Hungarian oak forests from the Central European perspective Jan Roleček, Vegetation Science Group,...

1
Hungarian oak forests from the Central Hungarian oak forests from the Central European perspective European perspective Jan Roleček, Vegetation Science Group, Institute of Botany and Zoology Masaryk University, Brno, Czech Republic [email protected] Who are we? The project is coordinated by the author at the Institute of Botany and Zoology in Brno, Czech Republic. We cooperate with the experts in all four concerned countries:Ilona Knollová & David Zelený, Institute of Botany and Zoology, Brno Wolfgang Willner, VINCA – Institut für Naturschutzforschung und Ökologie, Wien Franz Starlinger, Bundesamt für Wald, Wien János Csiky, Dept. of Systematic and Ecological Botany, University of Pécs Milan Valachovič, Institute of Botany, Slovak Academy of Sciences, Bratislava Jana Sadloňová, Institute of Botany, Slovak Academy of Sciences, Bratislava The project is supported by the Grant Agency of the Academy of Sciences of the Czech Republic grant B601630504 and by the long- term research plan MSM 0021622416. What we do? Since 2002, an extensive research of the variability of oak forest vegetation has been conducted in four Central European countries (Czech Republic, Slovakia, Austria, Hungary). The research combines traditional methods of phytosociological field research (Braun-Blanquet approach) and advanced methods of vegetation data analysis (multivariate analysis of vegetation variability, indicator species analysis, ecological analysis of vegetation using species traits, etc.). The aim of the research is to evaluate the overall variability of Central European oak forests, to determine the main gradients of their species composition on different spatial scales, and to draw syntaxonomic conclusions. Why we do that? The traditional phytosociology brings distorted view of the variability of vegetation, especially when moving to larger spatial scales (Chytrý 2001, Knollová & Chytrý 2004). Therefore it is not possible to describe the general patterns of species coexistence realistically in such large regions as e.g. Hungary by summarizing the results of local phytosociological studies. It is necessary to perform large-scale analysis of extensive phytosociological datasets then. On the other hand, the traditional phytosociology harbours indispensable knowledge on the local vegetation variability in the particular time and space. Thus the complementarity of the traditional and modern large-scale approaches must be emphasized. The challenge for the contemporary vegetation science is to make the best account of the advantages of both approaches. Cited literature Chytrý M. 2001: Phytosociological data give biased estimates of species richness. Journal of Vegetation Science 12: 439-444. Knollová I. & Chytrý M. 2004: Oak-hornbeam forests of the Czech Republic: geographical and ecological approaches to vegetation classification. Preslia 76: 291-311. Chytrý M., Tichý L., Holt J. & Botta-Dukát Z. (2002): Determination of diagnostic species with statistical fidelity measures. Journal of Vegetation Some preliminary results This is just to show rough patterns of variability and of geographic distribution of thermophilous oak forests in Central Europe, as they appear when you get the phytosociological data together (about 2500 relevés). We still have to improve the geographic and ecological representativeness of the data, to find the geographic coordinates of many relevés, and to acquire necessary environmental data. Who are we? The project is coordinated by the author at the Institute of Botany and Zoology in Brno, Czech Republic. We cooperate with the experts in all four concerned countries: Ilona Knollová & David Zelený, Institute of Botany and Zoology, Brno Wolfgang Willner, VINCA – Institut für Naturschutzforschung und Ökologie, Wien Franz Starlinger, Bundesamt für Wald, Wien János Csiky, Dept. of Systematic and Ecological Botany, University of Pécs Milan Valachovič, Institute of Botany, Slovak Academy of Sciences, Bratislava Jana Sadloňová, Institute of Botany, Slovak Academy of Sciences, Bratislava The project is supported by the Grant Agency of the Academy of Sciences of the Czech Republic grant B601630504 and by the long- term research plan MSM 0021622416. 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 Diagnostic species - group 2 Festuca ovina s.l. Luzula luzuloides Avenella flexuosa Hypericum perforatum Pinus sylvestris Hieracium lachenalii Quercus petraea agg. Galium pumilum s.l. Poa nemoralis Hieracium pilosella Dianthus carthusianorum s.l. Luzula campestris agg. Lychnis viscaria Agrostis capillaris Steris viscaria Diagnostic species - group 1 Hieracium pilosella Festuca ovina s.l. Linaria genistifolia Jasione montana Rumex acetosella Carex humilis Genista pilosa Luzula campestris agg. Phleum phleoides Dianthus carthusianorum s.l. Hypericum perforatum Trifolium alpestre Quercus petraea agg. Avenella flexuosa Sedum rupestre CZ AU CZ AU SK HU Diagnostic species - group 3 Poa nemoralis Quercus petraea agg. Lathyrus niger Hieracium sabaudum Lathyrus vernus Carex montana Veronica officinalis Festuca heterophylla Galium sylvaticum Stellaria holostea Viola reichenbach/riviniana Melampyrum pratense Luzula luzuloides Ajuga reptans Hieracium murorum CZ AU SK HU Diagnostic species - group 4 Luzula luzuloides Fagus sylvatica Hieracium murorum Avenella flexuosa Vaccinium myrtillus Quercus petraea agg. Digitalis grandiflora Senecio nemorensis agg. Calamagrostis arundinacea Hieracium racemosum Polypodium vulgare agg. Poa stiriaca Carpinus betulus Melampyrum pratense Sorbus aucuparia CZ AU SK HU Diagnostic species - group 5 Geum urbanum Brachypodium sylvaticum Carex michelii Ulmus minor Viola hirta Dictamnus albus Carex muricata agg. Euonymus europaea Buglossoides purpurocaerulea Ligustrum vulgare Crataegus species Fallopia sp. Prunus spinosa Chaerophyllum temulum Robinia pseudacacia CZ AU SK HU Diagnostic species - group 6 Prunus spinosa Euonymus europaea Acer tataricum Poa pratensis agg. Quercus cerris Phlomis tuberosa Quercus pubescens agg. Polygonatum latifolium Carex praecox s.l. Campanula bononiensis Fragaria viridis Filipendula vulgaris Alliaria petiolata Prunus fruticosa Pulmonaria mollis agg. AU SK HU Diagnostic species - group 7 Vicia cassubica Hypericum hirsutum Rosa canina s.l. Poa pratensis agg. Quercus cerris Rubus fruticosus agg. Prunus spinosa Poa nemoralis Acer tataricum Astragalus glycyphyllos Calamagrostis epigejos Lychnis coronaria Veronica officinalis Torilis japonica Carex muricata agg. CZ SK HU Diagnostic species - group 8 Geum urbanum Polygonatum latifolium Sambucus nigra Alliaria petiolata Acer campestre Quercus cerris Viola suavis Ulmus minor Buglossoides purpurocaerulea Galium aparine Viola mirabilis Geranium robertianum Euonymus europaea Lactuca quercina Brachypodium sylvaticum AU SK HU Diagnostic species - group 9 Teucrium chamaedrys Astragalus glycyphyllos Sorbus torminalis Ligustrum vulgare Cornus mas Vincetoxic hirundinaria Carex michelii Fragaria viridis Bupleurum falcatum Rosa canina s.l. Crataegus species Viburnum lantana Securigera varia Medicago falcata Galium glaucum CZ AU SK Diagnostic species - group 10 Potentilla alba Lathyrus niger Carex montana Quercus cerris Convallaria majalis Cornus sanguinea Prunus avium Serratula tinctoria Prunus spinosa Festuca heterophylla Asphodelus albus Galium sylvaticum Euphorbia angulata Iris variegata Crataegus species CZ AU SK HU Diagnostic species - group 11 Viola mirabilis Euonymus verrucosa Acer campestre Cornus sanguinea Fraxinus excelsior Geum urbanum Polygonatum latifolium Viburnum lantana Prunus avium Carpinus betulus Brachypodium sylvaticum Cruciata laevipes Ligustrum vulgare Galium odoratum Pulmonaria officinalis agg. CZ AU SK HU Diagnostic species - group 12 Seseli libanotis Buphthalmum salicifolium Hieracium bifidum Cyclamen purpurascens Viola collina Knautia drymeia Salvia glutinosa Sesleria albicans Calamagrostis varia Lonicera xylosteum Silene nemoralis Euphorbia angulata Carpinus betulus Silene nutans Corylus avellana AU Diagnostic species - group 13 Fraxinus excelsior Hepatica nobilis Euonymus verrucosa Cornus mas Quercus pubescens agg. Viburnum lantana Arabis turrita Tilia platyphyllos Hedera helix Viola odorata Carex alba Sorbus aria agg. Primula veris Berberis vulgaris Buglossoides purpurocaerulea AU HU Diagnostic species - group 14 Quercus robur Frangula alnus Potentilla erecta Molinia caerulea agg. Potentilla alba Carex pallescens Scrophularia nodosa Viburnum opulus Deschampsia cespitosa Selinum carvifolia Betonica officinalis Rubus caesius Fragaria vesca Campanula patula Viola reichen/riviniana CZ AU SK HU Diagnostic species - group 15 Quercus robur Anthoxanthum odoratum Peucedanum oreoselinum Carex fritschii Galium verum s.l. Iris variegata Achillea millefolium agg. Carex pallescens Clinopodium vulgare Festuca ovina s.l. Cerastium arvense Arrhenatherum elatius Festuca rubra Ranunculus polyanthemos Betonica officinalis CZ SK HU What do the charts say? The 20 charts present 15 most diagnostic species of 20 vegetation types of Central European (sub)thermophilous oak forests. Diagnostic species were determined using phi coefficient of fidelity (Chytrý et al. 2002). The vegetation types were extracted using cluster analysis (Ward´s method, Euclidean distance) of ordination scores of relevés on first four ordination axes of detrended correspondence analysis (species cover values log- transformed). The pie charts present the relative number of relevés of a particular vegetation type coming from a particular country: HU – Hungary, SK – Slovakia, AU – Austria, CZ – Czech Republic. The dendrogram shows the similarity between the 20 vegetation types presented in the charts. Diagnostic species - group 16 Carex humilis Sesleria albicans Sorbus aria agg. Teucrium chamaedrys Potentilla heptaphylla agg. Pimpinella saxifraga s.l. Asperula tinctoria Leontodon incanus Quercus pubescens agg. Genista pilosa Vincetoxic hirundinaria Galium austriacum Asperula cynanchica Arabis hirsuta agg. Salvia pratensis CZ AU SK Diagnostic species - group 17 Cornus mas Teucrium chamaedrys Quercus pubescens agg. Berberis vulgaris Erysimum odoratum Vincetoxicum hirundinaria Carex humilis Bupleurum falcatum Anthericum ramosum Brachypodium pinnatum Securigera varia Stachys recta Pyrethrum corymbosum Juniperus communis Campanula rapunculoides CZ AU SK Diagnostic species - group 18 Erysimum odoratum Quercus pubescens agg. Teucrium chamaedrys Prunus mahaleb Festuca pallens s.l. Galium album s.l. Stachys recta Helianthemum grandiflorum Cornus mas Carex humilis Inula ensifolia Fraxinus ornus Carex michelii Geranium sanguineum Waldsteinia geoides CZ HU SK Diagnostic species - group 19 Stachys recta Galium glaucum Thymus pannonicus agg. Festuca rupicola Phleum phleoides Teucrium chamaedrys Achillea millefolium agg. Agropyron intermedium agg. Anthemis tinctoria Sedum sexangulare Asperula cynanchica Allium flavum Linaria genistifolia Lactuca viminea Acinos arvensis CZ AU SK HU Diagnostic species - group 20 Poa compressa Festuca rupicola Galium verum s.l. Hieracium bauhinii Seseli osseum Dorycnium pentaphyllum agg. Poa pannonica Veronica chamaedrys agg. Veronica officinalis Poa pratensis agg. Carex caryophyllea Festuca valesiaca s.l. Juniperus communis Agrimonia eupatoria Trifolium campestre SK acidophilous subthermophilous oak forests thermophilous oak forests of deep +- eutrophic soils basiphilous subxerophilous oak forests subthermophilous oak forests of heavy or sandy soils xerothermic oak forests of rocky substrates

Transcript of Hungarian oak forests from the Central European perspective Jan Roleček, Vegetation Science Group,...

Page 1: Hungarian oak forests from the Central European perspective Jan Roleček, Vegetation Science Group, Institute of Botany and Zoology Masaryk University,

Hungarian oak forests from the Central European Hungarian oak forests from the Central European perspectiveperspective

Jan Roleček, Vegetation Science Group, Institute of Botany and Zoology Masaryk University, Brno, Czech Republic

[email protected]

Who are we?The project is coordinated by the author at the Institute of Botany and Zoology in Brno, Czech Republic. We cooperate with the experts in all four concerned countries:Ilona Knollová & David Zelený, Institute of Botany and Zoology, Brno Wolfgang Willner, VINCA – Institut für Naturschutzforschung und Ökologie, WienFranz Starlinger, Bundesamt für Wald, WienJános Csiky, Dept. of Systematic and Ecological Botany, University of PécsMilan Valachovič, Institute of Botany, Slovak Academy of Sciences, BratislavaJana Sadloňová, Institute of Botany, Slovak Academy of Sciences, Bratislava

The project is supported by the Grant Agency of the Academy of Sciences of the Czech Republic grant B601630504 and by the long-term research plan MSM 0021622416.

What we do?Since 2002, an extensive research of the variability of oak forest vegetation has been conducted in four Central European countries (Czech Republic, Slovakia, Austria, Hungary). The research combines traditional methods of phytosociological field research (Braun-Blanquet approach) and advanced methods of vegetation data analysis (multivariate analysis of vegetation variability, indicator species analysis, ecological analysis of vegetation using species traits, etc.). The aim of the research is to evaluate the overall variability of Central European oak forests, to determine the main gradients of their species composition on different spatial scales, and to draw syntaxonomic conclusions.

Why we do that? The traditional phytosociology brings distorted view of the variability of vegetation, especially when moving to larger spatial scales (Chytrý 2001, Knollová & Chytrý 2004). Therefore it is not possible to describe the general patterns of species coexistence realistically in such large regions as e.g. Hungary by summarizing the results of local phytosociological studies. It is necessary to perform large-scale analysis of extensive phytosociological datasets then.

On the other hand, the traditional phytosociology harbours indispensable knowledge on the local vegetation variability in the particular time and space. Thus the complementarity of the traditional and modern large-scale approaches must be emphasized. The challenge for the contemporary vegetation science is to make the best account of the advantages of both approaches.

Cit

ed

lite

ratu

reC

hyt

rý M

. 20

01:

Phy

toso

ciol

ogic

al d

ata

give

bia

sed

estim

ates

of

spec

ies

richn

ess.

Jou

rnal

of

Veg

etat

ion

Sci

ence

12:

439

-444

.K

no

llo

vá I

. &

Ch

ytrý

M.

2004

: O

ak-h

ornb

eam

for

ests

of

the

Cze

ch R

epub

lic:

geog

raph

ical

and

ec

olog

ical

app

roac

hes

to v

eget

atio

n cl

assi

ficat

ion.

Pre

slia

76:

291

-311

.C

hyt

rý M

., T

ich

ý L

., H

olt

J.

& B

ott

a-D

uká

t Z

. (2

002)

: D

eter

min

atio

n of

dia

gnos

tic s

peci

es w

ith

stat

istic

al f

idel

ity m

easu

res.

Jou

rnal

of

Veg

etat

ion

Sci

ence

13:

79-

90.

Some preliminary resultsThis is just to show rough patterns of variability and of geographic distribution of thermophilous oak forests in Central Europe, as they appear when you get the phytosociological data together (about 2500 relevés). We still have to improve the geographic and ecological representativeness of the data, to find the geographic coordinates of many relevés, and to acquire necessary environmental data.

Who are we?The project is coordinated by the author at the Institute of Botany and Zoology in Brno, Czech Republic. We cooperate with the experts in all four concerned countries:Ilona Knollová & David Zelený, Institute of Botany and Zoology, Brno Wolfgang Willner, VINCA – Institut für Naturschutzforschung und Ökologie, WienFranz Starlinger, Bundesamt für Wald, WienJános Csiky, Dept. of Systematic and Ecological Botany, University of PécsMilan Valachovič, Institute of Botany, Slovak Academy of Sciences, BratislavaJana Sadloňová, Institute of Botany, Slovak Academy of Sciences, Bratislava

The project is supported by the Grant Agency of the Academy of Sciences of the Czech Republic grant B601630504 and by the long-term research plan MSM 0021622416.

1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20

Diagnostic species - group 2

Festuca ovina s.l.

Luzula luzuloides

Avenella flexuosa

Hypericum perforatum

Pinus sylvestris

Hieracium lachenalii

Quercus petraea agg.

Galium pumilum s.l.

Poa nemoralis

Hieracium pilosella

Dianthus carthusianorum s.l.

Luzula campestris agg.

Lychnis viscaria

Agrostis capillaris

Steris viscaria

Diagnostic species - group 1

Hieracium pilosella

Festuca ovina s.l.

Linaria genistifolia

Jasione montana

Rumex acetosella

Carex humilis

Genista pilosa

Luzula campestris agg.

Phleum phleoides

Dianthus carthusianorum s.l.

Hypericum perforatum

Trifolium alpestre

Quercus petraea agg.

Avenella flexuosa

Sedum rupestre

CZAU

CZ AU

SKHU

Diagnostic species - group 3

Poa nemoralis

Quercus petraea agg.

Lathyrus niger

Hieracium sabaudum

Lathyrus vernus

Carex montana

Veronica officinalis

Festuca heterophylla

Galium sylvaticum

Stellaria holostea

Viola reichenbach/riviniana

Melampyrum pratense

Luzula luzuloides

Ajuga reptans

Hieracium murorum

CZ

AU

SK

HU

Diagnostic species - group 4

Luzula luzuloides

Fagus sylvatica

Hieracium murorum

Avenella flexuosa

Vaccinium myrtillus

Quercus petraea agg.

Digitalis grandiflora

Senecio nemorensis agg.

Calamagrostis arundinacea

Hieracium racemosum

Polypodium vulgare agg.

Poa stiriaca

Carpinus betulus

Melampyrum pratense

Sorbus aucuparia

CZ

AU

SK

HU

Diagnostic species - group 5

Geum urbanum

Brachypodium sylvaticum

Carex michelii

Ulmus minor

Viola hirta

Dictamnus albus

Carex muricata agg.

Euonymus europaea

Buglossoides purpurocaerulea

Ligustrum vulgare

Crataegus species

Fallopia sp.

Prunus spinosa

Chaerophyllum temulum

Robinia pseudacacia

CZ

AU

SK

HU

Diagnostic species - group 6

Prunus spinosa

Euonymus europaea

Acer tataricum

Poa pratensis agg.

Quercus cerris

Phlomis tuberosa

Quercus pubescens agg.

Polygonatum latifolium

Carex praecox s.l.

Campanula bononiensis

Fragaria viridis

Filipendula vulgaris

Alliaria petiolata

Prunus fruticosa

Pulmonaria mollis agg.

AUSK

HU

Diagnostic species - group 7

Vicia cassubica

Hypericum hirsutum

Rosa canina s.l.

Poa pratensis agg.

Quercus cerris

Rubus fruticosus agg.

Prunus spinosa

Poa nemoralis

Acer tataricum

Astragalus glycyphyllos

Calamagrostis epigejos

Lychnis coronaria

Veronica officinalis

Torilis japonica

Carex muricata agg.

CZ

SK

HU

Diagnostic species - group 8

Geum urbanum

Polygonatum latifolium

Sambucus nigra

Alliaria petiolata

Acer campestre

Quercus cerris

Viola suavis

Ulmus minor

Buglossoides purpurocaerulea

Galium aparine

Viola mirabilis

Geranium robertianum

Euonymus europaea

Lactuca quercina

Brachypodium sylvaticum

AU

SK

HU

Diagnostic species - group 9

Teucrium chamaedrys

Astragalus glycyphyllos

Sorbus torminalis

Ligustrum vulgare

Cornus mas

Vincetoxic hirundinaria

Carex michelii

Fragaria viridis

Bupleurum falcatum

Rosa canina s.l.

Crataegus species

Viburnum lantana

Securigera varia

Medicago falcata

Galium glaucum

CZ

AU

SK

Diagnostic species - group 10

Potentilla alba

Lathyrus niger

Carex montana

Quercus cerris

Convallaria majalis

Cornus sanguinea

Prunus avium

Serratula tinctoria

Prunus spinosa

Festuca heterophylla

Asphodelus albus

Galium sylvaticum

Euphorbia angulata

Iris variegata

Crataegus species

CZ

AU

SK

HU

Diagnostic species - group 11

Viola mirabilis

Euonymus verrucosa

Acer campestre

Cornus sanguinea

Fraxinus excelsior

Geum urbanum

Polygonatum latifolium

Viburnum lantana

Prunus avium

Carpinus betulus

Brachypodium sylvaticum

Cruciata laevipes

Ligustrum vulgare

Galium odoratum

Pulmonaria officinalis agg.

CZ

AUSK

HU

Diagnostic species - group 12

Seseli libanotis

Buphthalmum salicifolium

Hieracium bifidum

Cyclamen purpurascens

Viola collina

Knautia drymeia

Salvia glutinosa

Sesleria albicans

Calamagrostis varia

Lonicera xylosteum

Silene nemoralis

Euphorbia angulata

Carpinus betulus

Silene nutans

Corylus avellana

AU

Diagnostic species - group 13

Fraxinus excelsior

Hepatica nobilis

Euonymus verrucosa

Cornus mas

Quercus pubescens agg.

Viburnum lantana

Arabis turrita

Tilia platyphyllos

Hedera helix

Viola odorata

Carex alba

Sorbus aria agg.

Primula veris

Berberis vulgaris

Buglossoides purpurocaerulea

AU

HU

Diagnostic species - group 14

Quercus robur

Frangula alnus

Potentilla erecta

Molinia caerulea agg.

Potentilla alba

Carex pallescens

Scrophularia nodosa

Viburnum opulus

Deschampsia cespitosa

Selinum carvifolia

Betonica officinalis

Rubus caesius

Fragaria vesca

Campanula patula

Viola reichen/riviniana

CZ

AU SK

HU

Diagnostic species - group 15

Quercus robur

Anthoxanthum odoratum

Peucedanum oreoselinum

Carex fritschii

Galium verum s.l.

Iris variegata

Achillea millefolium agg.

Carex pallescens

Clinopodium vulgare

Festuca ovina s.l.

Cerastium arvense

Arrhenatherum elatius

Festuca rubra

Ranunculus polyanthemos

Betonica officinalis

CZ

SK

HU

What do the charts say?

The 20 charts present 15 most diagnostic species of 20 vegetation types of Central European (sub)thermophilous oak forests. Diagnostic species were determined using phi coefficient of fidelity (Chytrý et al. 2002). The vegetation types were extracted using cluster analysis (Ward´s method, Euclidean distance) of ordination scores of relevés on first four ordination axes of detrended correspondence analysis (species cover values log-transformed).

The pie charts present the relative number of relevés of a particular vegetation type coming from a particular country: HU – Hungary, SK – Slovakia, AU – Austria, CZ – Czech Republic.

The dendrogram shows the similarity between the 20 vegetation types presented in the charts.

Diagnostic species - group 16

Carex humilis

Sesleria albicans

Sorbus aria agg.

Teucrium chamaedrys

Potentilla heptaphylla agg.

Pimpinella saxifraga s.l.

Asperula tinctoria

Leontodon incanus

Quercus pubescens agg.

Genista pilosa

Vincetoxic hirundinaria

Galium austriacum

Asperula cynanchica

Arabis hirsuta agg.

Salvia pratensis

CZAU

SK

Diagnostic species - group 17

Cornus mas

Teucrium chamaedrys

Quercus pubescens agg.

Berberis vulgaris

Erysimum odoratum

Vincetoxicum hirundinaria

Carex humilis

Bupleurum falcatum

Anthericum ramosum

Brachypodium pinnatum

Securigera varia

Stachys recta

Pyrethrum corymbosum

Juniperus communis

Campanula rapunculoides

CZ

AUSK

Diagnostic species - group 18

Erysimum odoratum

Quercus pubescens agg.

Teucrium chamaedrys

Prunus mahaleb

Festuca pallens s.l.

Galium album s.l.

Stachys recta

Helianthemum grandiflorum

Cornus mas

Carex humilis

Inula ensifolia

Fraxinus ornus

Carex michelii

Geranium sanguineum

Waldsteinia geoides

CZ

HUSK

Diagnostic species - group 19

Stachys recta

Galium glaucum

Thymus pannonicus agg.

Festuca rupicola

Phleum phleoides

Teucrium chamaedrys

Achillea millefolium agg.

Agropyron intermedium agg.

Anthemis tinctoria

Sedum sexangulare

Asperula cynanchica

Allium flavum

Linaria genistifolia

Lactuca viminea

Acinos arvensis

CZ

AUSK

HU

Diagnostic species - group 20

Poa compressa

Festuca rupicola

Galium verum s.l.

Hieracium bauhinii

Seseli osseum

Dorycnium pentaphyllum agg.

Poa pannonica

Veronica chamaedrys agg.

Veronica officinalis

Poa pratensis agg.

Carex caryophyllea

Festuca valesiaca s.l.

Juniperus communis

Agrimonia eupatoria

Trifolium campestre

SK

acidophilous subthermophilous

oak forests

thermophilous oak forests of deep

+- eutrophic soils

basiphilous subxerophilous

oak forests

subthermophilous oak forests of heavy

or sandy soils

xerothermic oak forests

of

rock

y s

ub

stra

tes