Community Ecology Chapter 53. Community Ecology A community is an assemblage of species...
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Transcript of Community Ecology Chapter 53. Community Ecology A community is an assemblage of species...
Community Ecology• A community is an assemblage of
species (populations) living close enough together for potential interaction in a habitat
Community Ecology• The boundaries of a community can be
set by the experimental question (the decomposers on a rotting log, the benthic community of Lake Erie, the trees and shrubs in Yellowstone National Park)
• Communities vary in their species richness and relative abundance
Interspecific Interactions• Interspecific competition – competition
for resources between different species when resources are in short supply
Interspecific Competition• Competitive exclusion principle – The
concept that when populations of two similar species compete for the same limited resources, one population will use the resources more efficiently and have a reproductive advantage that will eventually lead to the elimination of the other population
The Ecological Niche• An organism’s ecological niche is
described as its place in an ecosystem – its habitat and use of biotic and abiotic resources
• The competitive exclusion principle holds that two species with identical niches cannot coexist in a community
Resource Partitioning• Resource partitioning – slight variations
in niche that allow ecologically similar species to coexist; competition is a selection factor in evolution
Predation• Predation – a predator eating prey• Includes herbivory and parasitism• Eating and being eaten are prerequisite
to reproductive success• Adaptations to increase success in
predation may include acute senses, speed and agility, camouflage coloration, and physical structure such as claws, fangs, teeth, and stingers
Predation• Plants may defend themselves with
mechanical devices, such as thorns, or chemical compounds
• Animals can defend against predation by hiding, fleeing, or defending
• Cryptic coloration allows animals to blend in to their environment
• Mechanical and chemical defenses discourage predation
• Aposematic coloration warns predators not to eat animals with chemical defenses
Predation• Mimicry may be used by prey to exploit
the warning coloration of other species• Batesian mimicry – a palatable or
harmless species mimics an unpalatable or harmful model
• Müllerian mimicry – two or more unpalatable species resemble each other
Cuckoo BeeCuckoo Bee Yellow JacketYellow Jacket
Parasites• A parasite derives its nourishment from
another organism, the host, which is harmed in the process
• Parasites that live within the host are endoparasites ; those that feed on the surface of a host are ectoparasites
• In parasitoidism, insects lay eggs on hosts, on which their larvae then feed.
• Parasites do not usually cause lethal harm to the host on which they feed
Mutualism• In mutualism, interactions between
species benefit both participants• Mutualistic interactions require the
coevolution of adaptations in both species
• Many cases of mutual symbiosis may have evolved when organisms became able to derive some benefits from their predator or parasite Mutualism between Mutualism between
acacia trees and antsacacia trees and ants
Commensalism• In commensalism, only one member
appears to benefit from the interaction• Examples include “hitchhiking” species
and species that feed on food incidentally exposed by anotherThe Patella gets it
food from the plant, the Euklonia, which is not harmed or damaged in the process.
Example of Example of MutualismMutualism
Trophic Structure• The trophic structure
of a community is its feeding relations
• A food chain shows the transfer of food energy from one trophic level to the next
What Limits the Length of a Food Chain?
• Within a food web, each food chain consists of only a few links (usually five or fewer)
• Two hypotheses– Energetic hypothesis – food chains are limited
by inefficiency of energy transfer (only about 10%) from one trophic level to the next
– Dynamic stability hypothesis – suggests that short food chains are more stable than long ones because an environmental disruption that reduces production at lower levels will be magnified at higher trophic levels as food supply is reduced all the way up the chain
Dominant Species• Species in a community that have the
highest abundance or largest biomass are a major influence on the distribution and abundance of other species
• A species may become a dominant species due to its competitive use of resources or success at avoiding predation
• What happens if the dominant specie is removed from a habitat such as the American chestnut trees?
Keystone Species• A keystone species has a large impact
on community structure as a result of its ecological role
• Experiments with a predatory sea star (Pisaster) demonstrates the importance of keystone species
Structures of Communities• The structure of a community may be
controlled bottom-up by nutrients or top-down by predators
• According to the bottom-up model of community organization, N → V → H → P, an increase in nutrients yields an increase in biomass at each succeeding trophic level
• The top-down model, N ← V ← H ← P, assumes that predation controls community organization; also called the trophic cascade model
Disturbance and Community Structure• The nonequilibrium model emphasizes
the nonstable, changing structure and composition of communities as a result of disturbances
• Disturbances such as fire, drought, storms, overgrazing, or human activity change resource availability, reduce or eliminate some populations, and may create opportunities for new species
• Small scale disturbances may enhance environmental patchiness and help maintain species diversity
Ecological Succession• Ecological succession is the transition in
species composition in a community, usually following some disturbance
• Primary succession occurs in areas previously devoid of life (new volcanic island or the moraine left by a retreating glacier)– Autotrophic bacteria → lichens → mosses →
grasses → shrubs → trees• Secondary succession occurs in existing
communities that have been disturbed by fire, logging, or farming, but the soil remains intact– Herbaceous species → woody shrubs → trees
Biodiversity• A communities biodiversity correlates
with its size and geographic location• Biodiversity is greatest in the tropics and
on large versus smaller islands• Biodiversity is measured by both species
richness and relative abundance• Ecologists measure biodiversity as
heterogeneity, which considers both diversity factors: richness and relative abundance
Biodiversity• Tropical habitats
support much larger numbers of species of plants, animals, and other organisms than do temperate and polar regions
Biodiversity• A species–area curve illustrates the
correlation between the size of a community and the number of species found there