Gould S.J. & Eldredge N. 1977 Punctuated equilibria: the tempo and mode of evolution reconsidered....
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Transcript of Gould S.J. & Eldredge N. 1977 Punctuated equilibria: the tempo and mode of evolution reconsidered....
![Page 1: Gould S.J. & Eldredge N. 1977 Punctuated equilibria: the tempo and mode of evolution reconsidered. Paleobiology 3: 115-151. Herhaling " most evolutionary.](https://reader034.fdocuments.in/reader034/viewer/2022051515/551bedb3550346c3588b6410/html5/thumbnails/1.jpg)
• Gould S.J. & Eldredge N. 1977Punctuated equilibria: the tempo and mode of evolution reconsidered.Paleobiology 3: 115-151.
Herhaling
"most evolutionary change is concentrated in rapid events of speciation in small, peripherally isolated populations"
"the norm for a species … is morphological stasis"
"paleontologists have worn blinders … ignored gaps in the stratigraphic sequences … and evolutinaru stasis"
![Page 2: Gould S.J. & Eldredge N. 1977 Punctuated equilibria: the tempo and mode of evolution reconsidered. Paleobiology 3: 115-151. Herhaling " most evolutionary.](https://reader034.fdocuments.in/reader034/viewer/2022051515/551bedb3550346c3588b6410/html5/thumbnails/2.jpg)
• Gould S.J. & Eldredge N. 1977Punctuated equilibria: the tempo and mode of evolution reconsidered.Paleobiology 3: 115-151.
Herhaling
Verpersoonlijking van de "vijand"
John Wyatt Durham
![Page 3: Gould S.J. & Eldredge N. 1977 Punctuated equilibria: the tempo and mode of evolution reconsidered. Paleobiology 3: 115-151. Herhaling " most evolutionary.](https://reader034.fdocuments.in/reader034/viewer/2022051515/551bedb3550346c3588b6410/html5/thumbnails/3.jpg)
Gryphaea
"…Trueman rejoiced in Gryphaea, but never mentioned the hundreds of Liassic species that showed no temporal change … "
![Page 4: Gould S.J. & Eldredge N. 1977 Punctuated equilibria: the tempo and mode of evolution reconsidered. Paleobiology 3: 115-151. Herhaling " most evolutionary.](https://reader034.fdocuments.in/reader034/viewer/2022051515/551bedb3550346c3588b6410/html5/thumbnails/4.jpg)
Micraster
"…Rowe monographed Micraster but spoke not a word about its legion of static colleagues in the English chalk … "
![Page 5: Gould S.J. & Eldredge N. 1977 Punctuated equilibria: the tempo and mode of evolution reconsidered. Paleobiology 3: 115-151. Herhaling " most evolutionary.](https://reader034.fdocuments.in/reader034/viewer/2022051515/551bedb3550346c3588b6410/html5/thumbnails/5.jpg)
"…Ziegler's celebrated study of the Silurian brachiopod Eocoelia … presents data for five characters. Three display a fluctuating pattern, one no trend at all, and the last a directional pattern. Only the last is graphed."
![Page 6: Gould S.J. & Eldredge N. 1977 Punctuated equilibria: the tempo and mode of evolution reconsidered. Paleobiology 3: 115-151. Herhaling " most evolutionary.](https://reader034.fdocuments.in/reader034/viewer/2022051515/551bedb3550346c3588b6410/html5/thumbnails/6.jpg)
Lepidolina multiseptata
Tomowo Ozawa
"…Ozawa measures nine characters … Four exhibit gradual trends; five do not. The first four are either graphed or listed in tables; the remaining five are discussed in qualitative terms only."
![Page 7: Gould S.J. & Eldredge N. 1977 Punctuated equilibria: the tempo and mode of evolution reconsidered. Paleobiology 3: 115-151. Herhaling " most evolutionary.](https://reader034.fdocuments.in/reader034/viewer/2022051515/551bedb3550346c3588b6410/html5/thumbnails/7.jpg)
• Gould S.J. & Eldredge N. 1977Punctuated equilibria: the tempo and mode of evolution reconsidered.Paleobiology 3: 115-151.
Herhaling
Verpersoonlijking van de "vijand"
Review van empirische testen
Studies op verkeerde schaal te kleine schaal ("neontologische" studies) te grote schaal (graduele series van soorten)
![Page 8: Gould S.J. & Eldredge N. 1977 Punctuated equilibria: the tempo and mode of evolution reconsidered. Paleobiology 3: 115-151. Herhaling " most evolutionary.](https://reader034.fdocuments.in/reader034/viewer/2022051515/551bedb3550346c3588b6410/html5/thumbnails/8.jpg)
• Gould S.J. & Eldredge N. 1977Punctuated equilibria: the tempo and mode of evolution reconsidered.Paleobiology 3: 115-151.
Herhaling
Verpersoonlijking van de "vijand"
Review van empirische testen
Studies op verkeerde schaal Studies met ontoereikende data(-analyse)
"…most cases presented as falsifications are circular because they rely upon the gradualistic presuppositions they claim to test…"
![Page 9: Gould S.J. & Eldredge N. 1977 Punctuated equilibria: the tempo and mode of evolution reconsidered. Paleobiology 3: 115-151. Herhaling " most evolutionary.](https://reader034.fdocuments.in/reader034/viewer/2022051515/551bedb3550346c3588b6410/html5/thumbnails/9.jpg)
• Gould S.J. & Eldredge N. 1977Punctuated equilibria: the tempo and mode of evolution reconsidered.Paleobiology 3: 115-151.
Herhaling
Verpersoonlijking van de "vijand"
Review empirische testen
Studies op verkeerde schaal Studies met ontoereikende data Mogelijks "ware" gevallen van gradualisme
ontworteling en reïnterpretatie
![Page 10: Gould S.J. & Eldredge N. 1977 Punctuated equilibria: the tempo and mode of evolution reconsidered. Paleobiology 3: 115-151. Herhaling " most evolutionary.](https://reader034.fdocuments.in/reader034/viewer/2022051515/551bedb3550346c3588b6410/html5/thumbnails/10.jpg)
• Gould S.J. & Eldredge N. 1977Punctuated equilibria: the tempo and mode of evolution reconsidered.Paleobiology 3: 115-151.
Herhaling
Verpersoonlijking van de "vijand"
Review empirische testen
Studies op verkeerde schaal Studies met ontoereikende data Mogelijks "ware" gevallen van gradualisme Studies die punctuated equilibria ondersteunen
![Page 11: Gould S.J. & Eldredge N. 1977 Punctuated equilibria: the tempo and mode of evolution reconsidered. Paleobiology 3: 115-151. Herhaling " most evolutionary.](https://reader034.fdocuments.in/reader034/viewer/2022051515/551bedb3550346c3588b6410/html5/thumbnails/11.jpg)
• Gould S.J. & Eldredge N. 1977Punctuated equilibria: the tempo and mode of evolution reconsidered.Paleobiology 3: 115-151.
Herhaling
Verpersoonlijking van de "vijand"
Review empirische testen
Nieuwe ideeën
Species selection
![Page 12: Gould S.J. & Eldredge N. 1977 Punctuated equilibria: the tempo and mode of evolution reconsidered. Paleobiology 3: 115-151. Herhaling " most evolutionary.](https://reader034.fdocuments.in/reader034/viewer/2022051515/551bedb3550346c3588b6410/html5/thumbnails/12.jpg)
• Gould S.J. & Eldredge N. 1977Punctuated equilibria: the tempo and mode of evolution reconsidered.Paleobiology 3: 115-151.
Herhaling
Verpersoonlijking van de "vijand"
Review empirische testen
Nieuwe ideeën
Species selection relative ability to speciate, not morphological flexibility
![Page 13: Gould S.J. & Eldredge N. 1977 Punctuated equilibria: the tempo and mode of evolution reconsidered. Paleobiology 3: 115-151. Herhaling " most evolutionary.](https://reader034.fdocuments.in/reader034/viewer/2022051515/551bedb3550346c3588b6410/html5/thumbnails/13.jpg)
• Gould S.J. & Eldredge N. 1977Punctuated equilibria: the tempo and mode of evolution reconsidered.Paleobiology 3: 115-151.
Herhaling
Verpersoonlijking van de "vijand"
Review empirische testen
Nieuwe ideeën
Species selection relative ability to speciate, not morphological flexibility "the need to translate micro to macroevolution guarantees that paleobiology shall provide essential theory"
![Page 14: Gould S.J. & Eldredge N. 1977 Punctuated equilibria: the tempo and mode of evolution reconsidered. Paleobiology 3: 115-151. Herhaling " most evolutionary.](https://reader034.fdocuments.in/reader034/viewer/2022051515/551bedb3550346c3588b6410/html5/thumbnails/14.jpg)
• Gould S.J. & Eldredge N. 1977Punctuated equilibria: the tempo and mode of evolution reconsidered.Paleobiology 3: 115-151.
Herhaling
Verpersoonlijking van de "vijand"
Review empirische testen
Nieuwe ideeën
Politiek
![Page 15: Gould S.J. & Eldredge N. 1977 Punctuated equilibria: the tempo and mode of evolution reconsidered. Paleobiology 3: 115-151. Herhaling " most evolutionary.](https://reader034.fdocuments.in/reader034/viewer/2022051515/551bedb3550346c3588b6410/html5/thumbnails/15.jpg)
• Gould S.J. & Eldredge N. 1977Punctuated equilibria: the tempo and mode of evolution reconsidered.Paleobiology 3: 115-151.
• Gould S.J. & Eldredge N. 1993Punctuated equilibria comes of age.Nature 366: 223-227.
Overwinning !
"punctuated equilibrium has been accepted by most of our colleagues … as a valuable addition to evolutionary theory"
![Page 16: Gould S.J. & Eldredge N. 1977 Punctuated equilibria: the tempo and mode of evolution reconsidered. Paleobiology 3: 115-151. Herhaling " most evolutionary.](https://reader034.fdocuments.in/reader034/viewer/2022051515/551bedb3550346c3588b6410/html5/thumbnails/16.jpg)
• Gould S.J. & Eldredge N. 1977Punctuated equilibria: the tempo and mode of evolution reconsidered.Paleobiology 3: 115-151.
• Gould S.J. & Eldredge N. 1993Punctuated equilibria comes of age.Nature 366: 223-227.
Overwinning !
"misunderstandings of colleagues … who failed to grasp …"
![Page 17: Gould S.J. & Eldredge N. 1977 Punctuated equilibria: the tempo and mode of evolution reconsidered. Paleobiology 3: 115-151. Herhaling " most evolutionary.](https://reader034.fdocuments.in/reader034/viewer/2022051515/551bedb3550346c3588b6410/html5/thumbnails/17.jpg)
• Gould S.J. & Eldredge N. 1977Punctuated equilibria: the tempo and mode of evolution reconsidered.Paleobiology 3: 115-151.
• Gould S.J. & Eldredge N. 1993Punctuated equilibria comes of age.Nature 366: 223-227.
Overwinning !
"the need to reformulate macroevolution, notably the key phenomenon of trends, as an accumulation of discrete speciation events treated as entities rather than undefinable segments of continua"
![Page 18: Gould S.J. & Eldredge N. 1977 Punctuated equilibria: the tempo and mode of evolution reconsidered. Paleobiology 3: 115-151. Herhaling " most evolutionary.](https://reader034.fdocuments.in/reader034/viewer/2022051515/551bedb3550346c3588b6410/html5/thumbnails/18.jpg)
• Gould S.J. & Eldredge N. 1977Punctuated equilibria: the tempo and mode of evolution reconsidered.Paleobiology 3: 115-151.
• Gould S.J. & Eldredge N. 1993Punctuated equilibria comes of age.Nature 366: 223-227.
Overwinning !
Macro-evolutie = species sorting
"Most macroevolution must be rendered by asking what kinds of species … did better than other (speciated more frequently, survived longer) … "
![Page 19: Gould S.J. & Eldredge N. 1977 Punctuated equilibria: the tempo and mode of evolution reconsidered. Paleobiology 3: 115-151. Herhaling " most evolutionary.](https://reader034.fdocuments.in/reader034/viewer/2022051515/551bedb3550346c3588b6410/html5/thumbnails/19.jpg)
• Gould S.J. & Eldredge N. 1977Punctuated equilibria: the tempo and mode of evolution reconsidered.Paleobiology 3: 115-151.
• Gould S.J. & Eldredge N. 1993Punctuated equilibria comes of age.Nature 366: 223-227.
Overwinning !
Macro-evolution = species sorting
Implicaties "levende fossielen" "darwins" nonsensicaal Cope's rule : kleine soorten speciëren sneller gecorreleerde punctuaties
![Page 20: Gould S.J. & Eldredge N. 1977 Punctuated equilibria: the tempo and mode of evolution reconsidered. Paleobiology 3: 115-151. Herhaling " most evolutionary.](https://reader034.fdocuments.in/reader034/viewer/2022051515/551bedb3550346c3588b6410/html5/thumbnails/20.jpg)
• Gould S.J. & Eldredge N. 1977Punctuated equilibria: the tempo and mode of evolution reconsidered.Paleobiology 3: 115-151.
• Gould S.J. & Eldredge N. 1993Punctuated equilibria comes of age.Nature 366: 223-227.
Overwinning !
Macro-evolution = species sorting
Implicaties
Empirische ondersteuning
![Page 21: Gould S.J. & Eldredge N. 1977 Punctuated equilibria: the tempo and mode of evolution reconsidered. Paleobiology 3: 115-151. Herhaling " most evolutionary.](https://reader034.fdocuments.in/reader034/viewer/2022051515/551bedb3550346c3588b6410/html5/thumbnails/21.jpg)
![Page 22: Gould S.J. & Eldredge N. 1977 Punctuated equilibria: the tempo and mode of evolution reconsidered. Paleobiology 3: 115-151. Herhaling " most evolutionary.](https://reader034.fdocuments.in/reader034/viewer/2022051515/551bedb3550346c3588b6410/html5/thumbnails/22.jpg)
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