Classical vs Levantine Neanderthals SLIDES
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Transcript of Classical vs Levantine Neanderthals SLIDES
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“CLASSIC” EUROPEANVS
“LEVANTINE” NEAR EASTERN NEANDERTHALS
Noelle Tankard
For Emergence of Modern Humans
University of Bristol, Dec 2010
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Neanderthal type specimen
• found in Feldhoffer grotto, Neander Valley, 1857
• later excavations found associated zygomaticomaxillary fragment
Image from American Museum of Natural History, Ian Tattersall
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Our understanding of Neanderthals
• Initially seen as a transitional stage through which humans passed
• primitive features emphasized, compared to chimpanzees
• Fossils found in other regions often labelled “Neanderthaloid” as a generic pre-Homo sapiens category in possession of prominent brow-ridges and low-vaulted brain cases
•“African Neanderthal” : Broken Hill skull, Zambia•“Eastern Neanderthals” : Solo skulls from Java
(Trinkaus 1979)
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... and with time...• less biased interpretations grew as body of data increased
• definition based on from Western Europe; now consider the “classical” or European Neanderthals
•By early 80s, commonly seen as subspecies of H. sapiens
Homo sapiens neanderthalensis
• Early genetic studies influenced trend to reconsider phylogenetic place relative to H. sapiens; differences from modern H. sap emphasized
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• “… one can systematically examine the large corpus of Neanderthal fossils… [with] present knowledge of the anatomical functions of bone and muscle… against a fuller chronological background… the picture that emerges is quite clear: a human population complex with a special pattern of anatomical features that extends without interruption from Gibraltar across Europe into the Near East and Western Asia.”
(Trinkaus 1979, 91)
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Classic Neanderthals
La Ferrassie, France: considered typical of “Classic” Neanderthals – also among largest in Europe
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The Neanderthal “clade”• Whether species or subspecies, clear morphological group with characteristic traits
• “Classical Neanderthals” clearly distinct from: present day modern H. Sapiens contemporary African H. Sapiens European Upper Palaeolithic populations
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Diagnostic traits of Neanderthals...
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Neanderthal range & specimens• “Classical” Western Europe
– France, Germany, Belgium, Netherlands, Spain– specimens in Italy and Greece sometimes referred to as “preNeanderthals”
• Eastern Europe– Croatia, Czech Republic, Ukraine, Slovenia, Slovakia
•“Levantine” and Near East– tend to be younger than “Classical”– also younger than H. sap in region
• Central / Western Asia–Uzbekistan Teshik Tash, Uzbekistan
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“Levantine” and Near Eastern Sites
• Term “Levantine” more common in non-English publications
– strictly: Israel, Palestine and Jordan while Near East stretches from Nile to northern Iran– Those who distinguish Levantine from other Near Eastern with intention to note Levant as “entry point” (from Europe) for Neanderthal migration “deeper” into the Near East (Vandermeesch 2007)
• Tabun, Amud, Kebara (Israel)• Shanidar (Iraq)• Dederiyeh (Syria)
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Interest in the Levant…• Long recognized as most likely place to find evidence of
Neanderthal and H. sap interaction – behavioural and/or genetic
• Chronologically, H. sap populations older than Neanderthal• Therefore, most analysis and study aimed at Levantine
Neanderthals vs Levantine H. sapiens– Skhul and Qafzeh: debate over classification of fossils found, number of species represented
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Shanidar
Shanidar 1
- 9 skeletons
Shanidar 135-40 yrs old at death
survived serious injuriesexcavated 50s-60s
ca 40-79 ky, but dates questionable
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Amud 1
- 1.8 m tall, 1740 cc
-Approx 25 yrs at death
-ESR on mammal tooth at 40-50 kyr
-Affinity to Shanidar
-“Mosaic” features
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Kebara1.7 m
25-35 yr at death
ca 60 ka
Possible burial
Similar to Amud, Shanidar,and Tabun but more robust
Hyoid bone
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Tabun
• C2 Mandible– Ambiguous, but Neanderthal
affinities• Tabun C, partial skeleton of
female – Possibly oldest Near Eastern
Neanderthal– Excavated 1932; stratigraphic
position controversial and dating unclear
– Skull fractured
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Dederiyeh
• Dederiyeh 1– Infant– ca 60 ky
Images from Akazawa’s website, reconstruction of find and “resuscitation” of find
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Computer model of maturation ofDederiyeh infant (to adult based on Amud 1)
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Vandermeesch 2007
• Majority of “Classic” traits – although not all to the same “degree of development”
• Several features characterized as “entering the range of modern human variation”– Higher skull– Transverse contour less oval
• Suprainiac fossa larger, less deep• Less occipital bunning than “Classical”
– occipital less convex and occipital torus less prominent– Overall “less stretched”
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• Other features described as “intermediate” between “Classic” Neanderthal and mod H. sap
Less facial prognathism
Cranial capacityvaries widely
Males taller
From Trinkaus 1979
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• Supraorbital torus• Occipital bunning• Facial prognathism• Long low brain case• Larger cranial capacity• Receding frontal• Suprainiac fossa• retromolar space
….. And others
•Not all of these traits appear in the same degree and same combination
•Homologous or derived?
•Absolute or continuum?
•Same “effect” can occur as a result of various “processes”: occipital bunning (Gunz 2007)
}…. Traits to distinguish from a H. sap
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Further questions• What is the significance of these differences?
• How to explain them?– Temporal variation: co-evolution into
Neanderthal form separate from “Classical”: • Earlier split from pre-Neanderthal population?
– Geographical variation?• Reversal of “cold climate” traits?
– Hybridization with moderns?
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• Are the Levantine Neanderthals more Homo sapiens – or simply less “Classical”?
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Comparative analyses with Levantine H. sapiens specimens….
... fail to demonstrate clear and definitive difference between Neanderthals and “moderns”
Kramer 2001: tested null hypothesis of two clades in Levant (Amud/Tabun vs Qafzeh/Skhul)
Wolpoff 2001: failed to disprove null hypothesis that taxonomy did not explain variation of sample overall BUT that variation within Qafzeh/Skhul was greater than variation comparing that sample to Levantine Neanderthals
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Issues
• Traits generally discussed as if a complete package, the Neanderthal “pattern” - specimens lacking all or some often dismissed as unclassifiable
• We are over-simplifying the situation by deliberately disregarding that which does not fit a perhaps arbitrary prefabricated designation
• How much variation within what level of taxonomic clade?