e0.
79A
cer
pseu
dopl
atan
usSy
cam
ore
map
leV
Non
e0.
72A
gros
tis c
apill
aris
Bro
wn-
top
bent
VN
one
0.65
Aila
nthu
s al
tissi
ma
Tree
-of-h
eave
nP
Vic
, NSW
, WA
0.89
Alli
um tr
ique
trum
Ang
led
onio
nV
Vic
, SA
, Tas
0.61
Alo
e sa
pona
ria
Alo
eS
Non
e0.
63A
ltern
anth
era
philo
xero
ides
Alli
gato
r wee
dN
one
Vic
, NSW
, QLD
, ACT
, SA
, NT,
WA
, Tas
0.69
Ant
hoxa
nthu
m o
dora
tum
Swee
t ver
nal-g
rass
VN
one
0.66
Asp
hode
lus
fistu
losu
sO
nion
-wee
dS
Vic
, NSW
, SA
, NT,
Tas
0.54
Bri
za m
axim
aLa
rge
quak
ing-
gras
sV
Non
e0.
74C
ardu
us p
ycno
ceph
alus
Slen
der t
hist
leS
Vic
, Tas
0.72
Car
duus
tenu
iflor
usSl
ende
r thi
stle
PV
ic, S
A, T
as0.
61C
arth
amus
lana
tus
Saff
ron
this
tleP
Vic
, NSW
, QLD
, NT,
WA
, Tas
0.51
Cen
taur
ea c
alci
trap
aSt
ar th
istle
SV
ic, N
SW0.
41C
estr
um p
arqu
iG
reen
poi
sonb
erry
Non
eV
ic, N
SW, Q
LD0.
73C
hrys
anth
emoi
des
mon
ilife
ra s
sp. m
onili
fera
Bon
esee
dV
Vic
, NSW
, QLD
, SA
, WA
0.80
Cir
sium
aca
rna
Sold
ier t
hist
leN
one
Vic
, SA
0.52
Cir
sium
arv
ense
Cal
iforn
ian
this
tleN
one
Vic
, NSW
, SA
, WA
, Tas
0.70
Cir
sium
vul
gare
Spea
r thi
stle
SV
ic, S
A, T
as0.
69C
onvo
lvul
us a
rven
sis
Com
mon
bin
dwee
dN
one
Vic
, SA
, WA
0.78
Cop
rosm
a re
pens
New
Zea
land
mirr
or-b
ush
VN
one
0.82
Cor
tade
ria
sello
ana
Pam
pas g
rass
VN
SW, W
A, T
as0.
95C
oton
east
er g
lauc
ophy
llus
Cot
onea
ster
VN
SW0.
81C
rata
egus
mon
ogyn
aH
awth
orn
VV
ic, S
A0.
70C
uscu
ta c
ampe
stri
sG
olde
n do
dder
SV
ic, N
SW, S
A, W
A, T
as0.
73C
ynar
a ca
rdun
culu
sA
rtich
oke
this
tleV
Vic
, SA
, WA
, Tas
0.66
Cyt
isus
sco
pari
usEn
glis
h br
oom
VV
ic, N
SW, S
A, A
CT, T
as0.
73
Pest
Pla
nt In
vasi
vene
ss A
sses
smen
t
Pre
pare
d by
the
Kei
th T
urnb
ull R
esea
rch
Inst
itute
, 200
2.
31
Scie
ntifi
c N
ame
Com
mon
Nam
eR
isk
Rat
ing
Nat
iona
l Sta
tus
Inva
sive
Inde
xD
atur
a st
ram
oniu
mC
omm
on th
orn-
appl
eS
Vic
, QLD
, WA
, NT,
Tas
0.61
Del
aire
a od
orat
aC
ape
ivy
VN
SW0.
72D
iplo
taxi
s te
nuifo
liaSa
nd m
usta
rdP
Vic
, SA
0.57
Dip
ogon
lign
osus
Dip
ogon
VW
A0.
82D
ipsa
cus
fullo
num
Wild
teas
elP
Vic
0.55
Ditt
rich
ia g
rave
olen
sSt
inkw
ort
PV
ic0.
61E
chiu
m p
lant
agin
eum
Pate
rson
's cu
rse
SV
ic, N
SW, S
A, W
A, N
T, T
as0.
66E
chiu
m v
ulga
reV
iper
's bu
glos
sP
Vic
, NSW
, Tas
0.66
Ehr
hart
a er
ecta
Pani
c ve
ldt g
rass
VN
one
0.77
Em
ex a
ustr
alis
Spin
y em
exS
Vic
, NSW
, QLD
, SA
, WA
, NT,
Tas
0.53
Era
gros
tis c
urvu
laA
fric
an lo
ve-g
rass
VV
ic, N
SW, S
A, W
A, A
CT,
Tas
0.71
Eri
ca lu
sita
nica
Span
ish
heat
hV
Non
e0.
77F
oeni
culu
m v
ulga
reFe
nnel
VV
ic, T
as0.
57G
alen
ia p
ubes
cens
Gal
enia
SN
SW0.
61G
enis
ta li
nifo
liaFl
ax-le
af b
room
VV
ic0.
70G
enis
ta m
onsp
essu
lana
Mon
tpel
lier b
room
VV
ic, N
SW, S
A, A
CT, T
as0.
70H
eder
a he
lixIv
yV
Non
e0.
76H
iera
cium
aur
antia
cum
Ora
nge
haw
kwee
dN
one
Non
e0.
62H
omer
ia fl
acci
daC
ape
tulip
(one
-leaf
)V
Vic
, NSW
, SA
, WA
, Tas
0.74
Hyp
eric
um a
ndro
saem
umTu
tsan
VV
ic, W
A0.
81H
yper
icum
per
fora
tum
St Jo
hn's
wor
tV
Vic
, NSW
, WA
, Tas
0.66
Hyp
eric
um te
trap
teru
mSq
uare
-ste
m S
t Joh
n's w
ort
PV
ic0.
55Il
ex a
quifo
lium
Hol
lyV
Non
e0.
75Ju
ncus
acu
tus
Shar
p ru
shV
Vic
0.52
Junc
us e
ffusu
sSo
ft ru
shV
Non
e0.
69La
vand
ula
stoe
chas
Topp
ed la
vend
erS
Vic
0.68
Leuc
anth
emum
vul
gare
Ox-
eye
dais
yP
Vic
0.72
Pest
Pla
nt In
vasi
vene
ss A
sses
smen
t
Pre
pare
d by
the
Kei
th T
urnb
ull R
esea
rch
Inst
itute
, 200
2.
32
Scie
ntifi
c N
ame
Com
mon
Nam
eR
isk
Rat
ing
Nat
iona
l Sta
tus
Inva
sive
Inde
xLe
yces
teri
a fo
rmos
aH
imal
ayan
hon
eysu
ckle
VN
one
0.80
Ligu
stru
m v
ulga
reEu
rope
an p
ivet
SN
one
0.75
Loni
cera
japo
nica
Japa
nese
hon
eysu
ckle
VN
one
0.83
Lyci
um fe
roci
ssim
umA
fric
an b
ox-th
orn
VV
ic, N
SW, Q
LD, S
A, W
A, N
T, T
as0.
67M
arru
bium
vul
gare
Hor
ehou
ndV
Vic
, NSW
, SA
, WA
, Tas
0.77
Mel
iant
hus
com
osus
Tufte
d ho
neyf
low
erN
one
Vic
0.49
Myr
ioph
yllu
m a
quat
icum
Parr
ot's
feat
her
PW
A, T
as0.
63M
yrsi
phyl
lum
asp
arag
oide
sB
ridal
cre
eper
VN
SW, S
A, W
A, T
as0.
79N
asse
lla h
yalin
aFi
ne n
eedl
e-gr
ass
SN
one
0.48
Nas
sella
nee
sian
aC
hile
an sp
ear-
gras
sV
NSW
0.69
Nas
sella
tric
hoto
ma
Serr
ated
tuss
ock
VV
ic, N
SW, S
A, A
CT, T
as0.
76O
nopo
rdum
aca
nthi
umH
eral
dic
this
tleN
one
Vic
, NSW
, Tas
0.57
Ono
pord
um a
caul
onSt
emle
ss o
nopo
rdon
SV
ic, N
SW, W
A, T
as0.
53O
punt
ia fi
cus-
indi
caPr
ickl
y-pe
arP
Non
e0.
76O
punt
ia r
obus
taW
heel
cac
tus
SV
ic, N
SW, Q
LD, S
A, W
A, N
T0.
66O
punt
ia s
tric
taC
omm
on p
rickl
y pe
arS
Vic
, NSW
, QLD
, SA
, WA
, NT
0.76
Oxa
lis p
es-c
apra
eSo
urso
bV
Vic
, SA
, Tas
0.74
Oxy
lobi
um la
nceo
latu
mO
xylo
bium
SN
one
0.62
Pas
palu
m d
ilata
tum
Pasp
alum
VN
one
0.73
Pen
nise
tum
cla
ndes
tinum
Kik
uyu
VN
one
0.67
Pha
lari
s aq
uatic
aTo
owoo
mba
can
ary-
gras
sV
Non
e0.
73P
hysa
lis v
isco
saPr
airie
gro
und
cher
ryN
one
Vic
, NSW
0.73
Pin
us p
inas
ter
Clu
ster
pin
eV
Non
e0.
79P
inus
rad
iata
Mon
tere
y pi
neV
Non
e0.
78P
ittos
poru
m u
ndul
atum
Swee
t pitt
ospo
rum
VN
SW, W
A0.
79P
olyg
ala
myr
tifol
iaM
yrtle
-leaf
milk
wor
tV
Non
e0.
76P
opul
us a
lba
Whi
te p
opla
rP
Non
e0.
66
Pest
Pla
nt In
vasi
vene
ss A
sses
smen
t
Pre
pare
d by
the
Kei
th T
urnb
ull R
esea
rch
Inst
itute
, 200
2.
33
Scie
ntifi
c N
ame
Com
mon
Nam
eR
isk
Rat
ing
Nat
iona
l Sta
tus
Inva
sive
Inde
xP
runu
s lu
sita
nica
Portu
gal l
aure
lS
Non
e0.
77R
esed
a lu
teol
aW
eld
Non
eV
ic, T
as0.
54R
ham
nus
alat
ernu
sIta
lian
buck
thor
nV
Non
e0.
82R
osa
rubi
gino
saSw
eet b
riar
VV
ic, N
SW, S
A, T
as0.
79R
ubus
frut
icos
us a
gg.
Bla
ckbe
rry
Non
eV
ic, N
SW, Q
LD, S
A, W
A, A
CT, T
as0.
77Sa
gitta
ria
gram
inea
Arr
owhe
adS
SA, W
A, T
as0.
58Sa
lix a
lba
Whi
te w
illow
SN
SW, S
A, A
CT
0.93
Salix
bab
ylon
ica
Wee
ping
will
owS
Non
e0.
93Sa
lix c
iner
eaG
rey
sallo
wV
NSW
, SA
, AC
T0.
92Sa
lix x
rub
ens
Cra
ck w
illow
VN
SW, S
A, A
CT
0.92
Salp
ichr
oa o
riga
nifo
liaPa
mpa
s lily
-of-
the-
valle
yV
Vic
, Tas
0.76
Scol
ymus
his
pani
cus
Gol
den
this
tleN
one
Vic
0.50
Sene
cio
jaco
baea
Rag
wor
tP
Vic
, NSW
, SA
, WA
, Tas
0.66
Sene
cio
pter
opho
rus
Win
ged
grou
ndse
lP
Vic
0.68
Sily
bum
mar
ianu
mV
arie
gate
d th
istle
SV
ic, S
A, W
A, T
as0.
76So
lanu
m e
laea
gnifo
lium
Silv
erle
af n
ight
shad
eN
one
Vic
, NSW
, SA
, Tas
0.67
Sola
num
linn
aean
umA
pple
of S
odom
SV
ic, W
A, T
as0.
36So
llya
hete
roph
ylla
Blu
ebel
l cre
eper
VN
one
0.81
Spar
tina
angl
ica
Spar
tina
VN
one
0.61
Spar
tina
x to
wns
endi
iSp
artin
aV
Non
e0.
58Sp
orob
olis
indi
cus
Rat
-tail
gras
sS
NSW
0.71
Trad
esca
ntia
alb
iflor
aW
ande
ring
cree
per
VN
SW0.
59Tr
ibul
us te
rres
tris
Cal
trop
Non
eV
ic, S
A, W
A, N
T, T
as0.
61U
lex
euro
paeu
sG
orse
VV
ic, N
SW, S
A, W
A, A
CT,
Tas
0.83
Ver
basc
um th
apsu
sG
reat
mul
lein
SV
ic0.
47V
erba
scum
vir
gatu
mTw
iggy
mul
lein
SN
one
0.64
Vin
ca m
ajor
Blu
e pe
riwin
kle
VN
one
0.67
Pest
Pla
nt In
vasi
vene
ss A
sses
smen
t
Pre
pare
d by
the
Kei
th T
urnb
ull R
esea
rch
Inst
itute
, 200
2.
34
Scie
ntifi
c N
ame
Com
mon
Nam
eR
isk
Rat
ing
Nat
iona
l Sta
tus
Inva
sive
Inde
xV
ulpi
a br
omoi
des
Squi
rrel
-tail
fesc
ueV
Non
e0.
67W
atso
nia
mer
iana
Wild
wat
soni
aS
Vic
, SA
, WA
0.40
Xan
thiu
m s
pino
sum
Bat
hurs
t bur
rS
Vic
, NSW
, QLD
, SA
, WA
, NT,
Tas
0.58
Xan
thiu
m s
trum
ariu
mN
oogo
ora
burr
Non
eV
ic, N
SW, Q
LD, S
A, W
A, N
T, A
CT, T
as0.
62L
egen
dR
isk
Rat
ing
(acc
ordi
ng to
Car
r et
al 1
992)
:V
- ve
ry se
rious
thre
at to
one
or m
ore
vege
tatio
n fo
rmat
ion
in V
icto
riaS
- ser
ious
thre
at to
one
or m
ore
vege
tatio
n fo
rmat
ion
in V
icto
riaP
- pot
entia
l thr
eat t
o on
e or
mor
e ve
geta
tion
form
atio
n in
Vic
toria
Non
e - N
ot p
rese
nt in
risk
ass
essm
ent
Nat
iona
l Sta
tus:
Vic
- N
oxio
us in
Vic
toria
NSW
- N
oxio
us in
New
Sou
th W
ales
Tas -
Nox
ious
in T
asm
ania
SA -
Nox
ious
in S
outh
Aus
tralia
WA
- N
oxio
us in
Wes
tern
Aus
tralia
QLD
- N
oxio
us in
Que
ensl
and
NT
- Nox
ious
in N
orth
ern
Terr
itory
AC
T - N
oxio
us in
Aus
tralia
n C
apita
l Ter
ritor
y
Pest Plant Invasiveness Assessment
Prepared by the Keith Turnbull Research Institute, 2002. 35
Appendix 3:
Examples of Present and PotentialDistribution Maps
Pest Plant Invasiveness Assessment
Prepared by the Keith Turnbull Research Institute, 2002. 36
List of species with present and potential distribution maps in the order they occur inAppendix 3. Updates to names are indicated.
Botanical name:
Acacia baileyana*Acer pseudoplatanus*Agrostis capillaris*Ailanthus altissimaAllium triquetrumAloe saponaria* = A. maculataAlternanthera philoxeroidesAnthoxanthum odoratum*Asphodelus fistulosus*Briza maxima*Carduus pycnocephalusCarduus tenuiflorusCarthamus lanatus*Centaurea calcitrapaCestrum parquiChrysanthemoides monilifera ssp. moniliferaCirsium acarna = Picnomon acarnaCirsium arvense*Cirsium vulgareConvolvulus arvensisCoprosma repens*Cortaderia selloanaCotoneaster glaucophyllus*Crataegus monogynaCuscuta campestrisCynara cardunculusCytisus scopariusDatura stramoniumDelairea odorata*Diplotaxis tenuifoliaDipogon lignosus*Dipsacus fullonumDittrichia graveolensEchium plantagineumEchium vulgareEhrharta erecta*Emex australisEragrostis curvulaErica lusitanica*Foeniculum vulgareGalenia pubescensGenista linifoliaGenista monspessulanaHedera helix*Hieracium aurantiacum
Pest Plant Invasiveness Assessment
Prepared by the Keith Turnbull Research Institute, 2002. 37
Homeria flaccida = Moraea flaccidaHypericum androsaemumHypericum perforatumHypericum tetrapterum*Ilex aquifolium*Juncus acutusJuncus effusus*Lavandula stoechasLeucanthemum vulgareLeycesteria formosaLigustrum vulgare*Lonicera japonica*Lycium ferocissimumMarrubium vulgareMelianthus comosusMyriophyllum aquaticum*Myrsiphyllum asparagoides = Asparagus asparagoidesNassella hyalinaNassella neesianaNassella trichotomaOnopordum acanthiumOnopordum acaulonOpuntia ficus-indicaOpuntia robustaOpuntia strictaOxalis pes-capraeOxylobium lanceolatum* = Callistachys lanceolataPaspalum dilatatum*Pennisetum clandestinum*Phalaris aquatica*Physalis viscosaPinus pinaster*Pinus radiata*Pittosporum undulatum*Polygala myrtifolia*Populus alba*Prunus lusitanica*Reseda luteolaRhamnus alaternus*Rosa rubiginosaRubus fruticosus agg.Sagittaria gramineaSalix albaSalix babylonicaSalix cinereaSalix x rubensSalpichroa origanifoliaScolymus hispanicusSenecio jacobaea
Pest Plant Invasiveness Assessment
Prepared by the Keith Turnbull Research Institute, 2002. 38
Senecio pterophorusSilybum marianumSolanum elaeagnifoliumSolanum linnaeanumSollya heterophylla*Spartina anglicaSpartina x townsendiiSporobolus indicus = S. africanusTradescantia albiflora* = T. fluminensisTribulus terrestrisUlex europaeusVerbascum thapsusVerbascum virgatum*Vinca major*Vulpia bromoides*Watsonia merianaXanthium spinosumXanthium strumarium = X. occidentale; X. orientale* Indicates a species with PV district boundaries on the potential distribution maps; the others
species, previously assessed by KTRI, have the CMA boundaries overlayed onto the potentialdistribution maps.
Pest Plant Invasiveness Assessment
Prepared by the Keith Turnbull Research Institute, 2002. 39
Maps in Appendix 3 are located in a separate document.