Gases Exchange in Plants 1
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Transcript of Gases Exchange in Plants 1
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GASES
EXCHANGE IN
PLANTSPRESENTED BY :Siti Sarah binti Sekeri
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Introduction
Gas exchange three major gases with the
environment
These are CO (for photosynthesis), O(for respiration) and water vapour
(transpiration)
These gases may be taken up or releaseby plants
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During the day, most plants
produce oxygen through
photosynthesis and use up
carbon dioxide During the night, when there is
no photosynthesis, oxygen isconsumed for respiratory
processes
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In aquatic plants, carbon dioxide
required for photosynthesis is
obtained from the surroundingwater in the form of hydrogen-
carbonate ions from the carbonicacid present
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In terrestrial plants, the carbondioxide is obtained from the
atmosphere through the stomata
found on the surface of leaves,
stems and branches
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Structure of Stoma
Stoma is a pore or aperture that
penetrates the epidermis of leaves,
branches and stems of green plants Normally, more stomata are found
on the lower epidermis compared to
the upper epidermis of leaves, and
much less on the epidermis of stems
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Gaseous exchange in plants
occurs mainly through porescalled stomata
They are found on the epidermis
of leaves & stems of flowering
plants
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Lenticels found in the bark ofstem & root hairs with thin walls
& large surface are also allowgaseous exchange to take place
Each stoma consist of a stomatapore surrounded by two guardcells
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Each guard cell is kidney-shapes
& contains chloroplasts
It has a thinner outer wall & a
thicker, less elastic, inner wall
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Changes in turgid pressure of
guard cells causes the opening or
closing of the stomatal pore
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Besides, stoma is a formed by
two specialized epidermal cells,
bordering each other The guard cells play an important
role in the opening and closing ofthe stoma
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Next, the guard cells are bean-
shaped, and are different fromother epidermal cells
These guard cells contain
chlorophyll, thus can
photosynthesise, whereas the
epidermal cells do not containchlorophyll and thus cannot
photosynthesise
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Another unique characteristic ofthe guard cell is that the inner
cellulose wall bordering thestoma is thicker than the outercellulose wall bordering the
epidermal cells around the guardcells
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Functions of Stoma
Stoma plays important role in the
gaseous exchange of respiratory
gases, in particular theabsorption of CO, and the
regulation of water in the plants
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O produced during
photosynthesis diffuses outthrough the stomatal pores into
the atmosphere, while COdiffuses in from the atmosphere
into the leaves
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Under water deficit conditions,
stoma closes automatically
This helps to prevent water loss
from the plants until the waterdeficit diminishes
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STOMATAL
OPENING
STOMATALCLOSING
STOMATAL
OPENING
AND
CLOSING
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STOMATAL OPENING
It has been observed that stomatal opening is
promoted by high light intensity and low
mesophyll CO levels.
Guard cells, which contain chloroplasts,
generate ATP by photophosphorylation during
photosynthesis.
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Blue light (BL) is absorbed by
BL photoreceptors
Which activate a proto pump
(H-ATPase) in the cell
membrane of the guard cell.
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ATPs generated by the light-
dependent reaction ofphotosynthesis are hydrolysed to
provide energy to drive the proton
pump. As proton (H) are pumped out of
the guard cells, the cells become
increasingly negatively charge.
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Potassium channels are activated
and K ions diffuse fromsubsidiary cells through the
channels down this
electrochemical gradient into the
guard cells.
Chloride ions (Cl) then enter tobalance the charge.
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In some plants, starch are
converted into malate (the oldhypothesis proposed that
starch was converted tosugar).
The older hypothesis is thestarch-sugar hypothesis.
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During the daylight, starch is
converted into soluble sugars
to lower the water potential of
cell sap.
The reverse process occurs at
night.
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The accumulation of K ions
(and malate ions) causes thewater potential in the guard
cells to become more
negative.
Water enters by osmosis from
neighbouring subsidiary cells
into the guard cells.
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The guard cells become turgid.
The outer wall of guard cell is thinnerand more elastic than the thicker
inner wall.
There are cellulose microfibrils
radially arranged around the cell wall
and the ends of the two guard cellsare joined.
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The increased turgor pressure
therefore causes the guard cells
to curve outward and the stomaopens.
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Stomatal closure
Stomatal closure can be triggered by
water stress, high temperature,
increasing CO levels in the leaf
mesophyll and low light intensity (night-
time).
The hormone abscisic acid (ABA) issecreted by plant cell when transpiration
rate is high and soil water is low.
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This often occurs around mid-day or
when a plant begins to wilt. The ABA binds the receptors at the
cell membrane of the guard cells.
This increases the permeability of
calcium channels in the cell
membrane.
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Calcium ions (Ca ) enter into the
guard cell. The influx of Ca also triggers the
release of Ca from the cell vacuole
into the cytosol.
The accumulation of Ca inhibits
the proton pump.
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Potassium ions (K) move out of
the guard cells into subsidiarycells.
In some plants, Cl and certainorganic ions e.g malate ions also
move out of the guard cells.
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In the older hypothesis, at night
glucose is converted to insolublestarch.
The water potential in the guard cell
increases.
Water diffuses out to neighbouring
subsidiary cells by osmosis.
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The turgor pressure in the guard cells
decreases, the cells become flaccidand the stoma closes.
At the night, the chloroplasts in the
guard cells do not photosynthesise.
Less ATP is produced.
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There is no active uptake of K
ions.
Instead, the K ions diffuse out of
the guard cells.
The cells become flaccid and the
stoma closes.
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Factors affecting the opening and
closing of stoma
Circadian rhythms
Some plants have biological clock whereby
stomata open during the day and close at
night
Plants were observed to continue opening
and closing their stoma even when exposed
to 24 hours of continuous light
However, some plants in hot dry conditions
(CAM plants) have an opposite rhythm
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Water balance
When there is a shortage of water,
plants will wilt. Wilting closes stomata
Concentration of CO
A low concentration of CO in thesurrounding air causes stomata to open
Light
Light causes stomata to open wherebyplants are more sensitive to blue light
compared to red light