Annotated Checklist of Vascular...
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National Park ServiceU.S. Department of the Interior
Natural Resource Program Center
Annotated Checklist of Vascular FloraPipe Spring National Monument
Natural Resource Technical Report NPS/NCPN/NRTR–2008/131
ON THE COVER
Wetland below West Cabin, Pipe Spring National Monument, Arizona. Photo by Walter Fertig.
Annotated Checklist of Vascular FloraPipe Spring National Monument
Natural Resource Technical Report NPS/NCPN/NRTR–2008/131
AuthorsWalter Fertig Moenave Botanical Consulting1117 W. Grand Canyon Dr.Kanab, UT 84741
Jason AlexanderCurator, UVU HerbariumUtah Valley University800 West University ParkwayOrem, UT 84058
Editing and DesignAlice Wondrak BielNorthern Colorado Plateau NetworkP.O. Box 848Moab, UT 84532
October 2008
U.S. Department of the InteriorNational Park ServiceNatural Resource Program CenterFort Collins, Colorado
ii Annotated Checklist of Vascular Flora: Pipe Spring National Monument
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Please cite this publication as:Fertig, W., and J. Alexander. 2008. Annotated checklist of vascular fl ora: Pipe Spring National Monument. Natural Resource Technical Report NPS/NCPN/NRTR–2008/131. National Park Service, Fort Collins, Colorado.
NPS D-222, October 2008
Contents iii
Contents
Figures ...........................................................................................................................................vTables ...........................................................................................................................................viiAbstract ........................................................................................................................................ ixAcknowledgements .....................................................................................................................xi
1 Introduction .............................................................................................................................11.1 Background and justifi cation .......................................................................................................11.2 Objectives ......................................................................................................................................11.3 Study area .....................................................................................................................................1
1.3.1 History and setting ........................................................................................................................ 11.3.2 Geology ......................................................................................................................................... 21.3.3 Vegetation ..................................................................................................................................... 21.3.4 Previous botanical studies ............................................................................................................ 4
2 Methods .....................................................................................................................................7
3 Results ........................................................................................................................................93.1 Summary of the fl ora of Pipe Spring National Monument .......................................................93.2 Life form and biogeographic diversity ......................................................................................103.3 Plants of special concern ............................................................................................................103.4 Noxious weeds ............................................................................................................................11
4 Discussion ................................................................................................................................13
5 Literature Cited .......................................................................................................................15
6 Appendices ..............................................................................................................................17Appendix A. Confi rmed, historical, and reported vascular plant taxa of Pipe Spring National Monument, organized by family and scientifi c name with taxonomic and biogeographic notes. .20Appendix B. Potential (unconfi rmed) taxa of Pipe Spring National Monument. .........................44Appendix C. Rejected (falsely reported or questionable) taxa of Pipe Spring National Monument. ........................................................................................................................................47Appendix D. Confi rmed, historical, and reported taxa of Pipe Spring National Monument organized by life form and common name with ecological notes. ...............................................48
iv Annotated Checklist of Vascular Flora: Pipe Spring National Monument
Contents v
FiguresFigure 1.3.1. Location of Pipe Spring National Monument, Mohave County, Arizona. Map created by
Aneth Wight of NCPN (derived from Evenden et al. 2002). ............................................................... 3
Figure 1.3.4. Increase in the number of plant taxa confi rmed as present in Pipe Spring National Monument by decade, 1930–2008. ...................................................................................................... 6
vi Annotated Checklist of Vascular Flora: Pipe Spring National Monument
Contents vii
TablesTable 1.3.4. Chronology of plant-collecting efforts in Pipe Spring National Monument. ............................... 5
Table 3.1. Statistical summary of the fl ora of Pipe Spring National Monument. ............................................ 9
Table 3.3. Plant species of conservation concern from Pipe Spring National Monument. ........................... 11
viii Annotated Checklist of Vascular Flora: Pipe Spring National Monument
Contents ix
Abstract
In 2005–2006, the National Park Service’s Northern Colorado Plateau Network received funding to develop a revised checklist of the vascular plant fl ora of Pipe Spring National Monument. This entailed a thorough review of existing literature, re-examination of specimens from the Pipe Spring National Monument herbarium and other state and national museum collections, and additional fi eld work to corroborate unverifi ed species reports or locate new species. Based on this study, 277 vascu-lar plant taxa have been confi rmed or reliably reported as occuring in the monument (215 are known from validated herbarium specimens and 62 from unvouchered literature reports). Another 15 species formerly considered part of the monument fl ora have now been shown to be falsely reported. The current fl ora of Pipe Spring National Monument represents 6.5% of the known fl ora of Arizona and 11% of the fl ora of northern Arizona and the Arizona Strip. Although no federally listed threatened or endangered species are known from the monument, two occur in the vicinity. Three plant species from Pipe Spring National Monument are currently tracked as species of concern by the State of Arizona’s Natural Heritage Program. Introduced plants make up nearly 21% of the monument fl ora (58 species), of which only 4 species are considered noxious weeds by the State of Arizona. Further surveys in Pipe Spring National Monument are likely to fi nd additional new species (46 are known from similar habi-tats outside the monument boundary), especially among late-season fl owering, weedy, or wetland taxa.
x Annotated Checklist of Vascular Flora: Pipe Spring National Monument
Contents xi
Acknowledgements
We would like to thank the following individuals for their assistance with this project. Botanist Dr. Angela Evenden, former leader of the Northern Colorado Plateau Network in Moab, Utah, was an active proponent of this project and helped secure funding for fi eld work and herbarium studies in 2006. John Hiscock and Andrea Bornemeier provided data from monument archives and assisted with research permits. Cheryl Decker and Denise Louie, of Zion National Park, assisted with fi eld surveys of the monument in 2001. Heather Karnes and An Glorieux helped with fi eld work and data entry in 2004 and 2005, respectively. Libby Nance, database specialist with the NCPN, assisted with development and quality control of the monument’s NPSpecies database. Libby Nance, Janet Coles, and Helen Thomas also provided data from recent vegetation mapping and wetland inventories on the monument. Drs. Stanley Welsh and Duane Atwood provided access to herbarium collections at Brigham Young University. Dr. Michael Piep provided access to collections at the Utah State University (Intermountain) herbarium. Dr. Ron Hartman and Ernie Nelson provided access and specimen loans from the Rocky Mountain Herbarium at the University of Wyoming. Thanks to the technical reviewers (Angela Evenden, Dr. Dusty Perkins, and Libby Nance) for their comments and suggestions. To any others whom we may have inadvertently forgotten, please accept our apologies (and thanks).
Research was conducted under permit # PISP-2006-SCI-0002 for study number PISP-00005. Funding was provided by the National Park Service through an agreement with the Rocky Mountains Coopera-tive Ecosystem Studies Unit and the University of Wyoming.
xii Annotated Checklist of Vascular Flora: Pipe Spring National Monument
Chapter 1: Introduction 1
1 Introduction In 2006, the National Park Service (NPS) estab-lished a cooperative agreement with the Univer-sity of Wyoming, through the Rocky Mountains Cooperative Ecosystem Studies Unit, to produce a series of technical reports summarizing the vas-cular plant fl ora of each of the 16 national park units of the Northern Colorado Plateau Network (NCPN). These reports are based on existing in-formation in the park service’s NPSpecies data-base system and augmented by further data min-ing, reviews of state and regional herbaria, and new fi eld surveys to resolve the status of many reported and unconfi rmed species. The objec-tive of these reports is to summarize the current state of knowledge of the fl ora of each park for the benefi t of NPS biologists and interpretive staff , outside researchers, and the general public. The following report addresses the vascular fl ora of Pipe Spring National Monument, in northern Arizona. Other NCPN park units are addressed in separate reports.
1.1 Background and justifi cationOne of the primary missions of NPS is “to con-serve unimpaired the natural and cultural resourc-es and values of the national park system for the enjoyment of this and future generations” (NPS 2000). Due to their high degree of protection and distribution across a wide variety of habitat types, the nation’s national park units make a signifi -cant contribution to the protection of biological diversity (Noss and Cooperrider 1994). Unfortu-nately, the full contribution of the NPS system is not known, as many park units have traditionally lacked comprehensive biological inventories or monitoring programs to identify the composition and condition of their biota. Knowing the status of park fl ora and fauna is critical for eff ective park management and for systematically identifying and fi lling gaps in the nation’s protected area net-work (Margules and Pressey 2000).
To rectify this shortcoming, Congress passed the National Parks Omnibus Management Act in 1998 to provide guidance and funding for an inventory and monitoring program across the entire Na-tional Park System. Since 2000, the NCPN Inven-tory and Monitoring Program has been develop-ing and verifying species lists of vertebrates and vascular plants for each park unit in the network
(Evenden et al. 2002). This eff ort has entailed re-viewing existing species checklists, corroborating the identifi cation of specimens in park museum and herbarium collections, locating additional specimen records from other museums, mining relevant literature and datasets, and conducting fi eld surveys to locate new taxa1 or confi rm un-substantiated reports. All of these data have been synthesized in NPSpecies, the park service’s stan-dardized database for documenting park status (present, probably present, unconfi rmed, falsely reported), abundance, residency, nativity, and rarity. With the production of a technical series of reports on the fl ora of each park, the NPSpecies data will now be available to a broader audience.
1.2 ObjectivesThe objectives of this project are to:
1. Document the current status (present, reported, potential, or falsely reported) of all vascular plant species that have been cited as occurring in Pipe Spring National Monument based on a review of herbarium collections and past literature; and
2. Develop an annotated checklist of the fl ora of Pipe Spring National Monument with supplemental information on taxonomic synonyms, distribution, within-park abun-dance, fl owering period, growth form, nativ-ity, habitat, and sources for each record.
1.3 Study area
1.3.1 History and setting
As one of the few reliable water sources in a large desert area, Pipe Spring has been a focal area of human activity for many centuries. Prior to about 300 B.C., the Arizona Strip was inhabited by semi-nomadic hunter-gatherers who left little physical evidence behind. Over the next thousand years, the region was inhabited by Ancestral Puebloan peoples who engaged in agriculture and hunting and produced abundant artifacts, including pit-house sites, woven baskets, pottery sherds, and arrowheads. Prolonged drought was one likely factor that brought an end to the Puebloan cul-ture in the local area by 1250 A.D. Subsequent inhabitants of the Arizona Strip were semi-no-madic Southern Paiute bands; the Kaibab Paiute occupied the area at the time of contact with
1Taxa refer to all named species, subspecies, and varieties.
2 Annotated Checklist of Vascular Flora: Pipe Spring National Monument
European cultures (McKoy 2000, Leach 1999).
Mormon missionary Jacob Hamblin passed through Pipe Spring in 1858; fi ve years later, St. George, Utah, resident James Whitmore estab-lished a cattle ranch there. In January 1866, Whit-more was murdered during a trip to retrieve live-stock driven off by an Indian party (Leach 1999, McKoy 2000). The Church of Jesus Christ of Latter-Day Saints acquired the property in 1870, constructed a fort (Winsor Castle) that controlled the outlet of Pipe Spring, and managed the site as a tithing ranch. During the 1870s, the native desert grasslands were subjected to grazing by more than 2,500 head of cattle and 11,000 sheep (Alexander 1998a). The ranch passed into private ownership in 1895, and remained as such with the establish-ment of the Kaibab Paiute Indian Reservation in 1909 (McKoy 2000).
While traveling to Zion National Park in 1920, National Park Service Director Stephen Mather hatched the idea of preserving the site as a nation-al monument. With the support of local boosters anticipating improved road access and tourism income, President Warren Harding established Pipe Spring National Monument in May 1923, to “serve as a memorial of western pioneer life.” (McKoy 2000). The monument’s mission has since been expanded to also convey American In-dian culture and history and to protect the spring and associated natural environment (Evenden et al. 2002).
Pipe Spring National Monument is located about 15 miles west of Fredonia in eastern Mohave County, Arizona (Figure 1.3.1). The monument is situated on the northern end of the Uinkaret Plateau, below the Vermilion Cliff s within the Ari-zona Strip (the northern portion of the state cut off by the Grand Canyon). The entire monument covers 40 acres (16 ha) and is embedded within the Kaibab Paiute Indian Reservation. Elevation within Pipe Spring National Monument ranges from 4,923 to 5,100 feet (1,500–1,555 m). Winter temperatures vary from 0 to 40°F, while summer temperatures reach highs of 90–115°F (Evenden et al. 2002).
1.3.2 Geology
The sandstone ridge to the northwest of Win-sor Castle is comprised of Jurassic-age Navajo Sandstone deposited as part of a Sahara-like dune system that extended from Nevada to Wyoming approximately 180 million years ago. The Navajo
formation overlies Kayenta strata derived from early Jurassic river sediments. Navajo Sandstone is more permeable than the Kayenta, and the con-tact between the two formations is notable as a source of springs throughout the Colorado Pla-teau. The juxtaposition of this contact zone with the Sevier Fault (separating the mesa from the adjacent Moenkopi and Chinle beds to the east) accounts for the size and fl ow of the three springs in the monument (Pipe Spring, itself, and the smaller Tunnel and Cabin springs). Holocene-age alluvium covers the eastern and southern ends of the monument, burying the Triassic Chinle and Moenkopi formations (Billingsley 2004).
1.3.3 Vegetation
Unfortunately, the pre-settlement vegetation of Pipe Spring and the Arizona Strip was never stud-ied in detail using modern scientifi c methods. Anecdotal reports by local ranchers describe the Strip as a “sea of grass” in the late 1860s (Alex-ander 1998a). In 1882, geologist Charles Dut-ton wrote that the Pipe Spring area had changed from being “covered with abundant grasses, af-fording rich pasturage to horses and cattle” to a place where “hardly a blade of grass is to be found within ten miles of the spring, unless upon the crags and mesas of the Vermilion Cliff s behind it” in just 10 years (Dutton 1882, cited in Alexander 1998a). Based on historic diaries, archives, and photographs, Alexander (1998a) reported that the mesa northwest of Pipe Spring was devoid of trees by the time NPS took over management in 1923, and the fl ats surrounding the fort were mostly bare ground.
Vegetation has slowly recovered in the monu-ment. Today, three main plant communities can be recognized. Sandy fl ats and south-facing mesa slopes are inhabited by desert shrub vegetation, dominated by greenish rabbitbrush (Chrysotham-nus nauseosus), fourwing saltbush (Atriplex ca-nescens), and sand sagebrush (Artemisia fi lifolia) (Alexander 1998b). Steeper Navajo Sandstone slopes are dominated by open Utah juniper (Juni-perus osteosperma) and two-needle pinyon (Pinus edulis) woodlands with an understory of turbi-nella live oak (Quercus turbinella) and Utah ser-viceberry (Amelanchier utahensis). Wet meadow vegetation consisting of baltic rush (Juncus arcti-cus), rabbitbrush, and scratchgrass (Muhlenbergia asperifolia) occurs below Cabin Spring (by West Cabin). A human-made pond, located just out-side Winsor Castle, is dominated by planted white poplar (Populus alba), Fremont cottonwood (P.
Chapter 1: Introduction 3
Figure 1.3.1. Location of Pipe Spring National Monument, Mohave County, Arizona. Map created by Aneth Wight of NCPN (derived from Evenden et al. 2002).
To St. George, UT
Visitor Center
To Fredonia, AZ389
Established May 31, 19231,500−1,555 meters elevation(4,923−5,100 feet elevation)
16 hectares (40 acres)0 0.1 0.2 0.3
Kilometers
0 0.1 0.2 0.3Miles
Legend:
Intermittent Streams
Paved Road
Unpaved Road
Unimproved/4WD Road
Park Boundary (NPS)
Kaibab Reservation (IR)
Pipe Spring National MonumentUtah
4 Annotated Checklist of Vascular Flora: Pipe Spring National Monument
fremontii), Carolina poplar (P. x canadensis), Si-berian elm (Ulmus pumila), and tree-of-heaven (Ailanthus altissima). Much of the remainder of the grounds is dedicated to cultivated orchards and a replica pioneer garden.
1.3.4 Previous botanical studies
The fi rst botanist to visit Pipe Spring was proba-bly Ellen Powell Thompson, sister of John Wesley Powell and resident of Kanab, Utah, in the early 1870s (Alexander 2004). It is not known wheth-er Thompson collected specimens at the future monument. Marcus E. Jones investigated the fl ora of Kanab and the Arizona Strip in the 1880s, but it is also unknown whether he specifi cally collect-ed at Pipe Spring. Albert Siler, a pioneer rancher and naturalist in southern Utah, collected the type specimen of Utahia sileri (later renamed Pediocactus sileri), Siler’s pincushion cactus, from “Cottonwood Springs and Pipe Springs, southern Utah” (actually Arizona) in 1883 (Benson 1982). This collection is now believed to have come from just outside the monument on the Kaibab Paiute Indian Reservation, as the exacting habitat condi-tions required by this rare cactus are not present within monument boundaries.
The holotype of Pipe Spring pricklypear (Opun-tia erinacea var. aurea, O. basilaris var. aurea, or O. aurea, depending on one’s taxonomic pref-erence) was collected “1/2 mile north of Pipe Spring” by McCabe and McCabe in 1930 (Ben-son 1982). This locality is either within or barely outside the monument’s boundary, but the spe-cies is well documented elsewhere in Pipe Spring National Monument. If accepted as being col-lected in the monument, the McCabe collection would stand as the earliest known specimen from the area (Table 1.3.4, Figure 1.3.4).
Leonard Heaton, custodian of Pipe Spring Na-tional Monument (PISP), began collecting plants for the PISP herbarium in 1934. Over the next seven years, Heaton and J. Whitehead collected 112 specimens for the herbarium, representing 98 species. Heaton (1935) compiled the fi rst check-list of monument fl ora in 1935, citing 42 species and estimating that another 100 remained to be
discovered (in hindsight, a reasonable prediction). Robert Peebles (co-author of Arizona Flora) and A. R. Purchase added six new species in 1936, and later found two more in a 1940 visit—including the fi rst report of Opuntia pinkavae (Pinkava’s pricklypear), not recognized as a new species to science until 1997. Utah botanist Walter Cottam and Kathy Parker also collected a few new taxa in the 1940s.
Over the next quarter century, relatively little bo-tanical work was done in the monument, except for some sporadic study in the early to mid-1960s. Ruth Nelson visited Pipe Spring twice starting in 1971, and added fi ve new species to the fl ora. Richard King, a range conservationist with the Fredonia offi ce of the Soil Conservation Service, compiled a species list in 1977 that included 82 plant species (with an emphasis on shrubs and grass species) (King 1977). King’s list contained 39 new species for the monument, of which all but 18 have since been confi rmed with vouchers.
The most intensive botanical surveys of Pipe Spring National Monument since the mid-1930s have taken place in the last decade. Zion National Park biologists Karen Mason and Bill LaBarre, who conducted a noxious-weed inventory in 1997, reported 13 new species for the monument (Kurth 1998). In 1998, Jason Alexander was hired by the NPS to research vegetation change within the monument and compile a revised species list. Alexander (1998b) documented 166 taxa for Pipe Spring National Monument, of which 63 repre-sented new species. In 2001, Alexander, Denise Louie of Zion National Park, and Angie Even-den of the NCPN staff in Moab, Utah, collected 116 vouchers for the PISP herbarium (more than doubling it in size) and recorded 22 new species for the fl ora. This team also verifi ed 60 previous literature reports with specimen vouchers (Table 1.3.4). More recently, we, along with Janet Coles of the NCPN vegetation staff , found 10 new spe-cies for the monument in fall 2007, and relocated 18 more species known only from historical or unvouchered reports.
Additional details on plant-collecting eff orts at PISP can be found in Table 1.3.4.
Chapter 1: Introduction 5
Table 1.3.4. Chronology of plant-collecting efforts in Pipe Spring National Monument.
Year Collector(s)# of new
taxa Comments
1883 A. L. Siler 0 Albert L. Siler collected the holotype of Pediocactus sileri from “Cottonwood Springs and Pipe Springs, southern Utah” [Arizona] in 1883. Based on the specifi c habitat requirements of this species, it is now believed that the collection came from the adjacent Kaibab Paiute Indian Reservation rather than Pipe Spring National Monument, itself.
1930 McCabe and McCabe
1 McCabe and McCabe collected the holotype of Opuntia erinacea var. aurea “1/2 mile north of Pipe Spring” within or very near Pipe Spring National Monument.
1934–1939 L. Heaton, J. Whitehead, A. R. Purchase, and R. H. Peebles
98 + 7 reported
Leonard Heaton, custodian of Pipe Spring National Monument, compiled a vascular plant species list with 42 species in 1935 and noted “there are about 100 more plants that have not been [n]amed or classifi ed...” (Heaton 1935). All but 7 of these species have been subsequently vouchered. From 1934–1939, Heaton and J. Whitehead collected 104 voucher specimens for the PISP herbarium and documented 92 plant taxa. In 1936, Robert H. Peebles discovered 4 new species for the monument and A.R. Purchase added 2 new grass species.
1940–1946 R. H. Peebles, W. Cottam, L. Heaton, and K. F. Parker
9 R. H. Peebles discovered 2 new species for PISP in 1940, including the fi rst record of Opuntia pinkavae (later described as a new taxon by Bruce Parfi tt in 1997). Walter Cottam and Leonard Heaton found 6 new taxa in 1941, including the earliest report of Lepidium montanum var. stellae (Stella’s pepperwort; later described as a new taxon by Stan Welsh in 1978). Kathy Parker discovered Convolvulus arvensis (fi eld bindweed) in the monument in 1946.
1961 F. Fanning 1 F. Fanning collected Amsonia jonesii (Jones’ amsonia) in the monument (specimen at the Desert Botanical Garden herbarium).
1965–1967 E. Lehto and M. Ganz
5 Ganz and Lehto made at least 16 collections at Pipe Spring (deposited at Arizona State University, University of Arizona, and Northern Arizona University).
1971–1973 R. A. Nelson and M. G. MacLeod
5 Ruth Nelson and M. G. MacLeod made collections at PISP during visits in 1971 and 1973 (specimens are deposited at Arizona State University and the University of Wyoming).
1977 R. J. King 21 + 18 reported
Richard King, range conservationist with the Soil Conservation Service in Fredonia, Arizona, compiled an annotated checklist of 82 species for the monument (King 1977), including 39 that had not previously been reported. In subsequent years, 21 of King’s new taxa were documented with vouchers.
1995 A. M. Phillips III 1 Arthur Phillips made 9 collections in the monument, including the fi rst of Rhus aromatica var. simplicifolia (squawbush).
1997 K. Mason and B. LaBarre
8 + 5 reported
Zion NP biological science technicians Karen Mason and Bill LaBarre conducted a weed survey of PISP, reporting 13 new species for the area (Kurth 1998), of which 8 have been subsequently vouchered.
1998 J. Alexander 34 + 29 reported
Alexander (1998b) examined historic records and conducted a ground survey of the monument to compile a revised checklist of 166 vascular plant taxa for PISP. This list included 63 new species, of which 33 were later vouchered (29 are still known only from literature reports).
2001 J. Alexander, D. Louie, and A. Evenden
22 Jason Alexander, Denise Louie, and Angie Evenden collected 116 voucher specimens (doubling the size of the PISP herbarium) and documented 22 new species as well as making vouchers of over 60 previously reported taxa.
2007 W. Fertig and J. Coles
10 Walter Fertig and Janet Coles conducted surveys and vegetation sampling and added 33 specimens for the PISP herbarium, including 10 new to the monument and 18 that were previously considered historical or had been reported without a voucher.
6 Annotated Checklist of Vascular Flora: Pipe Spring National Monument
Figure 1.3.4. Increase in the number of plant taxa confi rmed as present in Pipe Spring National Monument by decade, 1930–2008.
Nu
mb
er o
f va
scu
lar
pla
nt
taxa
250
200
150
100
50
01930 1940 1950 1960 1970 1980 1990 2000 2008
Year
Chapter 2: Methods 7
2 MethodsUnder contract with the National Park Service, Jason Alexander reviewed all mounted speci-mens in the PISP herbarium in 2004 to determine whether each was correctly identifi ed to species, subspecies, or variety following the nomenclature of Welsh et al. (2003). As necessary, he made cor-rections and annotations to each specimen and to the voucher records in PISP’s Automated Nation-al Cataloging System (ANCS+) and NPSpecies database. Based on Alexander’s herbarium work, we developed a revised checklist of species that were present in the monument (confi rmed by an authenticated voucher) and those that were re-ported in the literature but not corroborated with a voucher (equivalent to “probably present” in NPSpecies terminology). Species that were now known to be misidentifi ed or highly questionable were moved to a third list of rejected taxa. Finally, we developed a list of potential species (“uncon-fi rmed” in NPspecies) known from the vicinity of Pipe Spring National Monument using an unpub-lished list generated by Dr. N. Duane Atwood of Brigham Young University.
Each list was annotated with supplemental infor-mation on life form, geographic range (within the context of Arizona), population size in the monu-ment (derived from Jason Alexander’s NPSpecies dataset for PISP and fi eld observations), fl ower-ing period, distribution across general vegetation types, nativity, data source (collector and collec-tion number from a sample specimen or literature citation), year of earliest collection (from herbar-
ium records), and relevant synonymy and taxo-nomic problems (based on review of Barkworth et al. 2003, 2007, Barneby 1989, Cronquist 1994, Cronquist et al. 1972, 1977, 1984, 1997, Dorn 2001, Flora of North America Editorial Commit-tee 1993, 1997, 2000, 2002a, 2002b, 2003, 2005, 2006a, 2006b, 2006c, Holmgren et al. 2005, Mc-Dougall 1973, and Weber and Wittmann 2001). The annotations to these lists and label data from the PISP herbarium review were then incorpo-rated into the NPSpecies database for the monu-ment.
Work in 2007 focused on confi rmation of report-ed and potential species for the monument fl ora through fi eld surveys and review of collections from state and regional herbaria. We made one collecting trip to PISP, in October 2007, to target missing species, emphasizing under-sampled wet-land, weedy, and late-fl owering taxa. Herbarium studies were undertaken at Brigham Young Uni-versity, Utah State University, and the University of Wyoming in March, June, and October 2006, and February and March 2007. We also docu-mented monument specimens from the digital herbarium database of Arizona State University (http://seinet.asu.edu/seinet/collections/selec-tion.jsp?cat=plantae) and its links to comparable databases for the Desert Botanical Garden, Uni-versity of Arizona, and Northern Arizona Univer-sity. Finally, we conducted an internet search of the virtual herbarium database of the New York Botanical Garden (http://sciweb.nybg.org/sci-ence2/hcol/intf/index.asp) and examined plot data from the NCPN vegetation mapping crew (NCPN, in ed.).
8 Annotated Checklist of Vascular Flora: Pipe Spring National Monument
Chapter 3: Results 9
3 Results
3.1 Summary of the fl ora of Pipe Spring National Monument
In 2004, Jason Alexander completed a thorough review of the 227 vascular plant vouchers in the PISP herbarium (only 112 of which were col-lected prior to 2001). He found that 8 specimens were misidentifi ed (3.5%), 3 needed to be revised to a more current name (1.3%), 27 needed vari-ety or subspecies names added (11.9%), and 189 were correctly determined (83.3%). Based solely on these collections, the confi rmed fl ora of Pipe Spring National Monument stood at 162 taxa. From 2005 to 2007, we located an additional 27 taxa from collections at the University of Wyo-ming’s Rocky Mountain Herbarium and speci-
mens found through a database search of the Arizona State University digital database. Field surveys in 2007 netted another 26 new species (including 16 that were previously reported with-out a voucher). Overall, the documented fl ora of Pipe Spring National Monument has increased by 53 species (24.7%), bringing the total number of vouchered taxa to 215 (Table 3.1, Appendices A, D).
Currently, 62 additional taxa have been reported for PISP but have yet to be corroborated with a voucher specimen (Table 3.1, Appendix A). Of these, 7 were reported by Heaton (1935), 18 by King (1977), 5 by Kurth (1998), and 32 by Alexan-der (1998b). Taken together, the number of con-fi rmed and reported vascular plant taxa in Pipe Spring National Monument stands at 277 taxa (Table 3.1, Appendix A).
Table 3.1. Statistical summary of the fl ora of Pipe Spring National Monument.
CategoryNumber
confi rmed in park
Number additionally
reported for parkTotal
Taxonomic diversity
Total taxa (including varieties and subspecies)
215 62 277
Full species (excluding varieties and subspecies)
203 59 262
Families 54 2 56
Life form diversity
Tree taxa 8 2 10
Shrub taxa 34 6 40
Perennial forb taxa 72 26 98
Annual forb taxa 72 9 81
Perennial graminoid taxa 21 15 36
Annual graminoid taxa 8 4 12
Fern taxa 0 0 0
Biogeographic diversity
Introduced taxa 35 23 58
Native taxa
Locally endemic taxa 4 0 4
Regionally endemic taxa 14 7 21
Disjunct taxa 0 0 0
Peripheral taxa 4 1 5
Sparse taxa 1 0 1
Widespread taxa 157 31 188
Total native taxa 180 39 219The number of taxa and families is based on taxonomic concepts of Welsh et al. (2003). See Appendices, “Codes and Defi nitions”, for an explanation of the various categories.
10 Annotated Checklist of Vascular Flora: Pipe Spring National Monument
Entering the 2007 fi eld season, 41 taxa were rec-ognized as potentially occurring in PISP based on the proximity of the monument to known populations (Alexander NPSpecies database). Including fi ve additional species identifi ed by Duane Atwood, the current number of potential species for Pipe Spring National Monument is 46 (Appendix B).
Based on his evaluation of the PISP herbarium, Jason Alexander found 8 taxa that were falsely re-ported due to misidentifi cation. We consider an additional seven species cited (but not vouchered) by Heaton (1935) and King (1977) to be question-able based on the absence of suitable habitat or the distance of Pipe Spring National Monument to the known or expected range of each species (Appendix C).
The confi rmed and reported fl ora of Pipe Spring National Monument represents 6.5% of the 4,241 native and naturalized taxa documented for the state of Arizona by Kartesz (2003) and 34.8% of the state’s 161 plant families (Table 3.1). Within Mohave County, the monument fl ora captures 17.4% of the 1,588 taxa reported by Kartesz (2003). At the ecoregional scale, Pipe Spring Na-tional Monument contains 11.0% of the 2,517 species of seed plants and pteridophytes reported for northern Arizona by McDougall (1973) and the Flora of North America Editorial Committee (1993).
3.2 Life form and biogeographic diversity
Although ecologically dominant, tree and shrub species make up only 18% of the confi rmed and reported plant taxa of Pipe Spring National Mon-ument (Table 3.1). Perennial forbs are the most species-rich life form in the monument’s fl ora, with 98 known and reported taxa (35.4% of the total fl ora). Annual forbs and grass-like plants are unusually species-rich in the monument (relative to other fl oras in the Colorado Plateau), compris-ing 33.5% of all taxa. Like trees and shrubs, pe-rennial graminoids are ecologically important but relatively species-poor, representing just 13% of the total fl ora. No ferns or fern-allies have been found in PISP (Table 3.1).
Of the monument’s 219 native taxa, 188 (85.8%) are widespread across Arizona and western North America. Only four species (1.8% of the monument’s native fl ora) have extremely limited global distributions (local endemics found within
an area of less than 1° of latitude × 2° of longi-tude; see Appendix A for complete defi nitions of biogeographic categories). Regional endemics (largely restricted to the Colorado Plateau) com-prise an additional 9.6% of the monument fl ora (Table 3.1). Species at the edge of their range (pe-ripheral) or found sparsely across the state make up just 2.8% of the total fl ora.
3.3 Plants of special concernPipe Spring National Monument has no plants listed as threatened or endangered under the U.S. Endangered Species Act (ESA). Siler’s pincush-ion cactus (Pediocactus sileri), listed as threatened under the ESA, is known from adjacent areas of the Kaibab Pauite Indian Reservation (Siler’s type specimen comes from “Pipe Spring”), but is not thought to occur within the monument itself due to a lack of suitable gypsiferous Moenkopi soils (ARPC, no date). The federally threatened Jones’ cycladenia (Cycladenia humilis var. jonesii) is also known from Chinle soils west of Pipe Spring, but is unlikely to be found in the monument.
The Arizona Game and Fish Department (AGF)’s Heritage Data Management System (2008) tracks three cactus species from PISP as species of con-cern (Table 3.3). All are listed as Salvage Restrict-ed (collection prohibited except with a permit) under the 1993 Arizona Native Plant Law (ARPC, no date). Atwood’s wild-buckwheat (Eriogonum thompsoniae var. atwoodii) and Morton wild-buckwheat (E. mortonianum) are also tracked as species of concern by AGF and known from the Kaibab Paiute Indian Reservation, but have not been documented in Pipe Spring National Monu-ment.
Two recently described species with narrow geo-graphic ranges currently lack any protective status but might warrant management attention. Stella’s pepperwort (Lepidium montanum var. stellae) is restricted to southern Utah and northern Arizona (Welsh et al. 2003), where it typically occurs on gypsum-rich, Moenkopi-derived soils. Leon-ard Heaton collected this species at or near Pipe Spring in 1941 (suitable habitat is more likely just outside the monument), but it remained uniden-tifi ed until annotated by Jason Alexander in 2004. Pinkava’s pricklypear (Opuntia pinkavae) is also restricted to the Utah–Arizona border and was fi rst recorded at Pipe Spring in 1940 by Robert Peebles. The holotype of O. pinkavae comes from Bulrush Canyon, south of Pipe Spring (Parfi tt 1997).
Chapter 3: Results 11
3.4 Noxious weedsIntroduced plant species (not historically native to Arizona or North America) make up 20.9% of the confi rmed and reported fl ora of Pipe Spring National Monument (Table 3.1). This fi gure is substantially higher than the statewide average of 12.7% non-native species (Kartesz 2003). Of the 58 introduced plant species confi rmed or re-ported for PISP (Table 3.1), only four are listed as noxious by the State of Arizona (ADAPSD 2005). Scotch thistle (Onopordum acanthium) is on the state’s prohibited and restricted noxious weed lists, while fi eld bindweed (Convolvulus arven-sis), common purslane (Portulaca oleracea), and puncturevine (Tribulus terrestris) are listed as prohibited and regulated noxious weeds.
Kurth (1998) cited 34 non-native species of man-agement concern, based on a 1997 survey of the monument by Karen Mason and Bill LaBarre of Zion National Park. These consisted primar-ily of introduced ornamental trees that have be-come weedy or exotic forbs and grasses that are common agricultural or rangeland pests. Using a matrix of signifi cance of impact and feasibility of control, Kurth classifi ed these species into four groups: low threat/easy to control, low threat/hard to control, serious threat/easy to control, and serious threat/hard to control. Four species were recommended for immediate control: tree-of-heaven (Ailanthus altissima), white poplar (Populus alba), tamarisk (Tamarix chinensis), and Siberian elm (Ulmus pumila).
Table 3.3. Plant species of conservation concern from Pipe Spring National Monument.
Family Species Common nameTNC global rank1 Federal legal status AZ status2
Cactaceae Opuntia erinacea var. aurea (O. basilaris var. aurea)
Pipe Spring pricklypear
G3 None Salvage Restricted, S3
Cactaceae Opuntia whipplei var. whipplei Whipple’s cholla G4?T4? None Salvage Restricted, S1
Cactaceae Sclerocactus whipplei var. roseus (S. parvifl orus ssp. intermedius)
small-fl ower fi shhook cactus
G4T3? None Salvage Restricted, S2
This list is derived from Arizona Game and Fish Department Heritage Data Management System (2008) and Arizona Rare Plant Committee (no date).1TNC global rank assesses abundance and conservation priority on a scale of 1–5 (1 being extremely vulnerable and 5 being secure) for full species (G) and varieties or subspecies (T) across their entire range. A “?” indicates uncertainty in the rank. A comparable scoring system is used to measure state abundance (S) in Arizona under the column “AZ status”.2AZ status includes protection afforded under the 1993 Arizona Native Plant Law.
12 Annotated Checklist of Vascular Flora: Pipe Spring National Monument
Chapter 4: Discussion 13
4 DiscussionBeginning with Leonard Heaton’s fi rst species list in 1935, the known fl ora of Pipe Spring National Monument has grown from 42 species to 277 taxa (Tables 1.3.4 and 3.1, Figure 1.3.4, Appendix A). Based on available habitats and proximity, at least another 46 plant taxa may occur within the monument (Appendix B). Should all of these po-tential species be found, the documented fl ora of PISP would increase by nearly 14%. Many of the “missing” potential or reported taxa consist of cryptic species that fl ower late in the growing sea-son, or occur in undersampled wetland or weedy habitats. Confi rmation of reported species should remain a high priority, as some of these taxa may ultimately prove to be erroneous. Pipe Spring also has a high percentage of plants known only from
historic records (pre-dating 1970) that should be a priority for re-location to confi rm that they are still extant. Further examination of monu-ment collections from other herbaria in Arizona or major national collections (such as Harvard University’s Gray Herbarium, the New York Bo-tanical Garden, and the Smithsonian Institution’s U.S. National Herbarium) may result in the confi -mation of additional records of historical species for the monument. Additional fi eld studies would also be useful to better refi ne the abundance and distribution of species of high management interest (especially rare plants and invasive non-natives). Surveys following wet winters would likely lead to the discovery of new annual native and exotic species. Lastly, better documentation of plant species previously or currently grown in the monument’s orchard and pioneer garden is needed.
14 Annotated Checklist of Vascular Flora: Pipe Spring National Monument
Chapter 5: Literature Cited 15
5 Literature CitedAlexander, J. 1998a. Pipe Spring National Monu-
ment presettlement vegetation literature survey. Pipe Spring National Monument, Az. 43 pp.
—— . 1998b. Pipe Spring National Monument master vegetation species list. Report pre-pared for the National Park Service, Pipe Spring National Monument.
—— . 2004. A summary of fl oristics at Pipe Spring National Monument. Pipe Spring National Monument, Az. 3 pp.
Arizona Department of Agriculture, Plant Ser-vices Division (ADAPSD). 2005. Prohibited, regulated and restricted noxious weeds.
Arizona Game and Fish Department Heritage Data Management System. 2008. Special status species by taxon, scientifi c name. http://www.azgfd.gov/w_c/edits/hdms _spe-cies_lists.shtml
Arizona Rare Plant Committee (ARPC). No date. Arizona rare plant fi eld guide: A collabora-tion of agencies and organizations.
Barkworth, M. E., K. M. Capels, S. Long, and M. B. Piep, eds. 2003. Magnoliophyta: Com-melinidae (in part): Poaceae, part 2. Flora of North America north of Mexico, volume 25. New York: Oxford University Press. 783 pp.
Barkworth, M. E., K. M. Capels, S. Long, L. K. Anderton, and M. B. Piep, eds. 2007. Mag-noliophyta: Commelinidae (in part): Poa-ceae, part 1. Flora of North America north of Mexico, volume 24. New York: Oxford University Press. 911 pp.
Barneby, R. C. 1989. Volume 3, Part B. Fabales. In Cronquist, A., A. H. Holmgren, N. H. Holmgren, J. L. Reveal, and P. K. Holmgren, eds., Intermountain fl ora: Vascular plants of the Intermountain West, U.S.A. Bronx, N.Y.: New York Botanical Garden. 279 pp.
Benson, L. 1982. The cacti of the United States and Canada. Stanford, Ca.: Stanford Univer-sity Press. 1044 pp.
Billingsley, G. H., S. S. Priest, and T. J. Felger. 2004. Geologic map of Pipe Spring National Monument and the western Kaibab-Pai-ute Indian Reservation, Mohave County, Arizona. Pamphlet to accompany Scientifi c Investigations Map 2863. U.S. Geological
Survey, U.S. Department of the Interior.
Cronquist, A. 1994. Volume 5, Asterales. In Cronquist, A., A. H. Holmgren, N. H. Holmgren, J. L. Reveal, and P. K. Holmgren, eds., Intermountain fl ora: Vascular plants of the Intermountain West, U.S.A. Bronx, N.Y.: New York Botanical Garden. 496 pp.
Cronquist, A., A. H. Holmgren, N. H. Holmgren, and J. L. Reveal, eds. 1972. Volume 1, Geo-logical and botanical history of the region, its plant geography and a glossary: The vascular cryptogams and the gymnosperms. Intermountain fl ora: Vascular plants of the Intermountain West, U.S.A. Bronx, N.Y.: New York Botanical Garden. 270 pp.
Cronquist, A., A. H. Holmgren, N. H. Hol-mgren, J. L. Reveal, and P. K. Holmgren, eds. 1977. Volume 6, The Monocotyledons. Intermountain fl ora: Vascular plants of the Intermountain West, U.S.A. Bronx, N.Y.: New York Botanical Garden. 584 pp.
—— . 1984. Volume 4, Subclass Asteridae (except Asteraceae). Intermountain fl ora: Vascular plants of the Intermountain West, U.S.A. Bronx, N.Y.: New York Botanical Garden. 573 pp.
Cronquist, A., N. H. Holmgren, and P. K. Hol-mgren, eds. 1997. Volume 3, Part A, Subclass Rosidae (except Fabales). Intermountain fl ora: Vascular plants of the Intermountain West, U.S.A. Bronx, N.Y.: New York Botani-cal Garden. 446 pp.
Dorn, R. D. 2001. Vascular plants of Wyoming. Third edition. Cheyenne, Wy.: Mountain West Publishing. 412 pp.
Evenden, A., M. Miller, M. Beer, E. Nance, S. Daw, A. Wight, M. Estenson, and L. Cudlip. 2002. Northern Colorado Plateau Vital Signs Network and Prototype Cluster, plan for natural resources monitoring: Phase I Report [two volumes]. National Park Ser-vice, Northern Colorado Plateau Network, Moab, Ut. 138 pp + app.
Flora of North America Editorial Committee. 1993. Pteridophytes and gymnosperms. Flora of North America north of Mexico, Volume 2. New York: Oxford University Press. 475 pp.
—— . 1997. Magnoliophyta: Magnoliidae and Hamamelidae. Flora of North America north of Mexico, Volume 2. New York: Ox-
16 Annotated Checklist of Vascular Flora: Pipe Spring National Monument
ford University Press. 590 pp.
—— . 2000. Magnoliophyta: Alismatidae, Are-cidae, Commelinidae (in part), and Zingi-beridae. Flora of North America north of Mexico, Volume 2. New York: Oxford University Press. 352 pp.
—— . 2002a. Magnoliophyta: Commelinidae (in part): Cyperaceae. Flora of North America north of Mexico, Volume 2. New York: Ox-ford University Press. 608 pp.
—— . 2002b. Magnoliophyta: Liliidae: Liliales and Orchidales. Flora of North America north of Mexico, Volume 2. New York: Oxford University Press. 723 pp.
—— . 2003. Magnoliophyta: Caryophyllidae, part 1. Flora of North America north of Mexico, Volume 2. New York: Oxford University Press. 559 pp.
—— . 2005. Magnoliophyta: Caryophyllidae, part 2. Flora of North America north of Mexico, Volume 2. New York: Oxford University Press. 656 pp.
—— . 2006a. Magnoliophyta: Asteridae, part 6: Asteraceae, part 1. Flora of North America north of Mexico, Volume 2. New York: Ox-ford University Press. 579 pp.
—— . 2006b. Magnoliophyta: Asteridae, part 7: Asteraceae, part 2. Flora of North America north of Mexico, Volume 2. New York: Ox-ford University Press. 666 pp.
—— . 2006c. Magnoliophyta: Asteridae, part 8: Asteraceae, part 3. Flora of North America north of Mexico, Volume 2. New York: Ox-ford University Press. 616 pp.
Heaton, L. 1935. Plants from Pipe Spring Na-tional Monument.
Holmgren, N. H., P. K. Holmgren, and A. Cron-quist. 2005. Volume 2, Part B Subclass Dille-niidae. Intermountain fl ora: Vascular plants of the Intermountain West, U.S.A. Bronx, N.Y.: New York Botanical Garden. 488 pp.
Kartesz, J. T. 2003. Synthesis of the North Ameri-can fl ora, Version 2.0. http://www.bonap.org/synth.html.
Kearney, T. H., and R. H. Peebles. 1960. Arizona fl ora. Second edition. Berkeley: University of California Press. 1085 pp. (with supplement by J. T. Howell, E. McClintock, and collabo-rators).
King, R. J. 1977. The fl ora of Pipe Spring Na-tional Monument. Report prepared for the National Park Service by the Soil Conserva-tion Service, Fredonia, Arizona.
Kurth, L. 1998. Pipe Spring National Monu-ment non-native plants survey report, 1997. Report prepared for Pipe Spring National Monument by Zion National Park. 3 pp.
Leach, N. 1999. Pipe Spring National Monu-ment: An ancient oasis on a storied frontier. Springdale, Ut.: Zion Natural History As-sociation.
Margules, C. R., and R. L. Pressey. 2000. Sys-tematic conservation planning. Nature 405:243–253.
McDougall, W. B. 1973. Seed plants of northern Arizona. Flagstaff : Museum of Northern Arizona. 594 pp.
McKoy, K. L. 2000. Cultures at a crossroads: An administrative history of Pipe Spring National Monument. Cultural Resources Selections No. 15, Intermountain Region. Denver, Co.: National Park Service.
National Park Service (NPS). 2000. Management Policies. Washington, D.C.: U.S. Department of Interior, National Park Service. Publica-tion # NPS D1416. 137 pp.
Northern Colorado Plateau Network (NCPN). in ed. Pipe Spring National Monument veg-etation classifi cation and mapping project.
Noss, R. F., and A. Y. Cooperrider. 1994. Saving nature’s legacy: Protecting and restoring biodiversity. Washington, D.C.: Island Press. 416 pp.
Parfi tt, B. D. 1997. Opuntia pinkavae (Cacta-ceae), a new species from Arizona and Utah. Rhodora 99:223–228.
Weber, W. A., and R. C. Wittmann. 2001. Colo-rado fl ora: Western Slope. Third edition. Boulder: University Press of Colorado. 488 pp.
Welsh, S. L., N. D. Atwood, S. Goodrich, and L. C. Higgins. 2003. A Utah fl ora. Third edition, revised. Provo, Ut.: Brigham Young Univer-sity. 912 pp.
Appendices 17
6 AppendicesThe following appendices contain annotated checklists of the vascular fl ora of Pipe Spring Na-tional Monument.
Appendix A lists all species that have been con-fi rmed for the fl ora with an authenticated vouch-er specimen (including extant and historical re-cords), or that have been reliably reported for the monument in the literature (Heaton 1935, King 1977, Alexander 1998b). Vouchers were exam-ined from the PISP herbarium and collections from Brigham Young University (BRY), the Uni-versity of Wyoming (RM), and the digital muse-um databases of the New York Botanical Garden (NY), Northern Arizona University, Arizona State University, University of Arizona, and the Desert Botanical Garden. Species entries in Appendix A are organized alphabetically by family and genus.
Species listed in Appendix B are known from the vicinity of Pipe Spring National Monument, but have not previously been reported or confi rmed for the monument; that is, these species may po-tentially occur within PISP, based on the prox-imity of the monument to other populations or the presence of suitable habitat. Species on this list are derived from a checklist of potential spe-cies for NCPN parks prepared by Dr. N. Duane Atwood, of Brigham Young University, in 2001, and Jason Alexander for the PISP NPSpecies da-tabase. Some species from Appendix C could be considered as potentially occurring in the monu-ment, but are not included here.
Appendix C lists species that have been previ-ously reported for Pipe Spring National Monu-ment from published and unpublished literature (Alexander 1998a, Heaton 1935, King 1977) or from herbarium specimens, but are now consid-ered erroneous because they were based on mis-identifi ed specimens (False Reports or FalsRep), questionable literature records, or because the monument is well outside the known or expected range of the species (Questionable or Ques?).
Appendix D contains the same confi rmed, his-torical, and reported taxa as in Appendix A, but organized by life form rather than by family in or-der to better meet the needs of NPS interpretive specialists.
The codes and references for each fi eld in the ap-propriate appendices are summarized below.
Family (Appendices A, B, C, and D)Family concepts and nomenclature follow A Utah Flora (Welsh et al. 2003). Alternative family names and treatments from other regional fl oras (Dorn 2001, Flora of North America Editorial Commit-tee 1993, 2002b, Holmgren et al. 2005, Kearney and Peebles 1960, McDougall 1973,Weber and Wittman 2001) are listed in parentheses.
Species name (Appendices A, B, C, and D)Nomenclature for scientifi c names and taxonom-ic concepts are derived from Welsh et al. (2003). Scientifi c names are listed in alphabetical rather than phylogenetic order.
Synonyms/Taxonomic notes (Appendices A, B, and C)Pertinent synonyms are included for species with alternative scientifi c names in other recent state and regional fl oras (Barkworth et al. 2003, 2007, Barneby 1989, Cronquist 1994, Cronquist et al. 1972, 1977, 1984, 1997, Dorn 2001, Flora of North America Editorial Committee 1993, 1997, 2000, 2002a, 2002b, 2003, 2005, 2006a, 2006b, 2006c, Holmgren et al. 2005, Kearney and Peebles 1960, McDougall 1973, and Weber and Wittmann 2001). No eff ort has been made to identify ad-ditional synonyms from monographs and fl oras preceding the early 1970s.
Common name (Appendices A, B, C, and D)Common names are derived primarily from Welsh et al. (2003) and the Intermountain Flora (Barneby 1989, Cronquist 1994, Cronquist et al. 1972, 1977, 1984, 1997, Holmgren et al. 2005).
Life form (Appendices A, B, C, and D)Life form is based on the stature and growth habit of typical members of a species at reproductive maturity. Annual Forbs (AnnF) are broad-leaved dicots or monocots with non-woody stems that complete their life cycle (mature, fl ower, and die) in one year. Annual Graminoids (AnnG) are lin-ear-leaved, grass-like monocots that complete their life cycle (mature, fl ower, and die) in one year. Ferns and fern-allies (Fern) are non-fl ow-ering vascular plants that reproduce by spores. Perennial Forbs (PerF) are broad-leaved dicots or monocots with non-woody stems (at least above
18 Annotated Checklist of Vascular Flora: Pipe Spring National Monument
ground level) that live for multiple years. Perenni-al Graminoids (PerG) are linear-leaved, grass-like monocots that live for multiple years. Shrubs are woody perennials with one to many trunks aris-ing from near the base of the plant and are usually less than 3.5 m tall. Trees are woody perennials with a single stem or trunk and are typically over 3.5 m tall.
Range (Appendices A, B, and C)Range represents the distribution of a species within Arizona and the state’s contribution to the plant’s overall global range. Disjunct (Disj) taxa have their Arizona distribution separated from the main, contiguous portion of their range by a gap of more than 800 km (ca. 500 miles). Intro-duced (Intro) taxa are those that are not native to Arizona or North America but have become naturalized (breeding on their own without hu-man assistance). Local Endemics (LocEn) are taxa in which the entire global range is restricted to an area of less than 16,500 km2 (ca. 6,370 mi2, or 1° of latitude × 2° of longitude). Peripheral (Periph) taxa have a widespread global distribu-tion but occur at the margin of their contiguous range in Arizona and occupy less than 5% of the state’s area (usually only within a few miles of the state border). Regional Endemics (RegEn) are taxa with a global range of 16,500–250,000 km2 (an area about the size of the state of Wyoming). Sparse taxa occur widely across Arizona or North America but their range within Arizona is small and patchy, with populations restricted to special-ized or uncommon habitats. Widespread (Wide) taxa have global ranges exceeding 250,000 km2 and occur over at least 10% of the state.
Park status (Appendices A, C, and D)Park status identifi es whether a species is Pres-ent (Pres), Historical (Hist), or Reported (Rep) for Pipe Spring National Monument. Species are considered Present if a confi rmed voucher or documented observation has been made within monument boundaries since 1970. Historical species have been confi rmed with a voucher from the monument but have not been relocated since before 1970. Reported species are listed for the monument in the literature (Heaton 1935, King 1977, Alexander 1998b), but have not been cor-roborated with a voucher specimen. Species that may potentially occur in PISP are listed in Appen-dix B. Rejected taxa (listed in Appendix C) have either been falsely reported (FalsRep) in the lit-erature based on misidentifi ed herbarium speci-
mens or are considered Questionable (Ques?) because park reports are from well outside their known or expected range.
Pop. size (Appendix A)Population size refers to the abundance and fre-quency with which a species occurs within Pipe Spring National Monument. Abundance data are derived from the PISP NPSpecies database and personal observations. Abundant (Abun) taxa have large populations, have a broad ecological amplitude (often being dominant in one to sev-eral vegetation types), or occur extensively across the monument. Common (Com) taxa have large local populations but may be restricted to a single vegetation type or do not occur across the entire monument. Uncommon (Unc) taxa have low to medium-sized populations, typically have a nar-row ecological amplitude (or at least contribute relatively little to total vegetative cover), or have a small range across the monument. Rare taxa usually have low population numbers and are re-stricted to a single location within the monument. A “?” indicates that population size is unknown.
Source (Appendix A)Sources are the evidence for the assigned park status. In the case of Present and Historical taxa, sources are herbarium records (with collector, collection number, and repository) that were cor-roborated during this study. For reported taxa, the source is a published or unpublished refer-ence citing the species within the monument. Specimen repositories include the Pipe Spring National Monument herbarium (PISP), Brigham Young University (BRY), Utah State University (USU), New York Botanical Garden (NY), the University of Wyoming’s Rocky Mountain Her-barium (RM), Northern Arizona University, Ari-zona State University, University of Arizona, and Desert Botanical Garden.
Year doc. (Appendix A)Year documented indicates the fi rst year in which a species was confi rmed or reported to occur in Pipe Spring National Monument. This year does not necessarily correspond with the date of the voucher specimen cited under Source. “NA” indicates that the fi rst year is not known (used mostly for “reported” taxa).
Appendices 19
Comments (Appendices A, B, C, and D)This column includes additional comments on the status of a particular species in the monu-ment, the place of origin for introduced species, legal status, and other data.
Nativity (Appendix D)Nativity indicates whether a species is native to Arizona or introduced (Intro) based on McDou-gall (1973) and Kartesz (2003).
Flower time (Appendix D)Flower Time refers to the months in which a spe-cies is most likely to be in fl ower. NA indicates a species that is a non-fl owering fern or gymno-sperm.
Habitat (Appendix D)The distribution of species across generalized habitats in Pipe Spring National Monument is in-dicated by an “x” in the columns DesSh through Wet and is derived from the vegetation classifi ca-tion used by Alexander (1998b). DesSh = Desert shrub vegetation of fl ats and foothills below the monument’s Navajo Sandstone mesa dominated by greenish rabbitbrush (Chrysothamnus nauseo-sus var. consimilis), fourwing saltbush (Atriplex canescens), and sand sagebrush (Artemisia fi lifo-lia). PinJun = Pinyon-Juniper woodlands/rim-rock dominated by Utah juniper (Juniperus os-teosperma), Two-needle pinyon (Pinus edulis), turbinella live oak (Quercus turbinella), and Utah serviceberry (Amelanchier utahensis). Wet = Wet meadows associated with natural seeps and springs. Additional comments are included for species that are restricted to a particular soil/geo-logic type or are found mostly in disturbed sites.
20 Annotated Checklist of Vascular Flora: Pipe Spring National Monument
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3C
Am
aran
thac
eae
Am
aran
thus
blit
oide
sA
. gra
eciz
ans
pros
trat
e pi
gwee
dA
nnF
Wid
ePr
esU
ncA
lexa
nder
12
32 (P
ISP)
2001
Kin
g (1
977)
cit
ed
“Am
aran
thus
sp
.”
Am
aran
thac
eae
Am
aran
thu
s hy
poch
ondr
iacu
sgr
ain
amar
anth
Ann
FIn
tro
Pres
Unc
Fert
ig 2
3683
(P
ISP)
2007
Nat
ive
to S
W
USA
Ana
card
iace
aeRh
us a
rom
atic
a va
r. si
mpl
icif
olia
R. t
rilo
bata
var
. si
mpl
icif
olia
squa
wbu
shSh
rub
Wid
ePr
esU
ncPh
illip
s 95
-101
(N
AU
)19
95
Ana
card
iace
aeRh
us a
rom
atic
a va
r. tr
iloba
taR.
tri
loba
ta v
ar.
trilo
bata
squa
wbu
shSh
rub
Wid
eRe
p?
Hea
ton
(193
5)19
35
Apo
cyna
ceae
Am
soni
a jo
nesi
iJo
nes’
am
soni
aPe
rFRe
gEn
His
t?
Fann
ing
s.n.
(D
eser
t Bo
t G
arde
n)
1961
Apo
cyna
ceae
Am
soni
a to
men
tosa
va
r. st
enop
hylla
A. e
astw
oodi
ana
tom
ento
se
amso
nia
PerF
Spar
seH
ist
?H
eato
n s.
n.
(PIS
P)19
36PI
SP s
peci
men
is
the
“bre
vifo
lia”
form
Apo
cyna
ceae
Ap
ocy
nu
m
cann
abin
umIn
clud
es v
ars.
ca
nnab
inum
, gl
aber
rim
um, &
an
gust
ifol
ium
com
mon
do
gban
ePe
rFW
ide
Rep
?A
lexa
nder
(1
998b
)19
98
Asc
lepi
adac
eae
Asc
lepi
as la
tifo
liabr
oadl
eaf
milk
wee
dPe
rFW
ide
His
t?
Hea
ton
s.n.
(P
ISP)
1936
Asc
lepi
adac
eae
Asc
lep
ias
subv
erti
cilla
taA
. gal
ioid
esw
horl
ed
milk
wee
dPe
rFW
ide
Pres
Unc
Ale
xand
er
1225
(PIS
P)19
36
Appendices 21
Fam
ilySp
ecie
s na
me
Syno
nym
s/Ta
xono
mic
not
esCo
mm
onna
me
Life
fo
rmRa
nge
Park
st
atus
Pop.
si
zeSo
urce
Year
do
c.Co
mm
ents
Bora
gina
ceae
Cry
pta
nth
a ci
rcum
scis
sacu
shio
n cr
ypta
nth
Ann
FW
ide
His
t?
Hea
ton
s.n.
(P
ISP)
1936
Bora
gina
ceae
Cry
pta
nth
a co
nfer
tifl
ora
basi
n ye
llow
cr
ypta
nth
PerF
Wid
ePr
esU
ncN
elso
n 93
11
(RM
)19
37
Bora
gina
ceae
Cry
pta
nth
a cr
assi
sepa
la v
ar.
elac
hant
ha
thic
k-se
pale
d cr
ypta
nth
Ann
FW
ide
His
t?
Hea
ton
s.n.
(P
ISP)
1936
Bora
gina
ceae
Cryp
tant
ha fl
ava
Ore
ocar
ya fl
ava
plat
eau
yello
w
cryp
tant
hPe
rFW
ide
Rep
?A
lexa
nder
(1
998b
)N
A
Bora
gina
ceae
Cry
pta
nth
a fu
lvoc
anes
cens
Incl
udes
var
s.
fulv
ocan
esce
ns
& e
chin
oide
s,
Ore
ocar
ya
fulv
ocan
esce
ns
plat
eau
cryp
tant
hPe
rFRe
gEn
His
t?
Hea
ton
24
(PIS
P)19
37Ty
pe lo
calit
y of
va
r. ec
hino
ides
is
Par
ia C
anyo
n,
Gar
fi el
d Co
., U
T (J
ones
529
7p
POM
)
Bora
gina
ceae
Cryp
tant
ha g
raci
lissl
ende
r cr
ypta
nth
Ann
FW
ide
His
t?
Hea
ton
14
(PIS
P)19
37
Bora
gina
ceae
Cryp
tant
ha m
icra
ntha
redr
oot
cryp
tant
hA
nnF
Wid
eH
ist
?H
eato
n s.
n.
(PIS
P)19
36Ty
pe lo
calit
y is
“S
outh
ern
Uta
h”
(Par
ry 1
64 )
Bora
gina
ceae
Cryp
tant
ha p
tero
cary
a va
r. cy
clop
tera
win
gnut
cr
ypta
nth
Ann
FW
ide
His
t?
Hea
ton
13
(PIS
P)19
37
Bora
gina
ceae
Cryp
tant
ha p
tero
cary
a va
r. pt
eroc
arya
win
gnut
cr
ypta
nth
Ann
FW
ide
Pres
Unc
Even
den
&
Loui
e PI
SP 1
6 (P
ISP)
2001
Bora
gina
ceae
Hel
iotr
opiu
m
conv
olvu
lace
umEu
plo
ca
conv
olvu
lace
a ss
p.
calif
orni
ca
bind
wee
d he
liotr
ope
Ann
FW
ide
His
t?
Hea
ton
s.n.
(P
ISP)
1936
Bora
gina
ceae
Lapp
ula
occi
dent
alis
va
r. cu
pula
taL.
tex
ana,
L.
mar
gina
ta, L
. re
dow
skii
var.
cupu
lata
cups
eed
stic
ksee
dA
nnF
Wid
ePr
esU
ncEv
ende
n &
Lo
uie
PISP
14
(PIS
P)
1941
Bora
gina
ceae
Plag
iobo
thry
s jo
nesi
iJo
nes’
pop
corn
-fl
ower
Ann
FRe
gEn
Pres
Unc
Even
den
&
Loui
e PI
SP 1
5 (P
ISP)
2001
PISP
spe
cim
en
has
unus
ually
la
rge
fl ow
ers
App
endi
x A
. Con
fi rm
ed, h
isto
rica
l, an
d re
port
ed v
ascu
lar
plan
t ta
xa o
f Pi
pe S
prin
g N
atio
nal M
onum
ent,
org
aniz
ed b
y fa
mily
and
sci
enti
fi c
nam
e w
ith
taxo
nom
ic
and
biog
eogr
aphi
c no
tes,
con
t.
22 Annotated Checklist of Vascular Flora: Pipe Spring National Monument
Fam
ilySp
ecie
s na
me
Syno
nym
s/Ta
xono
mic
not
esCo
mm
onna
me
Life
fo
rmRa
nge
Park
st
atus
Pop.
si
zeSo
urce
Year
do
c.Co
mm
ents
Cact
acea
eEc
hino
cere
us
enge
lman
nii v
ar.
vari
egat
us
E. e
ngel
man
nii v
ar.
decu
mbe
ns. V
ars.
not
re
cogn
ized
in F
lora
of
Nor
th A
mer
ica
(200
3)
Gle
n Ca
nyon
he
dgeh
og
cact
us
PerF
RegE
nRe
p?
Ale
xand
er
(199
8b)
1998
Cact
acea
eEc
hino
cere
us
trig
loch
idia
tus
var.
mel
anac
anth
us
Incl
udes
var
. ine
rmis
. V
ars.
not
rec
ogni
zed
in F
lora
of
Nor
th
Am
eric
a (2
003)
clar
etcu
pPe
rFW
ide
Pres
Unc
Hea
ton
s.n.
(P
ISP)
1936
Cact
acea
eEc
hino
cere
us
trig
loch
idia
tus
var.
moj
aven
sis
Var
s. n
ot r
ecog
nize
d in
Flo
ra o
f N
orth
A
mer
ica
(200
3)
Moh
ave
clar
etcu
pPe
rFW
ide
His
t?
Peeb
les
1469
6 (A
RIZ)
1940
Cact
acea
eO
punt
ia e
rina
cea
var.
aure
aO
. aur
ea, O
. bas
ilari
s va
r. au
rea
Pipe
Spr
ing
pric
klyp
ear
PerF
LocE
nPr
esCo
mM
acLe
od 4
22
(ASU
)19
30Ty
pe lo
calit
y is
1/
2 m
ile N
of
Pipe
Spr
ing,
M
ohav
e Co
., A
Z (M
cCab
e &
M
cCab
e s.
n. D
S)
Cact
acea
eO
pu
nti
a er
inac
ea v
ar.
utah
ensi
sV
ar. n
ot r
ecog
nize
d in
Flo
ra o
f N
orth
A
mer
ica
(200
3);
cons
ider
ed a
few
-sp
ined
var
iant
of
O.
poly
acan
tha
Uta
h pr
ickl
ypea
rPe
rFW
ide
Pres
Com
Mac
Leod
428
(A
SU)
1971
Cact
acea
eO
punt
ia f
ragi
lisO
. fra
gilis
var
. br
achy
arth
abr
ittl
e pr
ickl
ypea
rPe
rFW
ide
Rep
?K
ing
(197
7),
Ale
xand
er
(199
8b)
1977
Cact
acea
eO
punt
ia p
haea
cant
ha
var.
maj
orV
ars.
not
rec
ogni
zed
in F
lora
of
Nor
th
Am
eric
a (2
003)
larg
e pr
ickl
ypea
rPe
rFW
ide
Pres
Unc
Gan
z 17
7 (A
SU)
1965
Als
o ci
ted
by
Ale
xand
er
(199
8b)
Cact
acea
eO
punt
ia p
inka
vae
O. b
asila
ris
var.
woo
dbur
yi,
cons
ider
ed p
art
of O
. m
acro
rhiz
a in
Uta
h
Flor
a (2
003)
Pink
ava’
s pr
ickl
ypea
rPe
rFLo
cEn
Pres
Unc
Leht
o s.
n.
(ASU
)19
40Ty
pe lo
calit
y is
N
W o
f Bu
lrus
h Ca
nyon
S o
f Pi
pe
Spri
ng, M
ohav
e Co
., A
Z (P
arfi
t 28
74 A
SU)
App
endi
x A
. Con
fi rm
ed, h
isto
rica
l, an
d re
port
ed v
ascu
lar
plan
t ta
xa o
f Pi
pe S
prin
g N
atio
nal M
onum
ent,
org
aniz
ed b
y fa
mily
and
sci
enti
fi c
nam
e w
ith
taxo
nom
ic
and
biog
eogr
aphi
c no
tes,
con
t.
Appendices 23
Fam
ilySp
ecie
s na
me
Syno
nym
s/Ta
xono
mic
not
esCo
mm
onna
me
Life
fo
rmRa
nge
Park
st
atus
Pop.
si
zeSo
urce
Year
do
c.Co
mm
ents
Cact
acea
eO
punt
ia w
hipp
lei v
ar.
whi
pple
iCy
lindr
opun
tia
whi
pple
iW
hipp
le’s
chol
laSh
rub
RegE
nPr
esU
ncK
ing
(197
7),
Ale
xand
er
(199
8b)
1977
Phot
ogra
phed
(b
ut n
ot
vouc
here
d) b
y W
. Fe
rtig
, 200
7
Cact
acea
eSc
lero
cact
us w
hipp
lei
var.
rose
usS.
par
vifl
oru
s,
S. w
hipp
lei v
ar.
inte
rmed
ius.
Incl
udes
S.
con
tort
us &
S.
terr
ae-c
anyo
nae
smal
lfl ow
er
fi sh
hook
cac
tus
PerF
Peri
phPr
esU
ncEv
ende
n &
Lo
uie
PISP
26
(PIS
P)
1998
Type
of
S.
inte
rmed
ius
is 9
m
i SW
of
Pipe
Sp
ring
, Moh
ave
Co.,
AZ
(Pee
bles
&
Par
ker
1471
2 CA
S)
Capp
arac
eae
(Cle
omac
eae)
Cleo
me
lute
aye
llow
bee
plan
tA
nnF
Wid
ePr
esU
ncA
lexa
nder
11
87 (P
ISP)
1935
Chen
opod
iace
aeA
trip
lex
arge
ntea
var
. ar
gent
eaA
. arg
ente
asi
lver
ora
chA
nnF
Wid
ePr
esU
ncA
lexa
nder
12
36 (P
ISP)
2001
Chen
opod
iace
aeA
trip
lex
cane
scen
s va
r. ca
nesc
ens
A. c
anes
cens
var
. oc
cide
ntal
isfo
urw
ing
salt
bush
Shru
bW
ide
Pres
Com
Ale
xand
er
1212
(PIS
P)19
66
Chen
opod
iace
aeCh
enop
odiu
m a
lbum
va
r. al
bum
C. a
lbum
lam
bsqu
arte
rA
nnF
Intr
oPr
esU
ncA
lexa
nder
12
61 (P
ISP)
1998
Nat
ive
to E
uras
ia
Chen
opod
iace
aeC
hen
op
od
ium
fr
emon
tii v
ar.
frem
onti
i
C. f
rem
onti
iFr
emon
t’s
goos
efoo
tA
nnF
Wid
ePr
esU
ncA
lexa
nder
11
92 (P
ISP)
2001
Chen
opod
iace
aeC
hen
op
od
ium
le
ptop
hyllu
mna
rrow
leaf
go
osef
oot
Ann
FW
ide
Rep
?H
eato
n (1
935)
1935
Chen
opod
iace
aeK
rasc
heni
nnik
ovia
la
nata
var
. sub
spin
osa
Euro
tia
lana
ta,
Cera
toid
es la
nata
var
. su
bspi
nosa
. Var
s. n
ot
reco
gniz
ed in
Flo
ra
of N
orth
Am
eric
a (2
003)
.
shru
bby
win
terf
atSh
rub
Wid
eH
ist
?Pe
eble
s 13
079
(ARI
Z)19
36Ty
pe lo
calit
y is
ne
ar S
t. G
eorg
e,
Was
hing
ton
Co.,
UT
(Goo
ddin
g 81
0 N
Y)
Chen
opod
iace
aeSa
lsol
a tr
agus
S. k
ali,
S. ib
eric
a, S
. pe
stif
er, S
. aus
tral
isRu
ssia
n th
istl
eA
nnF
Intr
oPr
esU
ncFe
rtig
236
76
(PIS
P)19
35N
ativ
e to
Asi
a
Chen
opod
iace
ae
(Sar
coba
tace
ae)
Sarc
ob
atu
s ve
rmic
ulat
usgr
ease
woo
dSh
rub
Wid
ePr
esU
ncA
lexa
nder
12
26 (P
ISP)
1935
Chen
opod
iace
aeSu
aeda
tor
reya
na v
ar.
torr
eyan
aS.
moq
uini
i, S.
nig
ra,
S. f
ruti
cosa
Torr
ey’s
seep
wee
dA
nnF
Wid
ePr
esU
ncA
lexa
nder
12
52 (P
ISP)
2001
App
endi
x A
. Con
fi rm
ed, h
isto
rica
l, an
d re
port
ed v
ascu
lar
plan
t ta
xa o
f Pi
pe S
prin
g N
atio
nal M
onum
ent,
org
aniz
ed b
y fa
mily
and
sci
enti
fi c
nam
e w
ith
taxo
nom
ic
and
biog
eogr
aphi
c no
tes,
con
t.
24 Annotated Checklist of Vascular Flora: Pipe Spring National Monument
Fam
ilySp
ecie
s na
me
Syno
nym
s/Ta
xono
mic
not
esCo
mm
onna
me
Life
fo
rmRa
nge
Park
st
atus
Pop.
si
zeSo
urce
Year
do
c.Co
mm
ents
Com
posi
tae
(Ast
erac
eae)
Am
bro
sia
acan
thic
arpa
Fran
seri
a ac
anth
icar
pabu
r ra
gwee
dA
nnF
Wid
ePr
esU
ncA
lexa
nder
11
85 (P
ISP)
1935
Com
posi
tae
(Ast
erac
eae)
Am
bro
sia
arte
mis
iifol
iaA
. art
emis
iifol
ia v
ar.
elat
ior
com
mon
ra
gwee
dA
nnF
Wid
ePr
esU
ncA
lexa
nder
11
86 (P
ISP)
1977
Com
posi
tae
(Ast
erac
eae)
Art
emis
ia b
igel
ovii
Bige
low
’s sa
gebr
ush
Shru
bW
ide
Pres
Unc
Ale
xand
er
(199
8b)
1977
Obs
erve
d in
fi el
d by
J. C
oles
and
W
. Fer
tig,
200
7
Com
posi
tae
(Ast
erac
eae)
Art
emis
ia c
ampe
stri
s va
r. sc
oule
rian
aA
. cam
pest
ris
ssp.
pac
ifi ca
, A.
paci
fi ca
, Olig
ospo
rus
cam
pest
ris
ssp.
pa
cifi
cus,
O. p
acifi
cus
sand
w
orm
woo
dPe
rFW
ide
Rep
?A
lexa
nder
(1
998b
)19
98O
bser
ved
in fi
eld
by J
. Col
es, 2
007
Com
posi
tae
(Ast
erac
eae)
Art
emis
ia d
racu
ncul
us
ssp.
gla
uca
Olig
osp
oru
s d
racu
ncu
lus
ssp.
gla
ucus
, A.
drac
uncu
loid
es. V
ar.
not
reco
gniz
ed in
Cr
onqu
ist
(199
4)
and
Flor
a of
Nor
th
Am
eric
a (2
006)
tarr
agon
PerF
Wid
eRe
p?
Kin
g (1
977)
1977
Com
posi
tae
(Ast
erac
eae)
Art
emis
ia fi
lifol
iaO
ligos
poru
s fi
lifol
ius
sand
sag
ebru
sh
Shru
bW
ide
Pres
Unc
Ale
xand
er
1257
(PIS
P)19
77
Com
posi
tae
(Ast
erac
eae)
Art
emis
ia lu
dovi
cian
a va
r. al
bula
A. l
udov
icia
na s
sp.
albu
laLo
uisi
ana
wor
mw
ood
PerF
Wid
ePr
esU
ncFe
rtig
236
81
(PIS
P)19
77
Com
posi
tae
(Ast
erac
eae)
Art
emis
ia t
ride
ntat
a va
r. tr
iden
tata
Seri
ph
idiu
m
trid
enta
tum
ssp
. tr
iden
tatu
m
basi
n bi
g sa
gebr
ush
Shru
bW
ide
Pres
Unc
Fert
ig 2
3674
(P
ISP)
2007
Com
posi
tae
(Ast
erac
eae)
Ast
er h
espe
rius
A. c
oeru
lesc
ens,
A
. lan
ceol
atus
va
r. h
esp
eriu
s,
Sym
ph
yotr
ich
um
la
nceo
latu
m v
ar.
hesp
eriu
m
Sisk
iyou
ast
erPe
rFW
ide
Pres
Rare
Ale
xand
er
1240
(PIS
P)20
01
Com
posi
tae
(Ast
erac
eae)
Ch
aen
acti
s ca
rpho
clin
iaC.
car
phoc
linia
var
. ca
rpho
clin
iaG
ray’
s ch
aena
ctis
Ann
FW
ide
Pres
Unc
Loui
e &
Ev
ende
n PI
SP-
10 (P
ISP)
1937
App
endi
x A
. Con
fi rm
ed, h
isto
rica
l, an
d re
port
ed v
ascu
lar
plan
t ta
xa o
f Pi
pe S
prin
g N
atio
nal M
onum
ent,
org
aniz
ed b
y fa
mily
and
sci
enti
fi c
nam
e w
ith
taxo
nom
ic
and
biog
eogr
aphi
c no
tes,
con
t.
Appendices 25
Fam
ilySp
ecie
s na
me
Syno
nym
s/Ta
xono
mic
not
esCo
mm
onna
me
Life
fo
rmRa
nge
Park
st
atus
Pop.
si
zeSo
urce
Year
do
c.Co
mm
ents
Com
posi
tae
(Ast
erac
eae)
Ch
aeto
pap
pa
eric
oide
sLe
ucel
ene
eric
oide
s,
Ast
er a
reno
sus
rose
-hea
thPe
rFW
ide
Pres
Unc
Ale
xand
er
1207
(PIS
P)19
39
Com
posi
tae
(Ast
erac
eae)
Chry
soth
amnu
s na
useo
sus
var.
cons
imili
s
Eric
amer
ia n
ause
osa
var.
cons
imili
s,
E. n
ause
osa
var.
oreo
phila
, C.
naus
eosu
s va
r. or
eoph
ilus
gree
nish
ra
bbit
brus
hSh
rub
Wid
ePr
esCo
mA
lexa
nder
12
19 (P
ISP)
1998
Com
posi
tae
(Ast
erac
eae)
Chry
soth
amnu
s na
useo
sus
var.
gnap
halo
des
C. n
ause
osus
va
r. ho
lole
ucus
, Er
icam
eria
nau
seos
a va
r. ho
lole
uca
gray
stem
ra
bbit
brus
hSh
rub
Wid
ePr
esU
ncFe
rtig
236
65
(PIS
P)20
07
Com
posi
tae
(Ast
erac
eae)
Chry
soth
amnu
s na
useo
sus
var.
grav
eole
ns
Eric
amer
ia n
ause
osa
var.
grav
eole
ns,
E. n
ause
osa
var.
glab
rata
glab
rate
ra
bbit
brus
hSh
rub
Wid
eRe
p?
Ale
xand
er
(199
8b)
1998
Com
posi
tae
(Ast
erac
eae)
Chry
soth
amnu
s na
useo
sus
var.
junc
eus
Eric
amer
ia n
ause
osa
var.
junc
earu
sh
rabb
itbr
ush
Shru
bRe
gEn
Pres
Unc
Fert
ig 2
3678
(P
ISP)
2007
Com
posi
tae
(Ast
erac
eae)
Chry
soth
amnu
s vi
scid
ifl or
us v
ar.
pube
rulu
s
Chry
soth
amnu
s vi
scid
ifl or
us s
sp.
pube
rulu
s
pube
rule
nt
rabb
itbr
ush
Shru
bW
ide
Pres
Unc
Ale
xand
er
1193
(PIS
P)20
01
Com
posi
tae
(Ast
erac
eae)
Chry
soth
amnu
s vi
scid
ifl or
us v
ar.
sten
ophy
llus
Incl
uded
in v
ar.
visc
idifl
orus
by
som
e au
thor
s. V
ar. n
ot
reco
gniz
ed in
Flo
ra
of N
orth
Am
eric
a (2
006)
.
slen
derl
eaf
rabb
itbr
ush
Shru
bW
ide
Pres
Com
Ale
xand
er
1247
(PIS
P)19
98
Com
posi
tae
(Ast
erac
eae)
Chry
soth
amnu
s vi
scid
ifl or
us v
ar.
visc
idifl
orus
gree
n ra
bbit
brus
hSh
rub
Wid
eRe
p?
Kin
g (1
977)
1977
Com
posi
tae
(Ast
erac
eae)
Cony
za c
anad
ensi
s va
r. gl
abra
taEr
iger
on c
anad
ensi
s,
Var
s. n
ot r
ecog
nize
d in
Flo
ra o
f N
orth
A
mer
ica
(200
6)
Cana
dian
ho
rsew
eed
Ann
FW
ide
Rep
?H
eato
n (1
935)
1935
Com
posi
tae
(Ast
erac
eae)
Erig
eron
div
erge
ns
var.
dive
rgen
sE.
div
erge
nssp
read
ing
dais
yPe
rFW
ide
Pres
Unc
Hea
ton
s.n.
(P
ISP)
1936
App
endi
x A
. Con
fi rm
ed, h
isto
rica
l, an
d re
port
ed v
ascu
lar
plan
t ta
xa o
f Pi
pe S
prin
g N
atio
nal M
onum
ent,
org
aniz
ed b
y fa
mily
and
sci
enti
fi c
nam
e w
ith
taxo
nom
ic
and
biog
eogr
aphi
c no
tes,
con
t.
26 Annotated Checklist of Vascular Flora: Pipe Spring National Monument
Fam
ilySp
ecie
s na
me
Syno
nym
s/Ta
xono
mic
not
esCo
mm
onna
me
Life
fo
rmRa
nge
Park
st
atus
Pop.
si
zeSo
urce
Year
do
c.Co
mm
ents
Com
posi
tae
(Ast
erac
eae)
Erig
eron
pum
ilus
var.
conc
innu
sE.
con
cinn
us v
ar.
conc
innu
s. In
clud
es
var.
subg
labe
r
Nav
ajo
fl ea
bane
PerF
Peri
phRe
p?
Ale
xand
er
(199
8b)
1998
Com
posi
tae
(Ast
erac
eae)
Gai
llard
ia a
rizo
nica
Ari
zona
bl
anke
tfl o
wer
Ann
FRe
gEn
Rep
?A
lexa
nder
(1
998b
)19
98
Com
posi
tae
(Ast
erac
eae)
Gai
llard
ia p
inna
tifi
daH
opi
blan
ketfl
ow
erPe
rFW
ide
Pres
Unc
Ale
xand
er
1208
(PIS
P)20
01
Com
posi
tae
(Ast
erac
eae)
Gri
ndel
ia s
quar
rosa
va
r. se
rrul
ata
Var
s. n
ot r
ecog
nize
d in
Flo
ra o
f N
orth
A
mer
ica
(200
6)
curl
ycup
gu
mw
eed
PerF
Wid
eRe
p?
Ale
xand
er
(199
8b)
1998
Com
posi
tae
(Ast
erac
eae)
Gu
tier
rezi
a m
icro
ceph
ala
Incl
udes
G. l
ucid
ath
read
sn
akew
eed
Shru
bW
ide
Pres
Unc
Ale
xand
er
1198
(PIS
P)19
35
Com
posi
tae
(Ast
erac
eae)
Gut
ierr
ezia
sar
othr
aebr
oom
sn
akew
eed
Shru
bW
ide
Pres
Com
Ale
xand
er
1258
(PIS
P)20
01
Com
posi
tae
(Ast
erac
eae)
Hel
iant
hus
annu
us
ssp.
lent
icul
aris
H. a
ridu
s; s
sp. n
ot
reco
gniz
ed in
Flo
ra
of N
orth
Am
eric
a (2
006)
com
mon
su
nfl o
wer
Ann
FW
ide
Pres
Unc
Ale
xand
er
1197
(PIS
P)19
35
Com
posi
tae
(Ast
erac
eae)
Hym
enop
appu
s fi
lifol
ius
var.
cine
reus
H. l
ugen
s, H
. fi
lifol
ius
var.
luge
ns, H
. fi li
foliu
s va
r. pa
ucifl
oru
s,
H. fi
lifo
lius
var.
meg
acep
halu
s,
H. fi
lifo
lius
var.
tom
ento
sus
com
mon
hy
alin
eher
bPe
rFW
ide
Pres
Unc
Ale
xand
er
1202
(PIS
P)19
98
Com
posi
tae
(Ast
erac
eae)
Iva
axill
aris
I.
axill
aris
var
. ax
illar
is. V
ars.
not
re
cogn
ized
in F
lora
of
Nor
th A
mer
ica
(200
6).
pove
rtyw
eed
PerF
Wid
ePr
esU
ncA
lexa
nder
12
17 (P
ISP)
1998
Com
posi
tae
(Ast
erac
eae)
Lact
uca
serr
iola
pric
kly
lett
uce
Ann
FIn
tro
Pres
Unc
Fert
ig 2
3690
(P
ISP)
1935
Nat
ive
to E
urop
e
Com
posi
tae
(Ast
erac
eae)
Layi
a gl
andu
losa
tidy
tips
Ann
FW
ide
Pres
Unc
Loui
e &
Ev
ende
n PI
SP 8
(P
ISP)
2001
App
endi
x A
. Con
fi rm
ed, h
isto
rica
l, an
d re
port
ed v
ascu
lar
plan
t ta
xa o
f Pi
pe S
prin
g N
atio
nal M
onum
ent,
org
aniz
ed b
y fa
mily
and
sci
enti
fi c
nam
e w
ith
taxo
nom
ic
and
biog
eogr
aphi
c no
tes,
con
t.
Appendices 27
Fam
ilySp
ecie
s na
me
Syno
nym
s/Ta
xono
mic
not
esCo
mm
onna
me
Life
fo
rmRa
nge
Park
st
atus
Pop.
si
zeSo
urce
Year
do
c.Co
mm
ents
Com
posi
tae
(Ast
erac
eae)
Mac
haer
anth
era
cane
scen
s va
r. ca
nesc
ens
Ast
er c
anes
cens
, D
iete
ria
cane
scen
s va
r. ca
nesc
ens
hoar
y as
ter
PerF
Wid
ePr
esU
ncEv
ende
n &
Lo
uie
PISP
31
(PIS
P)
1936
Com
posi
tae
(Ast
erac
eae)
Mac
haer
anth
era
tana
ceti
folia
Ast
er t
anac
etif
oliu
sta
nsyl
eaf
aste
rA
nnF
Wid
ePr
esU
ncA
lexa
nder
12
15 (P
ISP)
1998
Com
posi
tae
(Ast
erac
eae)
Mal
acot
hrix
gla
brat
aM
. cal
ifor
nica
var
. gl
abra
tafi
lifor
m d
eser
t-da
ndel
ion
Ann
FW
ide
Pres
Unc
Even
den
&
Loui
e PI
SP 2
7 (P
ISP)
2001
Com
posi
tae
(Ast
erac
eae)
Mal
aco
thri
x so
ncho
ides
sow
this
tle
dese
rt-
dand
elio
n
Ann
FW
ide
His
t?
Hea
ton
3 (P
ISP)
1937
Com
posi
tae
(Ast
erac
eae)
On
op
ord
um
ac
anth
ium
O. a
cant
hium
var
. ac
anth
ium
Scot
ch t
hist
lePe
rFIn
tro
Rep
?A
lexa
nder
(1
998b
)19
98A
Z st
ate
proh
ibit
ed
and
rest
rict
ed
noxi
ous
wee
d.
Nat
ive
to E
uras
ia
Com
posi
tae
(Ast
erac
eae)
Psilo
stro
phe
spar
sifl
ora
gree
nste
m
pape
rfl o
wer
PerF
RegE
nH
ist
?H
eato
n s.
n.
(PIS
P)19
36Ty
pe lo
calit
y is
“S
outh
ern
Uta
h”
(Tho
mps
on s
.n.
GH
)
Com
posi
tae
(Ast
erac
eae)
Sene
cio
doug
lasi
i var
. lo
ngilo
bus
S. lo
ngilo
bus,
S.
fl ac
cidu
s va
r. fl
acci
dus,
S. fl
acc
idu
s va
r. do
ugla
sii
Dou
glas
’ gr
ound
sel
PerF
Wid
ePr
esU
ncW
hite
head
s.n
. (P
ISP)
1935
Com
posi
tae
(Ast
erac
eae)
Solid
ago
sp.
gold
enro
dPe
rFW
ide
Rep
?A
lexa
nder
(1
998b
)19
98Id
enti
fi ca
tion
ne
eded
Com
posi
tae
(Ast
erac
eae)
Sonc
hus
aspe
rsp
inyl
eaf
sow
this
tle
Ann
FIn
tro
Pres
Unc
Even
den
&
Loui
e PI
SP 3
5 (P
ISP)
1997
Nat
ive
to E
urop
e
Com
posi
tae
(Ast
erac
eae)
Sonc
hus
oler
aceu
sco
mm
on
sow
this
tle
Ann
FIn
tro
Rep
?A
lexa
nder
(1
998b
)19
98N
ativ
e to
Eur
ope
Com
posi
tae
(Ast
erac
eae)
Step
hano
mer
ia e
xigu
aS.
exi
gua
ssp.
exi
gua
whi
te-p
lum
e w
ire
lett
uce
Ann
FW
ide
Pres
Unc
Ale
xand
er
1199
(PIS
P)20
01
Com
posi
tae
(Ast
erac
eae)
Step
hano
mer
ia
pauc
ifl or
abr
own-
plum
e w
ire
lett
uce
PerF
Wid
ePr
esU
ncFe
rtig
236
68
(PIS
P)19
98
App
endi
x A
. Con
fi rm
ed, h
isto
rica
l, an
d re
port
ed v
ascu
lar
plan
t ta
xa o
f Pi
pe S
prin
g N
atio
nal M
onum
ent,
org
aniz
ed b
y fa
mily
and
sci
enti
fi c
nam
e w
ith
taxo
nom
ic
and
biog
eogr
aphi
c no
tes,
con
t.
28 Annotated Checklist of Vascular Flora: Pipe Spring National Monument
Fam
ilySp
ecie
s na
me
Syno
nym
s/Ta
xono
mic
not
esCo
mm
onna
me
Life
fo
rmRa
nge
Park
st
atus
Pop.
si
zeSo
urce
Year
do
c.Co
mm
ents
Com
posi
tae
(Ast
erac
eae)
Thel
esp
erm
a su
bnud
um v
ar.
subn
udum
Var
s. n
ot r
ecog
nize
d in
Flo
ra o
f N
orth
A
mer
ica
(200
6)
scap
ose
gree
nthr
ead
PerF
Wid
ePr
esU
ncA
lexa
nder
12
04 (P
ISP)
1937
Type
loca
lity
is
“Sou
ther
n U
tah”
(P
arry
109
GH
)
Com
posi
tae
(Ast
erac
eae)
Tow
nsen
dia
inca
naho
ary
tow
nsen
dia
PerF
Wid
ePr
esU
ncLo
uie
&
Even
den
PISP
19
(PIS
P)
2001
Com
posi
tae
(Ast
erac
eae)
Trag
opog
on d
ubiu
sT.
dub
ius
var.
maj
orye
llow
sal
sify
PerF
Intr
oPr
esU
ncEv
ende
n &
Lo
uie
PISP
28
(PIS
P)
1997
Nat
ive
to E
urop
e
Com
posi
tae
(Ast
erac
eae)
Ver
besi
na e
ncel
ioid
es
var.
exau
ricu
lata
Xim
enes
ia
ence
lioid
esgo
lden
cr
ownb
eard
Ann
FW
ide
Pres
Unc
Ale
xand
er
1188
(PIS
P)19
35
Com
posi
tae
(Ast
erac
eae)
Xan
thiu
m s
trum
ariu
m
var.
cana
dens
eX
. pen
sylv
anic
um,
X. s
acch
arat
um.
Var
s. n
ot r
ecog
nize
d in
Flo
ra o
f N
orth
A
mer
ica
(200
6).
roug
h co
ckle
bur
Ann
FW
ide
Rep
?H
eato
n (1
935)
&
Kin
g (1
977)
1935
Conv
olvu
lace
aeCo
nvol
vulu
s ar
vens
isfi
eld
bind
wee
dPe
rFIn
tro
His
t?
Park
er 6
238
(ARI
Z)19
46A
Z st
ate
proh
ibit
ed
and
regu
late
d no
xiou
s w
eed.
N
ativ
e to
Eur
asia
Cruc
ifer
ae
(Bra
ssic
acea
e)A
rabi
s pe
renn
ans
var.
pere
nnan
sBo
eche
ra p
eren
nans
pere
nnia
l ro
ckcr
ess
PerF
Wid
eRe
p?
Ale
xand
er
(199
8b)
1998
Cruc
ifer
ae
(Bra
ssic
acea
e)Ca
psel
la b
ursa
-pas
tori
ssh
ephe
rd’s-
purs
eA
nnF
Intr
oH
ist
?H
eato
n 17
(P
ISP)
1937
Nat
ive
to E
urop
e
Cruc
ifer
ae
(Bra
ssic
acea
e)D
escu
rain
ia p
inna
ta
var.
glab
raw
este
rn
tans
ymus
tard
Ann
FW
ide
His
t?
Hea
ton
s.n.
(P
ISP)
1935
Cruc
ifer
ae
(Bra
ssic
acea
e)D
escu
rain
ia p
inna
ta
var.
inte
rmed
iaIn
clud
ed in
D.
pinn
ata
var.
osm
iaru
m b
y so
me
auth
ors
wes
tern
ta
nsym
usta
rdA
nnF
Peri
phH
ist
?H
eato
n 19
41-1
(P
ISP)
1939
Cruc
ifer
ae
(Bra
ssic
acea
e)D
escu
rain
ia p
inna
ta
var.
osm
iaru
mD
. pin
nata
ssp
. ha
licto
rum
wes
tern
ta
nsym
usta
rdA
nnF
Wid
ePr
esU
ncH
eato
n 19
41-2
(P
ISP)
1941
Cruc
ifer
ae
(Bra
ssic
acea
e)D
escu
rain
ia s
ophi
afl
ixw
eed
Ann
FIn
tro
Pres
Unc
Nel
son
9304
(R
M)
1971
Nat
ive
to E
urop
e
Cruc
ifer
ae
(Bra
ssic
acea
e)D
ithy
rea
wis
lizen
iD
imor
phoc
arpa
w
isliz
eni
spec
tacl
epod
Ann
FW
ide
Pres
Unc
Hea
ton
s.n.
(P
ISP)
1936
App
endi
x A
. Con
fi rm
ed, h
isto
rica
l, an
d re
port
ed v
ascu
lar
plan
t ta
xa o
f Pi
pe S
prin
g N
atio
nal M
onum
ent,
org
aniz
ed b
y fa
mily
and
sci
enti
fi c
nam
e w
ith
taxo
nom
ic
and
biog
eogr
aphi
c no
tes,
con
t.
Appendices 29
Fam
ilySp
ecie
s na
me
Syno
nym
s/Ta
xono
mic
not
esCo
mm
onna
me
Life
fo
rmRa
nge
Park
st
atus
Pop.
si
zeSo
urce
Year
do
c.Co
mm
ents
Cruc
ifer
ae
(Bra
ssic
acea
e)Er
ysim
um r
epan
dum
spre
adin
g w
allfl
ower
Ann
FIn
tro
His
t?
Hea
ton
s.n.
(P
ISP)
1939
Nat
ive
to E
urop
e
Cruc
ifer
ae
(Bra
ssic
acea
e)Le
pidi
um d
ensi
fl o
rum
va
r. ra
mos
umIn
clud
es v
ar.
mac
roca
rpum
prai
rie
pepp
erw
ort
Ann
FPe
riph
His
t?
Hea
ton
s.n.
(P
ISP)
1936
Cruc
ifer
ae
(Bra
ssic
acea
e)Le
pidi
um m
onta
num
va
r. jo
nesi
iJo
nes’
pe
pper
wor
tPe
rFW
ide
Pres
Com
Hea
ton
s.n.
(P
ISP)
1937
Type
loca
lity
is S
t. G
eorg
e,
Was
hing
ton
Co.,
UT
(Jon
es 1
636
NY
)
Cruc
ifer
ae
(Bra
ssic
acea
e)Le
pidi
um m
onta
num
va
r. st
ella
eL.
mon
tanu
m v
ar.
cine
reum
Stel
la’s
pepp
erw
ort
PerF
LocE
nH
ist
?H
eato
n 19
41-
17 (P
ISP)
1941
Type
loca
lity
is S
E of
Can
nonv
ille,
K
ane
Co.,
UT
(Rev
eal &
Rev
eal
4454
BRY
)
Cruc
ifer
ae
(Bra
ssic
acea
e)Le
pidi
um p
erfo
liatu
mcl
aspi
ng
pepp
erw
ort
Ann
FIn
tro
His
t?
Leht
o 71
72
(ARI
Z)19
66N
ativ
e to
Eur
ope
Cruc
ifer
ae
(Bra
ssic
acea
e)N
astu
rtiu
m o
ffi c
inal
eRo
ripp
a na
stur
tium
-aq
uati
cum
wat
ercr
ess
PerF
Intr
oPr
esU
ncFe
rtig
236
92
(PIS
P)19
97N
ativ
e to
Eur
ope
Cruc
ifer
ae
(Bra
ssic
acea
e)Si
sym
briu
m a
ltis
sim
umtu
mbl
e m
usta
rdA
nnF
Intr
oPr
esU
ncH
eato
n 19
41-
11 (P
ISP)
1941
Nat
ive
to E
urop
e
Cruc
ifer
ae
(Bra
ssic
acea
e)St
anle
ya p
inna
taS.
pin
nata
var
. pi
nnat
apr
ince
’s-pl
ume
PerF
Wid
ePr
esU
ncEv
ende
n &
Lo
uie
PISP
3
(PIS
P)
1939
Cruc
ifer
ae
(Bra
ssic
acea
e)St
rept
anth
ella
lo
ngir
ostr
islo
ng-b
eak
fi dd
le m
usta
rdA
nnF
Wid
eH
ist
?H
eato
n 18
(P
ISP)
1937
Cucu
rbit
acea
eCu
curb
ita
foet
idis
sim
abu
ffal
o go
urd
PerF
Wid
ePr
esU
ncA
lexa
nder
12
18 (P
ISP)
1935
Cupr
essa
ceae
Jun
iper
us
oste
ospe
rma
Sabi
na o
steo
sper
ma
Uta
h ju
nipe
rSh
rub
Wid
ePr
esU
ncA
lexa
nder
12
46 (P
ISP)
1935
Cupr
essa
ceae
Juni
peru
s sc
opul
orum
Sabi
na s
copu
loru
mRo
cky
Mou
ntai
n ju
nipe
r
Tree
Wid
eRe
p?
Ale
xand
er
(199
8b)
1998
Cype
race
aeCa
rex
nebr
asce
nsis
C. n
ebra
sken
sis
Neb
rask
a se
dge
PerG
Wid
ePr
esRa
reA
lexa
nder
12
44 (P
ISP)
1998
Cype
race
aeCa
rex
prae
grac
ilis
blac
kcre
eper
se
dge
PerG
Wid
ePr
esRa
reH
eato
n s.
n.
(PIS
P)19
38
App
endi
x A
. Con
fi rm
ed, h
isto
rica
l, an
d re
port
ed v
ascu
lar
plan
t ta
xa o
f Pi
pe S
prin
g N
atio
nal M
onum
ent,
org
aniz
ed b
y fa
mily
and
sci
enti
fi c
nam
e w
ith
taxo
nom
ic
and
biog
eogr
aphi
c no
tes,
con
t.
30 Annotated Checklist of Vascular Flora: Pipe Spring National Monument
Fam
ilySp
ecie
s na
me
Syno
nym
s/Ta
xono
mic
not
esCo
mm
onna
me
Life
fo
rmRa
nge
Park
st
atus
Pop.
si
zeSo
urce
Year
do
c.Co
mm
ents
Cype
race
aeCy
peru
s sp
.fl
at-s
edge
Ann
FIn
tro
Rep
?A
lexa
nder
(1
998b
)19
98Id
enti
fi ca
tion
ne
eded
Cype
race
aeEl
eoch
aris
par
ishi
iPa
rish
’s sp
iker
ush
PerG
Wid
ePr
esRa
reA
lexa
nder
12
42 (P
ISP)
1998
Cype
race
aeSc
irpu
s pu
ngen
s va
r. lo
ngis
pica
tus
Scri
pus
pung
ens
var.
poly
phyl
lus,
Sc
irpu
s am
eric
anus
va
r. lo
ngis
pica
tus,
Sc
hoen
ople
ctus
pu
ngen
s va
r. lo
ngis
pica
tus,
Sc
hoen
ople
ctus
pu
ngen
s va
r. po
lyph
yllu
s
thre
e-sq
uare
bu
lrus
hPe
rGW
ide
Pres
Rare
Ale
xand
er
1243
(PIS
P)19
98
Elae
agna
ceae
Shep
her
dia
ro
tund
ifol
iaro
undl
eaf
buff
alob
erry
Shru
bRe
gEn
Pres
Unc
Fert
ig 2
3667
(P
ISP)
1977
Type
loca
lity
is
uppe
r va
lley
of
the
Vir
gin,
Kan
e Co
., U
T (S
iler
s.n.
IS
C)
Ephe
drac
eae
Ephe
dra
torr
eyan
aTo
rrey
’s ep
hedr
aSh
rub
Wid
ePr
esRa
reFe
rtig
236
69
(PIS
P)20
07
Ephe
drac
eae
Ephe
dra
viri
dis
var.
viri
dis
E. v
irid
isgr
een
ephe
dra
Shru
bW
ide
Pres
Unc
Ale
xand
er
1201
(PIS
P)19
77
Ephe
drac
eae
Ephe
dra
viri
dis
var.
visc
ida
E. c
utle
riCu
tler
’s ep
hedr
aSh
rub
RegE
nPr
esU
ncG
anz
s.n.
(A
RIZ)
1965
Obs
erve
d ve
geta
tive
ly b
y Fe
rtig
, 200
7
Euph
orbi
acea
eC
ham
aesy
ce
albo
mar
gina
taEu
ph
orb
ia
albo
mar
gina
tara
ttle
snak
e-w
eed
PerF
Wid
eRe
p?
Ale
xand
er
(199
8b)
1998
Euph
orbi
acea
eCh
amae
syce
fen
dler
iEu
phor
bia
fend
leri
Fend
ler’s
spu
rge
PerF
Wid
ePr
esU
ncA
lexa
nder
12
03 (P
ISP)
1939
Euph
orbi
acea
eC
ham
aesy
ce
glyp
tosp
erm
aEu
ph
orb
ia
glyp
tosp
erm
ari
dge-
seed
ed
spur
geA
nnF
Wid
ePr
esCo
mH
eato
n s.
n.
(PIS
P)19
36
Euph
orbi
acea
eCh
amae
syce
pro
stra
taEu
phor
bia
pros
trat
a,
E. c
ham
aesy
cepr
ostr
ate
spur
geA
nnF
Intr
oPr
esU
ncFe
rtig
236
91
(PIS
P)20
07N
ativ
e to
eas
tern
N
orth
Am
eric
a
Faga
ceae
Que
rcus
tur
bine
llatu
rbin
ella
live
-oa
kSh
rub
Wid
ePr
esU
ncA
lexa
nder
12
49 (P
ISP)
1935
App
endi
x A
. Con
fi rm
ed, h
isto
rica
l, an
d re
port
ed v
ascu
lar
plan
t ta
xa o
f Pi
pe S
prin
g N
atio
nal M
onum
ent,
org
aniz
ed b
y fa
mily
and
sci
enti
fi c
nam
e w
ith
taxo
nom
ic
and
biog
eogr
aphi
c no
tes,
con
t.
Appendices 31
Fam
ilySp
ecie
s na
me
Syno
nym
s/Ta
xono
mic
not
esCo
mm
onna
me
Life
fo
rmRa
nge
Park
st
atus
Pop.
si
zeSo
urce
Year
do
c.Co
mm
ents
Fum
aria
ceae
Cory
dalis
aur
ea v
ar.
occi
dent
alis
C. c
urvi
siliq
ua s
sp.
occi
dent
alis
gold
en
cory
dalis
PerF
Wid
eH
ist
?H
eato
n s.
n.
(PIS
P)19
36
Gen
tian
acea
eSw
erti
a al
bom
argi
nata
Fras
era
albo
mar
gina
taw
hite
-mar
gine
d sw
erti
aPe
rFW
ide
His
t?
Peeb
les
1308
0 (A
RIZ)
1936
Type
loca
lity
is
near
St.
Geo
rge,
W
ashi
ngto
n Co
., U
T (P
alm
er s
.n.
US)
Gen
tian
acea
eSw
erti
a ut
ahen
sis
Fras
era
pani
cula
taU
tah
swer
tia
PerF
RegE
nRe
p?
Ale
xand
er
(199
8b)
1998
Type
loca
lity
is
Buck
skin
Mtn
s,
Kai
bab
Plat
eau
“on
the
sout
hern
ed
ge o
f U
tah”
, K
ane
Co.,
UT
(Jon
es s
.n. P
OM
)
Ger
ania
ceae
Erod
ium
cic
utar
ium
stor
k’s-
bill
Ann
FIn
tro
Pres
Com
Ale
xand
er
1213
(PIS
P)19
97N
ativ
e to
Eur
ope
Gra
min
eae
(Poa
ceae
)A
grop
yron
cri
stat
umA
. cri
stat
um s
sp.
cris
tatu
m, A
. cr
ista
tum
ssp
. pe
ctin
atum
, A.
pect
inat
um, A
. si
biri
cum
, A. f
ragi
le,
A. m
ongo
licum
, A.
dese
rtor
um
cres
ted
whe
atgr
ass
PerG
Intr
oRe
p?
Kin
g (1
977)
1977
Nat
ive
to E
uras
ia
Gra
min
eae
(Poa
ceae
)A
gros
tis
stol
onif
era
A. a
lba,
A. g
igan
tea
redt
opPe
rGIn
tro
Pres
Unc
Ale
xand
er
1241
(PIS
P)19
98N
ativ
e to
Eur
asia
an
d A
fric
a
Gra
min
eae
(Poa
ceae
)A
rist
ida
purp
urea
A. g
lauc
a, A
. pu
rpur
ea v
ar.
fend
leri
ana,
A.
fend
leri
ana,
A.
purp
urea
var
. lo
ngis
eta
purp
le t
hree
-aw
nPe
rGW
ide
Pres
Com
Ale
xand
er
1210
(PIS
P)19
41
Gra
min
eae
(Poa
ceae
)Bo
utel
oua
barb
ata
Chon
dros
um
barb
atum
. Inc
lude
s va
rs. b
arba
ta &
ro
thro
ckii
sixw
eeks
gra
ma
Ann
GW
ide
Pres
Unc
Fert
ig 2
3671
(P
ISP)
1935
App
endi
x A
. Con
fi rm
ed, h
isto
rica
l, an
d re
port
ed v
ascu
lar
plan
t ta
xa o
f Pi
pe S
prin
g N
atio
nal M
onum
ent,
org
aniz
ed b
y fa
mily
and
sci
enti
fi c
nam
e w
ith
taxo
nom
ic
and
biog
eogr
aphi
c no
tes,
con
t.
32 Annotated Checklist of Vascular Flora: Pipe Spring National Monument
Fam
ilySp
ecie
s na
me
Syno
nym
s/Ta
xono
mic
not
esCo
mm
onna
me
Life
fo
rmRa
nge
Park
st
atus
Pop.
si
zeSo
urce
Year
do
c.Co
mm
ents
Gra
min
eae
(Poa
ceae
)B
ou
telo
ua
curt
ipen
dula
Incl
udes
var
s.
curt
ipen
dula
&
caes
pito
sa
side
oats
gra
ma
PerG
Wid
eRe
p?
Kin
g (1
977)
1977
Gra
min
eae
(Poa
ceae
)Bo
utel
oua
erio
poda
Chon
dros
um
erio
podu
mbl
ack
gram
aPe
rGW
ide
Pres
Unc
Ale
xand
er
1250
(PIS
P)19
77
Gra
min
eae
(Poa
ceae
)Bo
utel
oua
grac
ilis
Chon
dros
um g
raci
lebl
ue g
ram
aPe
rGW
ide
Pres
Unc
Purc
hase
272
(A
RIZ)
1936
Gra
min
eae
(Poa
ceae
)Br
omus
dia
ndru
sB.
rig
idus
, Ani
sant
ha
dian
dra
ripg
ut b
rom
eA
nnG
Intr
oPr
esU
ncLo
uie
&
Even
den
PISP
22
(PIS
P)
1977
Nat
ive
to E
uras
ia
Gra
min
eae
(Poa
ceae
)Br
omus
iner
mis
var
. in
erm
isBr
omop
sis
iner
mis
, Br
omus
iner
mis
smoo
th b
rom
ePe
rGIn
tro
Rep
?K
ing
(197
7)19
77N
ativ
e to
Eur
asia
Gra
min
eae
(Poa
ceae
)Br
omus
rub
ens
red
brom
eA
nnG
Intr
oPr
esCo
mLo
uie
&
Even
den
PISP
34
(PIS
P)
1998
Nat
ive
to E
uras
ia
Gra
min
eae
(Poa
ceae
)Br
omus
tec
toru
mA
nisa
ntha
tec
toru
mch
eatg
rass
Ann
GIn
tro
Pres
Com
Hea
ton
1941
-14
(PIS
P)19
38N
ativ
e to
Eur
asia
Gra
min
eae
(Poa
ceae
)Ce
nchr
us lo
ngis
pinu
sC.
pau
cifl
orus
fi el
d sa
ndbu
rA
nnG
Wid
eRe
p?
Kur
th (1
998)
1997
Gra
min
eae
(Poa
ceae
)D
isti
chlis
spi
cata
D
. spi
cata
var
. str
icta
, D
. str
icta
dese
rt s
altg
rass
PerG
Wid
eRe
p?
Kin
g (1
977)
, A
lexa
nder
(1
998b
)
1977
Gra
min
eae
(Poa
ceae
)Ec
hino
chlo
a cr
us-g
alli
Incl
udes
E.
mic
rost
achy
a,
E. m
uric
ata
var.
mic
rost
achy
a
barn
yard
-gra
ssA
nnG
Intr
oRe
p?
Ale
xand
er
(199
8b)
1998
Nat
ive
to E
uras
ia
Gra
min
eae
(Poa
ceae
)El
ymus
elo
ngat
us
E. e
long
atus
var
. po
ntic
us, A
grop
yron
el
onga
tum
, Th
inop
yrum
po
ntic
um
tall
whe
atgr
ass
PerG
Intr
oRe
p?
Kur
th (1
998)
1997
Nat
ive
to E
uras
ia
Gra
min
eae
(Poa
ceae
)El
ymus
ely
moi
des
Sita
nion
hys
trix
, E.
ely
moi
des
var.
elym
oide
s
squi
rrel
tail
PerG
Wid
eRe
p?
Kin
g (1
977)
, A
lexa
nder
(1
998b
)
1977
App
endi
x A
. Con
fi rm
ed, h
isto
rica
l, an
d re
port
ed v
ascu
lar
plan
t ta
xa o
f Pi
pe S
prin
g N
atio
nal M
onum
ent,
org
aniz
ed b
y fa
mily
and
sci
enti
fi c
nam
e w
ith
taxo
nom
ic
and
biog
eogr
aphi
c no
tes,
con
t.
Appendices 33
Fam
ilySp
ecie
s na
me
Syno
nym
s/Ta
xono
mic
not
esCo
mm
onna
me
Life
fo
rmRa
nge
Park
st
atus
Pop.
si
zeSo
urce
Year
do
c.Co
mm
ents
Gra
min
eae
(Poa
ceae
)El
ymus
his
pidu
s A
grop
yron
in
term
ediu
m,
Elyt
rigi
a in
term
edia
, E.
his
pidu
s va
r. hi
spid
us, T
hino
pyru
m
inte
rmed
ium
inte
rmed
iate
w
heat
gras
sPe
rGIn
tro
Rep
?K
ing
(197
7)19
77N
ativ
e to
Eur
asia
Gra
min
eae
(Poa
ceae
)El
ymus
sm
ithi
iA
grop
yron
sm
ithi
i, Pa
scop
yrum
sm
ithi
i; in
clud
es v
ars.
sm
ith
ii &
pal
mer
i
wes
tern
w
heat
gras
sPe
rGW
ide
Rep
?K
ing
(197
7),
Ale
xand
er
(199
8b)
1977
Gra
min
eae
(Poa
ceae
)El
ymus
tra
chyc
aulu
s E.
tra
chyc
aulu
s va
r. tr
achy
caul
us,
E. a
lask
anus
ss
p. la
tigl
umis
, A
grop
yron
can
inum
, A
. sub
secu
ndum
, A
. tra
chyc
aulu
m, A
. la
tigl
ume
slen
der
whe
atgr
ass
PerG
Wid
eH
ist
?H
eato
n s.
n.
(PIS
P)19
38
Gra
min
eae
(Poa
ceae
)Er
agro
stis
cili
anen
sis
stin
kgra
ssA
nnG
Intr
oRe
p?
Kin
g (1
977)
1977
Nat
ive
to E
uras
ia
Gra
min
eae
(Poa
ceae
)Er
agro
stis
mex
ican
aE.
orc
utti
ana
Mex
ican
lo
vegr
ass
Ann
GIn
tro
Rep
?A
lexa
nder
(1
998b
)19
98N
ativ
e to
w
este
rn N
orth
A
mer
ica
Gra
min
eae
(Poa
ceae
)Er
agro
stis
pec
tina
cea
E. d
iffu
satu
fted
love
gras
sA
nnG
Wid
ePr
esU
ncFe
rtig
236
82
(PIS
P)19
77
Gra
min
eae
(Poa
ceae
)Er
ion
euro
n
pulc
hellu
mD
asyo
chlo
a pu
lche
lla,
Trid
ens
pulc
hellu
sfl
uffg
rass
PerG
Wid
eRe
p?
Hea
ton
(193
5)19
35
Gra
min
eae
(Poa
ceae
)Fe
stuc
a ar
undi
nace
aSc
hedo
noru
s p
ho
enix
, S.
arun
dina
ceus
, Lol
ium
ar
undi
nace
um
tall
fesc
uePe
rGIn
tro
Rep
?K
ing
(197
7),
Ale
xand
er
(199
8b)
1977
Nat
ive
to E
urop
e
Gra
min
eae
(Poa
ceae
)H
ilari
a ja
mes
iiPl
eura
phis
jam
esii
galle
taPe
rGW
ide
Pres
Unc
Ale
xand
er
1209
(PIS
P)19
36
Gra
min
eae
(Poa
ceae
)H
orde
um ju
batu
mCr
ites
ion
juba
tum
foxt
ail b
arle
yPe
rGW
ide
Rep
?K
ing
(197
7)19
77
App
endi
x A
. Con
fi rm
ed, h
isto
rica
l, an
d re
port
ed v
ascu
lar
plan
t ta
xa o
f Pi
pe S
prin
g N
atio
nal M
onum
ent,
org
aniz
ed b
y fa
mily
and
sci
enti
fi c
nam
e w
ith
taxo
nom
ic
and
biog
eogr
aphi
c no
tes,
con
t.
34 Annotated Checklist of Vascular Flora: Pipe Spring National Monument
Fam
ilySp
ecie
s na
me
Syno
nym
s/Ta
xono
mic
not
esCo
mm
onna
me
Life
fo
rmRa
nge
Park
st
atus
Pop.
si
zeSo
urce
Year
do
c.Co
mm
ents
Gra
min
eae
(Poa
ceae
)H
orde
um m
urin
umH
. lep
orin
um,
H. m
urin
um s
sp.
lepo
rinu
m, C
rite
sion
gl
aucu
m
rabb
it b
arle
yA
nnG
Intr
oPr
esU
ncH
eato
n 19
41-3
(P
ISP)
1938
Gra
min
eae
(Poa
ceae
)M
uhle
nber
gia
aspe
rifo
liasc
ratc
hgra
ssPe
rGW
ide
Pres
Unc
Ale
xand
er
1223
(PIS
P)19
77
Gra
min
eae
(Poa
ceae
)M
uhle
nber
gia
port
eri
bush
muh
lyPe
rGW
ide
Pres
Unc
Fert
ig 2
3680
(P
ISP)
1977
Gra
min
eae
(Poa
ceae
)M
unro
a sq
uarr
osa
Mon
roa
squa
rros
afa
lse
buff
alog
rass
Ann
GW
ide
Pres
Rare
Fert
ig 2
3670
(P
ISP)
2007
Gra
min
eae
(Poa
ceae
)Po
a fe
ndle
rian
aP.
long
iligu
la, P
. fe
ndle
rian
a ss
p.
long
iligu
la
mut
tong
rass
PerG
Wid
eRe
p?
Ale
xand
er
(199
8b)
1998
Gra
min
eae
(Poa
ceae
)Po
a pr
aten
sis
Poa
agas
size
nsis
Ken
tuck
y bl
uegr
ass
PerG
Intr
oRe
p?
Kin
g (1
977)
, A
lexa
nder
(1
998b
)
1977
Nat
ive
to E
urop
e
Gra
min
eae
(Poa
ceae
)Po
lyp
og
on
m
onsp
elie
nsis
rabb
itsf
oot-
gras
sA
nnG
Intr
oPr
esU
ncFe
rtig
236
86
(PIS
P)19
98N
ativ
e to
Eur
asia
an
d A
fric
a
Gra
min
eae
(Poa
ceae
)Po
lyp
og
on
se
miv
erti
cilla
tus
Ag
rost
is
sem
iver
tici
llata
, P.
viri
dis
wat
er
poly
pogo
nPe
rGIn
tro
Rep
?A
lexa
nder
(1
998b
)19
98N
ativ
e to
Eur
asia
an
d A
fric
a
Gra
min
eae
(Poa
ceae
)Sp
orob
olus
air
oide
s va
r. ai
roid
esS.
air
oide
sal
kali
saca
ton
PerG
Wid
eRe
p?
Kin
g (1
977)
, A
lexa
nder
(1
998b
)
1977
Gra
min
eae
(Poa
ceae
)Sp
orob
olus
con
trac
tus
spik
e dr
opse
edPe
rGW
ide
Pres
Unc
Ale
xand
er
1221
(PIS
P)19
36
Gra
min
eae
(Poa
ceae
)Sp
oro
bo
lus
cryp
tand
rus
sand
dro
psee
dPe
rGW
ide
Pres
Unc
Purc
hase
271
(A
RIZ)
1936
Gra
min
eae
(Poa
ceae
)Sp
orob
olus
fl ex
uosu
sm
esa
drop
seed
PerG
Wid
ePr
esU
ncA
lexa
nder
12
20 (P
ISP)
1977
Gra
min
eae
(Poa
ceae
)St
ipa
com
ata
var.
com
ata
Hes
pero
stip
a co
mat
a va
r. co
mat
ane
edle
-and
-th
read
PerG
Wid
ePr
esU
ncLo
uie
&
Even
den
PISP
28
(PIS
P)
1977
Gra
min
eae
(Poa
ceae
)St
ipa
hym
enoi
des
Ory
zops
is
hym
eno
ides
, A
chn
ath
eru
m
hym
enoi
des
Indi
an r
iceg
rass
PerG
Wid
ePr
esU
ncK
ing
(197
7)19
77O
bser
ved
vege
tati
vely
by
Fert
ig, 2
007
App
endi
x A
. Con
fi rm
ed, h
isto
rica
l, an
d re
port
ed v
ascu
lar
plan
t ta
xa o
f Pi
pe S
prin
g N
atio
nal M
onum
ent,
org
aniz
ed b
y fa
mily
and
sci
enti
fi c
nam
e w
ith
taxo
nom
ic
and
biog
eogr
aphi
c no
tes,
con
t.
Appendices 35
Fam
ilySp
ecie
s na
me
Syno
nym
s/Ta
xono
mic
not
esCo
mm
onna
me
Life
fo
rmRa
nge
Park
st
atus
Pop.
si
zeSo
urce
Year
do
c.Co
mm
ents
Gra
min
eae
(Poa
ceae
)St
ipa
spec
iosa
Ach
nath
erum
sp
ecio
sum
, Jar
ava
spec
iosu
m
dese
rt
need
legr
ass
PerG
Wid
ePr
esU
ncLo
uie
&
Even
den
PISP
23
(PIS
P)
1977
Hyd
roph
ylla
ceae
Phac
elia
cre
nula
ta v
ar.
corr
ug
ata
P. c
orru
gata
corr
ugat
e ph
acel
iaA
nnF
Wid
eH
ist
?H
eato
n s.
n.
(PIS
P)19
37
Hyd
roph
ylla
ceae
Phac
elia
cre
nula
ta v
ar.
cren
ulat
aP.
cre
nula
tacr
enul
ate
phac
elia
Ann
FW
ide
Pres
Unc
Loui
e &
Ev
ende
n PI
SP 6
(P
ISP)
2001
Hyd
roph
ylla
ceae
Phac
elia
ives
iana
Ives
’ pha
celia
Ann
FW
ide
Pres
Unc
Loui
e &
Ev
ende
n PI
SP
12 (P
ISP)
1937
Hyd
roph
ylla
ceae
Phac
elia
pul
chel
la v
ar.
good
ding
iiG
oodd
ing’
s pr
etty
pha
celia
Ann
FRe
gEn
His
t?
Hea
ton
7 (P
ISP)
1937
Irid
acea
eSi
syri
nchi
um
dem
issu
mbl
ue-e
yed
gras
sPe
rFW
ide
Rep
?A
lexa
nder
(1
998b
)19
98
Junc
acea
eJu
ncus
arc
ticu
sJ.
bal
ticu
s va
r. m
onta
nus,
J.
arct
icus
var
. bal
ticu
s,
J. a
rcti
cus
ssp.
at
er; i
nclu
des
var.
mex
ican
us
Balt
ic r
ush
PerG
Wid
ePr
esU
ncEv
ende
n &
Lo
uie
33 (P
ISP)
1998
Junc
acea
eJu
ncus
ens
ifol
ius
var.
brun
nesc
ens
J. s
axim
onta
nus
form
a br
unne
scen
s.
Incl
uded
in J
. en
sifo
lius
var.
mon
tanu
s by
som
e au
thor
s
Trac
y’s
rush
PerG
Wid
eH
ist
?H
eato
n s.
n.
(PIS
P)19
38
Junc
acea
eJu
ncus
tor
reyi
Torr
ey’s
rush
PerG
Wid
ePr
esU
ncFe
rtig
236
93
(PIS
P)19
98
Labi
atae
(L
amia
ceae
)M
arru
bium
vul
gare
com
mon
ho
reho
und
PerF
Intr
oPr
esU
ncA
lexa
nder
12
35 (P
ISP)
1935
Nat
ive
to E
uras
ia
Labi
atae
(L
amia
ceae
)M
enth
a sp
icat
asp
earm
int
PerF
Intr
oPr
esRa
reA
lexa
nder
11
90 (P
ISP)
1998
Nat
ive
to E
urop
e
Labi
atae
(L
amia
ceae
)Sa
lvia
dor
rii s
sp. d
orri
iS.
car
nosa
Dor
r’s s
age
Shru
bW
ide
Pres
Unc
Hea
ton
1 (P
ISP)
1937
Legu
min
osae
(F
abac
eae)
Ast
raga
lus
amph
ioxy
s va
r. ve
sper
tinu
sSh
eldo
n’s
milk
vetc
hPe
rFRe
gEn
His
t?
Hea
ton
s.n.
(P
ISP)
1934
App
endi
x A
. Con
fi rm
ed, h
isto
rica
l, an
d re
port
ed v
ascu
lar
plan
t ta
xa o
f Pi
pe S
prin
g N
atio
nal M
onum
ent,
org
aniz
ed b
y fa
mily
and
sci
enti
fi c
nam
e w
ith
taxo
nom
ic
and
biog
eogr
aphi
c no
tes,
con
t.
36 Annotated Checklist of Vascular Flora: Pipe Spring National Monument
Fam
ilySp
ecie
s na
me
Syno
nym
s/Ta
xono
mic
not
esCo
mm
onna
me
Life
fo
rmRa
nge
Park
st
atus
Pop.
si
zeSo
urce
Year
do
c.Co
mm
ents
Legu
min
osae
(F
abac
eae)
Ast
raga
lus
cera
mic
us
var.
cera
mic
uspa
inte
d m
ilkve
tch
PerF
Wid
ePr
esU
ncEv
ende
n &
Lo
uie
PISP
21
(PIS
P)
1998
Legu
min
osae
(F
abac
eae)
Ast
raga
lus
prae
long
us
var.
prae
long
usst
inki
ng
milk
vetc
hPe
rFW
ide
Rep
?A
lexa
nder
&
Dec
ker
1444
(P
ISP)
1998
Legu
min
osae
(F
abac
eae)
Gly
cyrr
hiza
lepi
dota
G
. lep
idot
a va
r. le
pido
taN
utta
ll’s
licor
ice
PerF
Wid
eRe
p?
Ale
xand
er
(199
8b)
1998
Legu
min
osae
(F
abac
eae)
Lupi
nus
pusi
llus
var.
pusi
llus
L. p
usill
us s
sp.
pusi
llus
dwar
f lu
pine
Ann
FW
ide
His
t?
Hea
ton
2 (P
ISP)
1937
Legu
min
osae
(F
abac
eae)
Med
icag
o sa
tiva
M. s
ativ
a ss
p. s
ativ
aal
falf
aPe
rFIn
tro
Rep
?K
urth
(199
8)19
97N
ativ
e to
Eur
ope
Legu
min
osae
(F
abac
eae)
Mel
ilotu
s al
baM
. alb
us, i
nclu
ded
in M
. offi
cin
alis
by
som
e au
thor
s
whi
te
swee
tclo
ver
Ann
FIn
tro
Rep
?K
ing
(197
7),
Ale
xand
er
(199
8b)
1977
Nat
ive
to E
urop
e
Legu
min
osae
(F
abac
eae)
Mel
ilotu
s of
fi ci
nalis
yello
w
swee
tclo
ver
PerF
Intr
oRe
p?
Ale
xand
er
(199
8b)
1998
Nat
ive
to E
urop
e
Legu
min
osae
(F
abac
eae)
Pete
ria
thom
pson
iae
Thom
pson
’s pe
teri
aPe
rFRe
gEn
Pres
Rare
Ale
xand
er
1251
(PIS
P)20
01Ty
pe lo
calit
y is
K
anab
, Kan
e Co
., U
T (T
hom
pson
s.
n. G
H)
Legu
min
osae
(F
abac
eae)
Robi
nia
pseu
doac
acia
blac
k lo
cust
Tree
Intr
oPr
esU
ncA
lexa
nder
12
56 (P
ISP)
1998
Nat
ive
to e
aste
rn
Nor
th A
mer
ica
Lilia
ceae
(T
hem
idac
eae)
And
rost
ephi
um
brev
ifl or
umBr
odia
ea p
ayso
nii
pink
fun
nel-l
ilyPe
rFW
ide
His
t?
Peeb
les
1307
8 (U
niv
Ari
zona
)19
36Ty
pe lo
calit
y is
nea
r K
anab
, K
ane
Co.,
UT
(Tho
mps
on s
.n.)
Lilia
ceae
(C
aloc
hort
acea
e)Ca
loch
ortu
s fl
exuo
sus
sinu
ous
mar
ipos
aPe
rFW
ide
Pres
Unc
Loui
e &
Ev
ende
n PI
SP 1
(P
ISP)
1935
Type
loca
lity
is n
ear
Kan
ab,
Kan
e Co
., U
T (T
hom
pson
s.n
. G
H)
Lina
ceae
Linu
m a
rist
atum
Mes
yniu
m a
rist
atum
broo
mfl
axA
nnF
Wid
ePr
esU
ncFe
rtig
236
75
(PIS
P)19
98
Loas
acea
eM
entz
elia
alb
icau
lisM
. obs
cura
, Acr
olas
ia
albi
caul
us, A
. gra
cilis
whi
te-s
tem
bl
azin
gsta
rA
nnF
Wid
ePr
esU
ncEv
ende
n &
Lo
uie
PISP
17
(PIS
P)
1937
App
endi
x A
. Con
fi rm
ed, h
isto
rica
l, an
d re
port
ed v
ascu
lar
plan
t ta
xa o
f Pi
pe S
prin
g N
atio
nal M
onum
ent,
org
aniz
ed b
y fa
mily
and
sci
enti
fi c
nam
e w
ith
taxo
nom
ic
and
biog
eogr
aphi
c no
tes,
con
t.
Appendices 37
Fam
ilySp
ecie
s na
me
Syno
nym
s/Ta
xono
mic
not
esCo
mm
onna
me
Life
fo
rmRa
nge
Park
st
atus
Pop.
si
zeSo
urce
Year
do
c.Co
mm
ents
Loas
acea
eM
entz
elia
inte
gra
M. m
ulti
fl or
a va
r. in
tegr
a, M
. pum
ila
var.
inte
gra
Vir
gin
stic
klea
fPe
rFW
ide
His
t?
Leht
o 70
37
(ARI
Z)19
66Ty
pe lo
calit
y is
Roc
kvill
e,
Was
hing
ton
Co.,
UT
(Jon
es 6
082c
)
Loas
acea
eM
entz
elia
mul
tifl
ora
M. p
umila
var
. m
ulti
fl o
ra, M
. m
ulti
fl or
a va
r. m
ulti
fl or
a, N
utta
llia
mul
tifl
ora
dese
rt s
tick
leaf
PerF
Wid
ePr
esU
ncA
lexa
nder
12
14 (P
ISP)
1998
Mal
vace
aeA
ltha
ea r
osea
Alc
ea r
osea
holly
hock
PerF
Intr
oRe
p?
Kur
th (1
998)
1997
Nat
ive
to C
hina
Mal
vace
aeM
alva
neg
lect
aco
mm
on
mal
low
Ann
FIn
tro
Pres
Com
Ale
xand
er
1231
(PIS
P)19
97N
ativ
e to
Eur
asia
Mal
vace
aeSp
haer
alce
a am
bigu
a va
r. am
bigu
aM
ohav
e gl
obem
allo
wPe
rFW
ide
Pres
Unc
Ale
xand
er
1255
(PIS
P)19
35
Mal
vace
aeSp
haer
alce
a pa
rvif
olia
smal
l-lea
f gl
obem
allo
wPe
rFW
ide
His
t?
Whi
tehe
ad s
.n.
(PIS
P)19
36
Nyc
tagi
nace
aeA
bron
ia f
ragr
ans
A. f
ragr
ans
var.
ellip
tica
, A. n
ana
var.
harr
isii,
incl
udes
A.
ellip
tica
frag
rant
san
d-ve
rben
aPe
rFW
ide
Pres
Unc
Hea
ton
20
(PIS
P)19
37O
bser
ved
vege
tati
vely
by
Fert
ig, 2
007
Nyc
tagi
nace
aeM
irab
ilis
mul
tifl
ora
M. m
ulti
fl o
ra
var.
mul
tifl
ora
, Q
uam
oclid
ion
mul
tifl
orum
show
y fo
ur-
o’cl
ock
PerF
Wid
ePr
esU
ncLo
uie
&
Even
den
PISP
18
(PIS
P)
1935
Ole
acea
eM
enod
ora
scab
raro
ugh
men
odor
aPe
rFW
ide
Pres
Unc
Ale
xand
er
1200
(PIS
P)19
98
Ona
grac
eae
Cam
isso
nia
ch
amae
neri
oide
sO
eno
ther
a ch
amae
neri
oide
ssl
ende
r-po
d ca
mis
soni
aA
nnF
Wid
ePr
esU
ncEv
ende
n &
Lo
uie
PISP
24
(PIS
P)
2001
Ona
grac
eae
Cam
isso
nia
parr
yiO
enot
hera
par
ryi
Parr
y’s
cam
isso
nia
Ann
FRe
gEn
His
t?
Hea
ton
25
(PIS
P)19
36Ty
pe lo
calit
y is
ne
ar S
t. G
eorg
e,
Was
hing
ton
Co.,
UT
(Par
ry 7
2 G
H)
Ona
grac
eae
Cam
isso
nia
parv
ula
Oen
othe
ra c
onto
rta
var.
fl ex
uosa
tiny
cam
isso
nia
Ann
FPe
riph
His
t?
Hea
ton
s.n.
(P
ISP)
1936
App
endi
x A
. Con
fi rm
ed, h
isto
rica
l, an
d re
port
ed v
ascu
lar
plan
t ta
xa o
f Pi
pe S
prin
g N
atio
nal M
onum
ent,
org
aniz
ed b
y fa
mily
and
sci
enti
fi c
nam
e w
ith
taxo
nom
ic
and
biog
eogr
aphi
c no
tes,
con
t.
38 Annotated Checklist of Vascular Flora: Pipe Spring National Monument
Fam
ilySp
ecie
s na
me
Syno
nym
s/Ta
xono
mic
not
esCo
mm
onna
me
Life
fo
rmRa
nge
Park
st
atus
Pop.
si
zeSo
urce
Year
do
c.Co
mm
ents
Ona
grac
eae
Epilo
bium
cili
atum
E. c
iliat
um v
ar.
cilia
tum
, E. w
atso
nii,
E. a
deno
caul
on
nort
hern
w
illow
-her
b Pe
rFW
ide
Rep
?A
lexa
nder
(1
998b
)19
98
Ona
grac
eae
Gau
ra c
occi
nea
scar
let
gaur
aPe
rFW
ide
His
t?
Hea
ton
s.n.
(P
ISP)
1939
Ona
grac
eae
Gau
ra p
arvi
fl or
aG
. mol
lissm
all-fl
ow
ered
ga
ura
Ann
FW
ide
Pres
Unc
Ale
xand
er
1224
(PIS
P)19
98
Ona
grac
eae
Oen
othe
ra a
lbic
aulis
whi
te-s
tem
ev
enin
g pr
imro
se
Ann
FW
ide
His
t?
Hea
ton
s.n.
(P
ISP)
1936
Ona
grac
eae
Oen
othe
ra c
aesp
itos
a va
r. na
vajo
ensi
sPa
ria
even
ing
prim
rose
PerF
RegE
nPr
esU
ncEv
ende
n &
Lo
uie
PISP
20
(PIS
P)
1998
Type
loca
lity
is P
aria
Riv
er,
Kan
e Co
., U
T (C
ronq
uist
102
02
MO
)
Ona
grac
eae
Oen
othe
ra c
alif
orni
caCa
lifor
nia
even
ing
prim
rose
PerF
Wid
ePr
esU
ncA
lexa
nder
12
05 (P
ISP)
2001
Ona
grac
eae
Oen
othe
ra p
allid
a va
r. pa
llida
Incl
udes
var
. ru
ncin
ata
pale
eve
ning
pr
imro
sePe
rFW
ide
Pres
Unc
Ale
xand
er
1206
(PIS
P)19
36
Ona
grac
eae
Oen
othe
ra p
rim
iver
is
var.
bufo
nus
Oen
othe
ra p
rim
iver
isea
rly
even
ing
prim
rose
Ann
FRe
gEn
Rep
?A
lexa
nder
(1
998b
)N
A
Oro
banc
hace
aeO
rob
anch
e lu
dovi
cian
aO
. lud
ovic
iana
var
. lu
dovi
cian
a, O
. co
oper
i; in
clud
es O
. m
ulti
fl or
a fr
om U
tah
Fl
ora
(199
3) a
nd v
ars.
co
oper
i & a
rane
osa
Coop
er’s
broo
mra
pePe
rFW
ide
Pres
Unc
Whi
tehe
ad s
.n.
(PIS
P)19
35
Papa
vera
ceae
Esch
scho
lzia
m
inut
ifl or
apy
gmy
gold
en-
popp
yA
nnF
Wid
ePr
esU
ncLo
uie
&
Even
den
PISP
5
(PIS
P)
1935
Pina
ceae
Pinu
s ed
ulis
two-
need
le
piny
on
Tree
Wid
ePr
esU
ncA
lexa
nder
12
48 (P
ISP)
1935
Plan
tagi
nace
aePl
anta
go la
nceo
lata
Engl
ish
plan
tain
PerF
Intr
oRe
p?
Ale
xand
er
(199
8b)
1998
Nat
ive
to E
uras
ia
Plan
tagi
nace
aePl
anta
go m
ajor
Incl
udes
var
. pa
chyp
hylla
com
mon
pl
anta
inPe
rFIn
tro
Pres
Rare
Ale
xand
er
1189
(PIS
P)19
35M
ostl
y na
tive
to
Euro
pe
App
endi
x A
. Con
fi rm
ed, h
isto
rica
l, an
d re
port
ed v
ascu
lar
plan
t ta
xa o
f Pi
pe S
prin
g N
atio
nal M
onum
ent,
org
aniz
ed b
y fa
mily
and
sci
enti
fi c
nam
e w
ith
taxo
nom
ic
and
biog
eogr
aphi
c no
tes,
con
t.
Appendices 39
Fam
ilySp
ecie
s na
me
Syno
nym
s/Ta
xono
mic
not
esCo
mm
onna
me
Life
fo
rmRa
nge
Park
st
atus
Pop.
si
zeSo
urce
Year
do
c.Co
mm
ents
Pole
mon
iace
aeEr
iast
rum
ere
mic
umM
ohav
e er
iast
rum
Ann
FW
ide
His
t?
Hea
ton
s.n.
(P
ISP)
1941
Pole
mon
iace
aeG
ilia
inco
nspi
cua
Incl
udes
var
s.
inco
nspi
ca &
sin
uat
a,
G. o
phth
alm
oide
s, G
. si
nuat
a, G
ilia
clok
eyi
shy
gilia
Ann
FW
ide
Pres
Unc
Even
den
&
Loui
e PI
SP 1
3 (P
ISP)
1936
If r
ecog
nize
d,
plan
ts f
rom
PIS
P re
pres
ent
vars
. in
cons
picu
a an
d si
nuat
a
Pole
mon
iace
aeG
ilia
lept
omer
ia v
ar.
lept
omer
iaA
licie
lla le
ptom
eria
com
mon
gili
aA
nnF
Wid
eH
ist
?H
eato
n s.
n.
(PIS
P)19
37
Pole
mon
iace
aeIp
omop
sis
long
ifl or
aG
ilia
long
ifl or
alo
ng-fl
ow
ered
gi
liaPe
rFW
ide
Pres
Unc
Ale
xand
er
1211
(PIS
P)19
36
Pole
mon
iace
aeIp
omop
sis
poly
clad
onG
ilia
poly
clad
onsp
read
ing
gilia
Ann
FW
ide
His
t?
Hea
ton
30
(PIS
P)19
37
Pole
mon
iace
aeLa
nglo
isia
set
osis
sim
aM
ohav
e la
nglo
isia
Ann
FW
ide
His
t?
Hea
ton
21
(PIS
P)19
37
Poly
gala
ceae
Poly
gala
sub
spin
osa
P. s
ubsp
inos
a va
r. su
bspi
nosa
cush
ion
milk
wor
tPe
rFW
ide
His
t?
Hea
ton
s.n.
(P
ISP)
1939
Poly
gona
ceae
Cent
rost
egia
thu
rber
iCh
oriz
anth
e th
urbe
riTh
urbe
r’s
spin
efl o
wer
Ann
FW
ide
His
t?
Hea
ton
19
(PIS
P)19
37
Poly
gona
ceae
Erio
gonu
m c
ernu
um
var.
cern
uum
Var
s. n
ot r
ecog
nize
d in
Flo
ra o
f N
orth
A
mer
ica
(200
5)
nodd
ing
wild
bu
ckw
heat
Ann
FW
ide
His
t?
Hea
ton
s.n.
(P
ISP)
1936
Poly
gona
ceae
Erio
go
nu
m
cory
mbo
sum
var
. au
reum
E. a
ureu
m, i
nclu
des
var.
glut
inos
umgo
lden
wild
bu
ckw
heat
Shru
bRe
gEn
Rep
?K
ing
(197
7),
Ale
xand
er
(199
8b)
1977
Type
loca
lity
is S
t. G
eorg
e,
Was
hing
ton
Co.,
UT
(Jon
es
6091
PO
M).
Cons
ider
ed a
lo
cal e
ndem
ic
rest
rict
ed t
o th
e Sh
ivw
its
area
of
Was
hing
ton
Co.
in F
lora
of
Nor
th
Am
eric
a (2
005)
.
App
endi
x A
. Con
fi rm
ed, h
isto
rica
l, an
d re
port
ed v
ascu
lar
plan
t ta
xa o
f Pi
pe S
prin
g N
atio
nal M
onum
ent,
org
aniz
ed b
y fa
mily
and
sci
enti
fi c
nam
e w
ith
taxo
nom
ic
and
biog
eogr
aphi
c no
tes,
con
t.
40 Annotated Checklist of Vascular Flora: Pipe Spring National Monument
Fam
ilySp
ecie
s na
me
Syno
nym
s/Ta
xono
mic
not
esCo
mm
onna
me
Life
fo
rmRa
nge
Park
st
atus
Pop.
si
zeSo
urce
Year
do
c.Co
mm
ents
Poly
gona
ceae
Erio
go
nu
m
cory
mbo
sum
var
. co
rym
bosu
m
Incl
udes
var
s.
velu
tinu
m,
diva
rica
tum
, er
ectu
m, &
dav
idse
an
d E.
lanc
ifol
ium
cris
plea
f w
ild
buck
whe
atSh
rub
Wid
ePr
esU
ncA
lexa
nder
12
28 (P
ISP)
2001
Poly
gona
ceae
Erio
gonu
m d
efl e
xum
E. d
efl e
xum
var
. de
fl ex
umsk
elet
onw
eed
wild
buc
kwhe
atA
nnF
Wid
eH
ist
?H
eato
n s.
n.
(PIS
P)19
36
Poly
gona
ceae
Erio
gonu
m in
fl at
um
va
r. in
fl at
umE.
infl
atum
dese
rt t
rum
pet
PerF
Wid
ePr
esU
ncEv
ende
n &
Lo
uie
PISP
36
(PIS
P)
1935
Poly
gona
ceae
Erio
go
nu
m
mic
roth
ecum
var
. si
mps
onii
E. s
imps
onii,
E.
mic
roth
ecum
var
. fo
liosu
m; i
nclu
des
E.
phoe
nice
um
slen
der
wild
bu
ckw
heat
PerF
Wid
ePr
esU
ncA
lexa
nder
12
27 (P
ISP)
1977
Poly
gona
ceae
Erio
go
nu
m
palm
eria
num
Palm
er’s
wild
bu
ckw
heat
Ann
FW
ide
Pres
Unc
Loui
e &
Ev
ende
n PI
SP 7
(P
ISP)
2001
Poly
gona
ceae
Poly
go
nu
m
amph
ibiu
mP.
am
phib
ium
va
r. em
ersu
m, P
. co
ccin
eum
, Per
sica
ria
cocc
inea
, Per
sica
ria
amph
ibia
wat
er
smar
twee
dPe
rFW
ide
Rep
?H
eato
n (1
935)
1935
Poly
gona
ceae
Poly
gonu
m a
vicu
lare
P. a
rena
stru
m,
P. a
vicu
lare
ssp
. av
icul
are
yard
kno
twee
dA
nnF
Intr
oPr
esU
ncA
lexa
nder
12
30 (P
ISP)
1998
Nat
ive
to E
uras
ia.
PISP
spe
cim
en
is “
aren
astr
um”
phas
e.
Poly
gona
ceae
Rum
ex c
risp
uscu
rly
dock
PerF
Intr
oRe
p?
Kur
th (1
998)
1997
Nat
ive
to E
uras
ia
Poly
gona
ceae
Rum
ex h
ymen
osep
alus
cana
igre
doc
kPe
rFW
ide
Pres
Unc
Fert
ig 2
3672
(P
ISP)
1998
Port
ulac
acea
ePo
rtul
aca
oler
acea
com
mon
pu
rsla
neA
nnF
Intr
oPr
esCo
mFe
rtig
236
63
(PIS
P)19
98A
Z st
ate
proh
ibit
ed
and
regu
late
d no
xiou
s w
eed.
N
ativ
e to
eas
tern
N
orth
Am
eric
a
Ranu
ncul
acea
e (H
elle
bora
ceae
)D
elph
iniu
m a
nder
soni
i va
r. sc
apos
umD
. sca
posu
mpa
le la
rksp
urPe
rFW
ide
His
t?
Peeb
les
1307
7 (A
RIZ)
1936
App
endi
x A
. Con
fi rm
ed, h
isto
rica
l, an
d re
port
ed v
ascu
lar
plan
t ta
xa o
f Pi
pe S
prin
g N
atio
nal M
onum
ent,
org
aniz
ed b
y fa
mily
and
sci
enti
fi c
nam
e w
ith
taxo
nom
ic
and
biog
eogr
aphi
c no
tes,
con
t.
Appendices 41
Fam
ilySp
ecie
s na
me
Syno
nym
s/Ta
xono
mic
not
esCo
mm
onna
me
Life
fo
rmRa
nge
Park
st
atus
Pop.
si
zeSo
urce
Year
do
c.Co
mm
ents
Rosa
ceae
Am
elan
chie
r ut
ahen
sis
Uta
h se
rvic
eber
rySh
rub
Wid
ePr
esU
ncN
elso
n 93
08
(RM
)19
71Ty
pe lo
calit
y is
Toq
uerv
ille,
W
ashi
ngto
n Co
., U
T (J
ones
171
6 G
H)
Rosa
ceae
Fallu
gia
para
doxa
Apa
che
plum
eSh
rub
Wid
ePr
esU
ncLo
uie
&
Even
den
PISP
38
(PIS
P)
1936
Rosa
ceae
Pote
ntill
a gr
acili
s va
r. el
mer
iP.
pec
tini
sect
aco
mb-
leaf
ci
nque
foil
PerF
Wid
eH
ist
?H
eato
n s.
n.
(PIS
P)19
39
Rosa
ceae
Prun
us a
mer
ican
aA
mer
ican
plu
mSh
rub
Intr
oPr
esU
ncFe
rtig
236
89
(PIS
P)19
98N
ativ
e to
cen
tral
N
orth
Am
eric
a
Rosa
ceae
Purs
hia
mex
ican
a va
r. st
ansb
urya
naP.
sta
nsbu
rian
a,
Cow
ania
mex
ican
acl
iff-
rose
Shru
bW
ide
Pres
Unc
Hea
ton
s.n.
(P
ISP)
1936
Rosa
ceae
Rosa
woo
dsii
R. a
rizo
nica
, R.
woo
dsii
var.
ultr
amon
tana
. In
clud
es R
. man
ca.
Woo
ds’ r
ose
Shru
bW
ide
Pres
Unc
Ale
xand
er
1239
(PIS
P)19
77Ci
ted
by K
ing
(197
7) a
s R
. ar
izon
ica
Salic
acea
ePo
pulu
s al
baw
hite
pop
lar
Tree
Intr
oPr
esU
ncA
lexa
nder
12
53 (P
ISP)
1997
Nat
ive
to E
uras
ia
Salic
acea
ePo
pulu
s fr
emon
tii v
ar.
frem
onti
iP.
fre
mon
tii,
P.
delt
oide
s va
r. fr
emon
tii
Frem
ont
cott
onw
ood
Tree
Wid
ePr
esU
ncA
lexa
nder
12
60 (P
ISP)
1998
Salic
acea
ePo
pulu
s ni
gra
Lom
bard
y po
plar
Tree
Intr
oRe
p?
Ale
xand
er
(199
8b)
1998
Nat
ive
to E
uras
ia
Salic
acea
ePo
pulu
s x
cana
dens
isH
ybri
d be
twee
n P.
de
ltoi
des
x P.
nig
raCa
rolin
a po
plar
Tree
Intr
oPr
esU
ncFe
rtig
236
87
(PIS
P)20
07
Salic
acea
ePo
pulu
s x
jack
iiH
ybri
d be
twee
n P.
bal
sam
ifer
a x
P.
delt
oide
s
balm
of
Gile
adTr
eeIn
tro
Pres
Unc
Nel
son
s.n.
(R
M)
1973
Salic
acea
eSa
lix e
xigu
a va
r. st
enop
hylla
S. e
xigu
a va
r. ex
igua
coyo
te w
illow
Shru
bW
ide
Pres
Unc
Ale
xand
er
1259
(PIS
P)19
98
Saxi
frag
acea
e (G
ross
ular
iace
ae)
Ribe
s au
reum
R.
aur
eum
var
. au
reum
gold
en c
urra
ntSh
rub
Wid
ePr
esU
ncFe
rtig
236
85
(PIS
P)20
07
Saxi
frag
acea
e (G
ross
ular
iace
ae)
Ribe
s sa
tivu
mR.
rub
rum
gard
en c
urra
ntSh
rub
Intr
oRe
p?
Ale
xand
er
(199
8b)
1998
Nat
ive
to E
urop
e
App
endi
x A
. Con
fi rm
ed, h
isto
rica
l, an
d re
port
ed v
ascu
lar
plan
t ta
xa o
f Pi
pe S
prin
g N
atio
nal M
onum
ent,
org
aniz
ed b
y fa
mily
and
sci
enti
fi c
nam
e w
ith
taxo
nom
ic
and
biog
eogr
aphi
c no
tes,
con
t.
42 Annotated Checklist of Vascular Flora: Pipe Spring National Monument
Fam
ilySp
ecie
s na
me
Syno
nym
s/Ta
xono
mic
not
esCo
mm
onna
me
Life
fo
rmRa
nge
Park
st
atus
Pop.
si
zeSo
urce
Year
do
c.Co
mm
ents
Scro
phul
aria
ceae
Cast
illej
a ch
rom
osa
C. a
pple
gate
i ss
p. m
arti
nii,
C.
angu
stif
olia
var
. du
bia
dese
rt
pain
tbru
shPe
rFW
ide
Pres
Unc
Even
den
&
Loui
e 29
(PIS
P)20
01
Scro
phul
aria
ceae
Cast
illej
a lin
ariif
olia
C. li
nari
aefo
liaW
yom
ing
pain
tbru
shPe
rFW
ide
Pres
Unc
Ale
xand
er
1195
(PIS
P)19
98
Scro
phul
aria
ceae
Cord
ylan
thus
pa
rvifl
orus
smal
lfl ow
er
bird
’s-be
akA
nnF
Wid
ePr
esU
ncA
lexa
nder
12
29 (P
ISP)
1935
Scro
phul
aria
ceae
Pens
tem
on la
evis
smoo
th
pens
tem
onPe
rFRe
gEn
Rep
?A
lexa
nder
(1
998b
)19
98Ty
pe lo
calit
y is
Spr
ingd
ale,
W
ashi
ngto
n Co
., U
T (J
ones
525
0 in
pa
rt U
S)
Scro
phul
aria
ceae
Pens
tem
on p
alm
eri
var.
egla
ndul
osus
Palm
er’s
pens
tem
onPe
rFRe
gEn
Pres
Unc
Fert
ig 2
3666
(P
ISP)
1998
Type
loca
lity
is
2.5
mile
s N
of
Kan
ab, K
ane
Co.,
UT
(Mag
uire
, M
agui
re, &
Pi
rani
an 1
2279
U
TC)
Scro
phul
aria
ceae
Pens
tem
on u
tahe
nsis
Uta
h pe
nste
mon
PerF
Wid
eH
ist
?H
eato
n 11
(P
ISP)
1937
Sim
arou
bace
aeA
ilant
hus
alti
ssim
atr
ee-o
f-he
aven
Tree
Intr
oPr
esU
ncFe
rtig
236
84
(PIS
P)19
35N
ativ
e to
Chi
na
Sola
nace
aeCh
amae
sara
cha
coro
nopu
sfa
lse
nigh
tsha
dePe
rFW
ide
Pres
Rare
Fert
ig 2
3695
(P
ISP)
2007
Sola
nace
aeD
atur
a w
righ
tii
Dat
ura
met
eloi
des
ange
l’s t
rum
pet
Ann
FW
ide
Pres
Unc
Hea
ton
(193
5)19
35
Sola
nace
aeLy
cium
pal
lidum
pale
wol
fber
rySh
rub
Wid
ePr
esU
ncFe
rtig
236
73
(PIS
P)19
77
Sola
nace
aeN
ico
tian
a tr
igon
ophy
llaN
. obt
usif
olia
; in
clud
es v
ars.
tr
igon
ophy
lla &
pa
lmer
i
dese
rt t
obac
coA
nnF
Wid
eH
ist
?H
eato
n s.
n.
(PIS
P)19
39
Sola
nace
aePh
ysal
is c
rass
ifol
iath
ick-
leav
ed
grou
nd-c
herr
yPe
rFW
ide
Rep
?A
lexa
nder
(1
998b
)19
98
Sola
nace
aePh
ysal
is h
eder
ifol
ia
var.
palm
eri
P. h
eder
aefo
lia v
ar.
palm
eri
Palm
er’s
grou
nd-c
herr
yPe
rFRe
gEn
Rep
?H
eato
n (1
935)
1935
App
endi
x A
. Con
fi rm
ed, h
isto
rica
l, an
d re
port
ed v
ascu
lar
plan
t ta
xa o
f Pi
pe S
prin
g N
atio
nal M
onum
ent,
org
aniz
ed b
y fa
mily
and
sci
enti
fi c
nam
e w
ith
taxo
nom
ic
and
biog
eogr
aphi
c no
tes,
con
t.
Appendices 43
Fam
ilySp
ecie
s na
me
Syno
nym
s/Ta
xono
mic
not
esCo
mm
onna
me
Life
fo
rmRa
nge
Park
st
atus
Pop.
si
zeSo
urce
Year
do
c.Co
mm
ents
Sola
nace
aeSo
lanu
m s
arra
choi
des
S. v
illos
um, S
. ph
ysal
ifol
ium
var
. ni
tidi
bacc
atum
grou
nd-c
herr
y ni
ghts
hade
Ann
FIn
tro
Rep
?A
lexa
nder
(1
998b
)N
AN
ativ
e to
Sou
th
Am
eric
a
Tam
aric
acea
eTa
mar
ix c
hine
nsis
T. p
enta
ndra
, T.
ram
osis
sim
afi
ve-s
tam
en
tam
aris
kSh
rub
Intr
oRe
p?
Kin
g (1
977)
, K
urth
(199
8)19
77N
ativ
e to
Eur
asia
Ulm
acea
eU
lmus
pum
ilaSi
beri
an e
lmTr
eeIn
tro
Pres
Unc
Nel
son
s.n.
(R
M)
1973
Nat
ive
to A
sia
Um
belli
fera
e (A
piac
eae)
Beru
la e
rect
a va
r. in
cisa
B. e
rect
acu
tlea
f w
ater
-pa
rsni
pPe
rFW
ide
Pres
Rare
Ale
xand
er
1245
(PIS
P)19
98
Um
belli
fera
e (A
piac
eae)
Cym
op
teru
s m
ulti
nerv
atus
purp
le-n
erve
d sp
ring
-par
sley
PerF
Wid
ePr
esU
ncLo
uie
&
Even
den
PISP
-2
(PIS
P)
1937
Ver
bena
ceae
Ver
bena
bra
ctea
tapr
ostr
ate
verv
ain
PerF
Wid
ePr
esU
ncA
lexa
nder
12
16 (P
ISP)
1997
Vis
cace
aePh
orad
endr
on
juni
peri
num
juni
per
mis
tlet
oePe
rFW
ide
Pres
Unc
Fert
ig 2
3677
(P
ISP)
1998
Typi
cally
par
asit
ic
on J
un
iper
us
oste
ospe
rma
Zygo
phyl
lace
aeLa
rrea
tri
dent
ata
creo
sote
bus
hSh
rub
Wid
ePr
esRa
reA
lexa
nder
12
37 (P
ISP)
2001
Zygo
phyl
lace
aeTr
ibul
us t
erre
stri
spu
nctu
revi
neA
nnF
Intr
oPr
esU
ncA
lexa
nder
12
33 (P
ISP)
1997
AZ
stat
e pr
ohib
ited
an
d re
gula
ted
noxi
ous
wee
d.
Nat
ive
to E
uras
ia.
App
endi
x A
. Con
fi rm
ed, h
isto
rica
l, an
d re
port
ed v
ascu
lar
plan
t ta
xa o
f Pi
pe S
prin
g N
atio
nal M
onum
ent,
org
aniz
ed b
y fa
mily
and
sci
enti
fi c
nam
e w
ith
taxo
nom
ic
and
biog
eogr
aphi
c no
tes,
con
t.
44 Annotated Checklist of Vascular Flora: Pipe Spring National Monument
App
endi
x B.
Pot
enti
al (u
ncon
fi rm
ed) t
axa
of P
ipe
Spri
ng N
atio
nal M
onum
ent.
Fam
ilySp
ecie
s na
me
Syno
nym
s/Ta
xono
mic
not
esCo
mm
on n
ame
Life
fo
rmRa
nge
Com
men
ts
Am
aran
thac
eae
Am
aran
thus
alb
ustu
mbl
e pi
gwee
dA
nnF
Intr
oN
ativ
e to
tro
pica
l Am
eric
a
Am
aran
thac
eae
Am
aran
thu
s re
trofl
exu
sre
droo
t pi
gwee
dA
nnF
Intr
oN
ativ
e to
Cen
tral
Am
eric
a
Asc
lepi
adac
eae
Asc
lepi
as s
peci
osa
show
y m
ilkw
eed
PerF
Wid
e
Bora
gina
ceae
Cryp
tant
ha h
umili
sO
reoc
arya
hum
ilis
dwar
f cr
ypta
nth
PerF
Peri
ph
Bora
gina
ceae
Cryp
tant
ha r
ecur
vata
recu
rved
cry
ptan
thA
nnF
Wid
e
Bora
gina
ceae
Cyno
glos
sum
offi
cin
ale
com
mon
hou
nd’s-
tong
uePe
rFIn
tro
Nat
ive
to E
uras
ia
Bora
gina
ceae
Lith
ospe
rmum
inci
sum
show
y st
ones
eed
PerF
Wid
e
Cact
acea
ePe
dioc
actu
s si
leri
Uta
hia
sile
riSi
ler’s
pin
cush
ion
cact
usPe
rFLo
cEn
Type
loca
lity
is P
ipe
Spri
ng,
Moh
ave
Co.,
AZ
(Sile
r s.
n.
MO
). U
SFW
S: T
hrea
tene
d
Capp
arac
eae
(Cle
omac
eae)
Cleo
me
serr
ulat
aRo
cky
Mou
ntai
n be
epla
ntA
bbF
Wid
e
Chen
opod
iace
aeA
trip
lex
conf
erti
folia
shad
scal
eSh
rub
Wid
e
Chen
opod
iace
aeBa
ssia
sco
pari
aK
ochi
a sc
opar
ia s
sp. s
cop
aria
, B.
sie
vers
iana
sum
mer
-cyp
ress
Ann
FIn
tro
Nat
ive
to E
uras
ia
Chen
opod
iace
aeC
hen
op
od
ium
de
sicc
atum
Som
e au
thor
s in
clud
e C
. pr
ater
icol
ade
sert
goo
sefo
otA
nnF
Peri
ph
Chen
opod
iace
aeCh
enop
odiu
m g
lauc
um
var.
salin
umC.
sal
inum
oakl
eaf
goos
efoo
tA
nnF
Wid
e
Chen
opod
iace
aeH
alog
eton
glo
mer
atus
halo
geto
nA
nnF
Intr
oN
ativ
e to
Eur
asia
Chen
opod
iace
aeK
rasc
heni
nnik
ovia
la
nata
var
. lan
ata
Cera
toid
es la
nata
, Eur
otia
la
nata
. Var
s. n
ot r
ecog
nize
d in
Flo
ra o
f N
orth
Am
eric
a (2
003)
.
win
terf
atSh
rub
Wid
e
Chen
opod
iace
aeSa
lsol
a pa
ulse
nii
barb
wir
e Ru
ssia
n th
istl
eA
nnF
Intr
oN
ativ
e to
Eur
asia
Com
posi
tae
(Ast
erac
eae)
Aca
mpt
opap
pus
spha
eroc
epha
lus
A. s
phae
roce
phal
us v
ar.
spha
eroc
epha
lus
gold
enhe
adSh
rub
Wid
e
Com
posi
tae
(Ast
erac
eae)
Arc
tium
min
usbu
rdoc
kPe
rFIn
tro
Nat
ive
to E
uras
ia
Com
posi
tae
(Ast
erac
eae)
Art
emis
ia s
pine
scen
sPi
crot
ham
nus
dese
rtor
umbu
dsag
eSh
rub
Peri
ph
Com
posi
tae
(Ast
erac
eae)
Chae
nact
is d
ougl
asii
Incl
udes
var
s. d
ougl
asii
&
mon
tana
hoar
y du
sty-
mai
den
PerF
Wid
e
Com
posi
tae
(Ast
erac
eae)
Chae
nact
is f
rem
onti
iFr
emon
t’s d
usty
-mai
den
Ann
FW
ide
Com
posi
tae
(Ast
erac
eae)
Chae
nact
is s
tevi
oide
sst
evia
dus
ty-m
aide
nA
nnF
Wid
e
Appendices 45
Fam
ilySp
ecie
s na
me
Syno
nym
s/Ta
xono
mic
not
esCo
mm
on n
ame
Life
fo
rmRa
nge
Com
men
ts
Com
posi
tae
(Ast
erac
eae)
Gly
ptop
leur
a se
tulo
sase
tose
cru
stw
eed
Ann
FW
ide
Type
loca
lity
is n
ear
St.
Geo
rge,
Was
hing
ton
Co.,
UT
(Pal
mer
6 G
H)
Com
posi
tae
(Ast
erac
eae)
Hel
iant
hus
peti
olar
is
ssp.
fal
lax
Incl
udes
var
. pet
iola
ris
prai
rie
sunfl
ow
erA
nnF
Wid
e
Com
posi
tae
(Ast
erac
eae)
Tara
xacu
m o
ffi c
inal
eco
mm
on d
ande
lion
PerF
Intr
oN
ativ
e to
Eur
asia
Cruc
ifer
ae (B
rass
icac
eae)
Chor
ispo
ra t
enel
labl
ue m
usta
rdA
nnF
Intr
oN
ativ
e to
Asi
a
Cruc
ifer
ae (B
rass
icac
eae)
Lepi
dium
vir
gini
cum
va
r. pu
besc
ens
poor
-man
’s pe
pper
wor
tA
nnF
Wid
e
Cruc
ifer
ae (B
rass
icac
eae)
Mal
colm
ia a
fric
ana
Afr
ican
mus
tard
Ann
FIn
tro
Nat
ive
to A
fric
a
Cruc
ifer
ae (B
rass
icac
eae)
Scho
enoc
ram
be
linif
olia
Sisy
mbr
ium
lini
foliu
mfl
ax-le
aved
pla
insm
usta
rdPe
rFW
ide
Gra
min
eae
(Poa
ceae
)Er
ione
uron
pilo
sum
hair
y tr
iden
sPe
rGW
ide
Gra
min
eae
(Poa
ceae
)Fe
stuc
a oc
tofl
ora
Vul
pia
octo
fl or
a; in
clud
es
vars
. gla
uca,
hir
tella
, &
octo
fl or
a
sixw
eeks
fes
cue
Ann
GW
ide
Gra
min
eae
(Poa
ceae
)Tr
itic
um a
esti
vum
whe
atA
nnG
Intr
oN
ativ
e to
Eur
asia
Legu
min
osae
(Fab
acea
e)A
stra
galu
s m
ollis
sim
us
var.
thom
pson
iae
woo
lly m
ilkve
tch
PerF
Wid
eTy
pe lo
calit
y is
Kan
ab,
Kan
e Co
., U
T (T
hom
pson
s.
n. G
H)
Legu
min
osae
(Fab
acea
e)Lu
pinu
s ar
gent
eus
var.
arge
nteu
ssi
lver
y lu
pine
PerF
Wid
e
Nyc
tagi
nace
aeTr
ipte
roca
lyx
mic
rant
hus
Abr
onia
mic
rant
ha, T
. pe
dunc
ulat
ussm
all-fl
ow
er s
andp
uffs
Ann
FW
ide
Ona
grac
eae
Oen
oth
era
coro
nopi
folia
hair
y-th
roat
eve
ning
-pr
imro
sePe
rFW
ide
Plan
tagi
nace
aePl
anta
go p
atag
onic
aP.
pat
agon
ica
var.
pata
goni
ca, P
. pur
shii;
in
clud
es v
ars.
gna
phal
oide
s,
spin
ulos
a, &
bre
visc
apa
woo
lly p
lant
ain
Ann
FW
ide
Pole
mon
iace
aeLi
nant
hus
dich
otom
usev
enin
g-sn
owA
nnF
Wid
e
Pole
mon
iace
aePh
lox
long
ifol
iaIn
clud
es v
ars.
long
ifol
ia &
st
ansb
uryi
long
-leaf
phl
oxPe
rFW
ide
Poly
gona
ceae
Erio
go
nu
m
cory
mbo
sum
var
. th
omps
onia
e
E. t
hom
pson
iae
var.
thom
pson
iae
Thom
pson
’s w
ild
buck
whe
atSh
rub
RegE
nTy
pe lo
calit
y is
nea
r K
anab
, K
ane
Co.,
UT
(Tho
mps
on
s.n.
GH
). U
SFW
S: 3
C
App
endi
x B.
Pot
enti
al (u
ncon
fi rm
ed) t
axa
of P
ipe
Spri
ng N
atio
nal M
onum
ent,
con
t.
46 Annotated Checklist of Vascular Flora: Pipe Spring National Monument
Fam
ilySp
ecie
s na
me
Syno
nym
s/Ta
xono
mic
not
esCo
mm
on n
ame
Life
fo
rmRa
nge
Com
men
ts
Poly
gona
ceae
Poly
go
nu
m
lapa
thif
oliu
mPe
rsic
aria
lapa
thif
olia
will
ow-w
eed
Ann
FIn
tro
Nat
ive
to E
uras
ia
Ranu
ncul
acea
eRa
nunc
ulus
inam
oenu
s R.
inam
oenu
s va
r. in
amoe
nus.
Incl
udes
var
. al
peop
hilu
s.
drab
but
terc
upPe
rFW
ide
Ranu
ncul
acea
eRa
nunc
ulus
tes
ticu
latu
sCe
rato
ceph
ala
test
icul
ata,
C.
orth
ocer
asbu
r bu
tter
cup
Ann
FIn
tro
Nat
ive
to E
uras
ia
Sant
alac
eae
Com
andr
a um
bella
ta
var.
palli
daC.
pal
lida
bast
ard
toad
fl ax
PerF
Wid
e
Scro
phul
aria
ceae
Cast
illej
a sc
abri
da v
ar.
scab
rida
East
woo
d’s
pain
tbru
shPe
rFRe
gEn
Sola
nace
aeLy
cium
and
erso
nii
And
erso
n’s
wol
fber
rySh
rub
Wid
e
App
endi
x B.
Pot
enti
al (u
ncon
fi rm
ed) t
axa
of P
ipe
Spri
ng N
atio
nal M
onum
ent,
con
t.
Appendices 47
App
endi
x C.
Rej
ecte
d (f
alse
ly r
epor
ted
or q
uest
iona
ble)
tax
a of
Pip
e Sp
ring
Nat
iona
l Mon
umen
t.
Fam
ilySp
ecie
s na
me
Syno
nym
s/Ta
xono
mic
not
esCo
mm
on n
ame
Life
fo
rmRa
nge
Park
st
atus
Com
men
ts
Aga
vace
ae
(Lili
acea
e)Yu
cca
angu
stis
sim
aY.
ang
usti
ssim
a va
r. an
gust
issi
ma
narr
owle
aved
yuc
caSh
rub
RegE
nQ
ues?
Repo
rted
by
Kin
g (1
977)
, but
pro
babl
y ba
sed
on Y
. ka
nabe
nsis
.
Bora
gina
ceae
Cry
pta
nth
a ba
rbig
era
bear
ded
cryp
tant
haA
nnF
Wid
eFa
lsRe
pRe
port
fro
m P
ISP
herb
ariu
m b
ased
on
a m
isid
enti
fi ed
spe
cim
en o
f Pl
agio
both
rys
jone
sii.
Type
lo
calit
y is
St.
Geo
rge,
Was
hing
ton
Co.,
UT
(Par
ry 1
71
GH
).
Cact
acea
eO
punt
ia b
asila
ris
O. b
asila
ris
var.
basi
lari
sbe
aver
tail
cact
usPe
rFW
ide
Que
s?Re
port
ed b
y K
ing
(197
7), b
ut p
roba
bly
base
d on
O.
erin
acea
var
. aur
ea.
Chen
opod
iace
aeA
trip
lex
gard
neri
va
r. cu
neat
aA
. cun
eata
valle
y sa
ltbu
shSh
rub
RegE
nQ
ues?
Repo
rted
by
Hea
ton
(193
5), b
ut p
roba
bly
base
d on
A
. can
esce
ns.
Com
posi
tae
(Ast
erac
eae)
Mad
ia g
lom
erat
am
ount
ain
tarw
eeed
Ann
FPe
riph
Fals
Rep
Repo
rt f
rom
Ale
xand
er (1
998a
) bas
ed o
n a
mis
iden
tifi
ed s
peci
men
of
Cord
ylan
thus
par
vifl
orus
.
Com
posi
tae
(Ast
erac
eae)
Sonc
hus
arve
nsis
spin
y-le
af s
owth
istl
eA
nnF
Intr
oFa
lsRe
pRe
port
fro
m P
ISP
herb
ariu
m b
ased
on
a m
isid
enti
fi ed
spe
cim
en o
f S.
asp
er. N
ativ
e to
Eur
ope.
A
Z st
ate
proh
ibit
ed n
oxio
us w
eed.
Faga
ceae
Que
rcus
dum
osa
coas
tal s
age
scru
b oa
kSh
rub
RegE
nQ
ues?
Repo
rted
by
Hea
ton
(193
5), b
ut p
roba
bly
base
d on
Q
. tur
bine
lla. Q
. dum
osa
occu
rs in
Cal
ifor
nia
but
is
not
know
n fr
om A
rizo
na.
Hyd
roph
ylla
ceae
Phac
elia
am
bigu
aP.
cre
nula
ta v
ar.
ambi
gua
Jone
s’ p
hace
liaA
nnF
Wid
eFa
lsRe
pRe
port
fro
m P
ISP
herb
ariu
m b
ased
on
a m
isid
enti
fi ed
spe
cim
en o
f P.
cre
nula
ta v
ar. c
renu
lata
.
Loas
acea
eM
entz
elia
nit
ens
M. n
iten
s va
r. jo
nesi
icu
rvep
od s
tick
leaf
Ann
FW
ide
Fals
Rep
Repo
rt f
rom
PIS
P he
rbar
ium
bas
ed o
n a
mis
iden
tifi
ed s
peci
men
of
M. a
lbic
aulis
.
Loas
acea
eM
entz
elia
pu
mila
var
. pu
mila
M. p
umila
Wyo
min
g st
ickl
eaf
PerF
RegE
nFa
lsRe
pRe
port
fro
m A
lexa
nder
(199
8a) b
ased
on
a m
isid
enti
fi ed
spe
cim
en o
f M
. mul
tifl
ora.
Not
kno
wn
from
Ari
zona
Mal
vace
aeSp
haer
alce
a la
xaca
liche
glo
be m
allo
wPe
rFW
ide
Que
s?Re
port
ed b
y K
ing
(197
7), b
ut p
roba
bly
base
d on
S.
ambi
gua.
Ona
grac
eae
Oen
oth
era
cesp
itos
a va
r. m
argi
nata
long
-tub
e ev
enin
g-pr
imro
sePe
rFW
ide
Fals
Rep
Repo
rt f
rom
PIS
P he
rbar
ium
bas
ed o
n a
mis
iden
tifi
ed s
peci
men
of
O. c
espi
tosa
var
. na
vajo
ensi
s.
Pole
mon
iace
aeG
ilia
scop
ulor
umro
ck g
ilia
Ann
FW
ide
Fals
Rep
Repo
rt f
rom
PIS
P he
rbar
ium
bas
ed o
n a
mis
iden
tifi
ed s
peci
men
of
G. i
ncon
spic
ua.T
ype
loca
lity
is S
t. G
eorg
e, W
ashi
ngto
n Co
., U
T (J
ones
16
59 P
OM
).
Scro
phul
aria
ceae
Cord
ylan
thus
ne
vini
iN
evin
’s bi
rd’s-
beak
Ann
FRe
gEn
Que
s?Re
port
ed b
y H
eato
n (1
935)
, but
pro
babl
y ba
sed
on
C. p
arvi
fl or
us.
Um
belli
fera
e (A
piac
eae)
Cym
op
teru
s pu
rpur
eus
C. p
urpu
reus
var
. pu
rpur
eus
Colo
rado
Pla
teau
sp
ring
-par
sley
PerF
RegE
nFa
lsRe
pRe
port
fro
m P
ISP
herb
ariu
m b
ased
on
a m
isid
enti
fi ed
spe
cim
en o
f C.
mul
tine
rvat
us.
48 Annotated Checklist of Vascular Flora: Pipe Spring National Monument
App
endi
x D
. Con
fi rm
ed, h
isto
rica
l, an
d re
port
ed t
axa
of P
ipe
Spri
ng N
atio
nal M
onum
ent
orga
nize
d by
life
for
m a
nd c
omm
on n
ame
wit
h ec
olog
ical
not
es.
Fam
ilySp
ecie
s na
me
Com
mon
nam
eN
ativ
ity
Park
st
atus
Flow
er
tim
eD
esSh
Pin
Jun
Wet
Com
men
ts
Tree
s
Cupr
essa
ceae
Juni
peru
s sc
opul
orum
Rock
y M
ount
ain
juni
per
Nat
ive
Rep
NA
xx
Legu
min
osae
(Fab
acea
e)Ro
bini
a ps
eudo
acac
iabl
ack
locu
stIn
tro
Pres
May
–Jun
Cult
ivat
ed
Pina
ceae
Pinu
s ed
ulis
two-
need
le p
inyo
n N
ativ
ePr
esN
Ax
Salic
acea
ePo
pulu
s al
baw
hite
pop
lar
Intr
oPr
esM
ar–J
unCu
ltiv
ated
Salic
acea
ePo
pulu
s fr
emon
tii v
ar.
frem
onti
iFr
emon
t co
tton
woo
dN
ativ
ePr
esM
ar–J
unx
Salic
acea
ePo
pulu
s ni
gra
Lom
bard
y po
plar
Intr
oRe
pA
pr–M
ayCu
ltiv
ated
Salic
acea
ePo
pulu
s x
cana
dens
isCa
rolin
a po
plar
Intr
oPr
esA
pr–M
ayCu
ltiv
ated
Salic
acea
ePo
pulu
s x
jack
iiba
lm o
f G
ilead
Intr
oPr
esA
pr–M
ayCu
ltiv
ated
Sim
arou
bace
aeA
ilant
hus
alti
ssim
atr
ee-o
f-he
aven
Intr
oPr
esJu
n–Ju
lD
istu
rbed
are
as
Ulm
acea
eU
lmus
pum
ilaSi
beri
an e
lmIn
tro
Pres
Mar
–Apr
xD
istu
rbed
are
as
Shru
bs
Aga
vace
ae (L
iliac
eae)
Yucc
a ba
ccat
ada
til y
ucca
Nat
ive
Pres
Apr
–Jun
xx
Aga
vace
ae (L
iliac
eae)
Yucc
a ka
nabe
nsis
Kan
ab y
ucca
Nat
ive
Pres
May
–Jul
xx
Ana
card
iace
aeRh
us a
rom
atic
a va
r. si
mpl
icif
olia
squa
wbu
shN
ativ
ePr
esM
ay–J
ulx
Ana
card
iace
aeRh
us a
rom
atic
a va
r. tr
iloba
tasq
uaw
bush
Nat
ive
Rep
May
–Jul
x
Cact
acea
eO
punt
ia w
hipp
lei v
ar.
whi
pple
iW
hipp
le’s
chol
laN
ativ
ePr
esM
ay–J
ulx
Chen
opod
iace
aeA
trip
lex
cane
scen
s va
r. ca
nesc
ens
four
win
g sa
ltbu
shN
ativ
ePr
esM
ay–S
epx
x
Chen
opod
iace
aeK
rasc
heni
nnik
ovia
lana
ta v
ar.
subs
pino
sash
rubb
y w
inte
rfat
Nat
ive
His
tM
ay–J
ulx
Chen
opod
iace
ae
(Sar
coba
tace
ae)
Sarc
obat
us v
erm
icul
atus
grea
sew
ood
Nat
ive
Pres
May
–Aug
x
Com
posi
tae
(Ast
erac
eae)
Art
emis
ia b
igel
ovii
Bige
low
’s sa
gebr
ush
Nat
ive
Pres
Aug
–Oct
xx
Com
posi
tae
(Ast
erac
eae)
Art
emis
ia fi
lifol
iasa
nd s
ageb
rush
N
ativ
ePr
esA
ug–O
ctx
Com
posi
tae
(Ast
erac
eae)
Art
emis
ia t
ride
ntat
a va
r. tr
iden
tata
basi
n bi
g sa
gebr
ush
Nat
ive
Pres
Jul–
Oct
x
Com
posi
tae
(Ast
erac
eae)
Chry
soth
amnu
s na
useo
sus
var.
cons
imili
sgr
eeni
sh r
abbi
tbru
shN
ativ
ePr
esA
ug–O
ctx
Appendices 49
App
endi
x D
. Con
fi rm
ed, h
isto
rica
l, an
d re
port
ed t
axa
of P
ipe
Spri
ng N
atio
nal M
onum
ent
orga
nize
d by
life
for
m a
nd c
omm
on n
ame
wit
h ec
olog
ical
not
es, c
ont.
Fam
ilySp
ecie
s na
me
Com
mon
nam
eN
ativ
ity
Park
st
atus
Flow
er
tim
eD
esSh
Pin
Jun
Wet
Com
men
ts
Shru
bs, c
ont.
Com
posi
tae
(Ast
erac
eae)
Chry
soth
amnu
s na
useo
sus
var.
gnap
halo
des
gray
stem
rab
bitb
rush
Nat
ive
Pres
Aug
–Oct
x
Com
posi
tae
(Ast
erac
eae)
Chry
soth
amnu
s na
useo
sus
var.
grav
eole
nsgl
abra
te r
abbi
tbru
shN
ativ
eRe
pA
ug–O
ctx
xx
Com
posi
tae
(Ast
erac
eae)
Chry
soth
amnu
s na
useo
sus
var.
junc
eus
rush
rab
bitb
rush
Nat
ive
Pres
Aug
–Oct
x
Com
posi
tae
(Ast
erac
eae)
Chry
soth
amnu
s vi
scid
ifl o
rus
var.
pube
rulu
spu
beru
lent
rab
bitb
rush
Nat
ive
Pres
Jul–
Sep
x
Com
posi
tae
(Ast
erac
eae)
Chry
soth
amnu
s vi
scid
ifl o
rus
var.
sten
ophy
llus
slen
derl
eaf
rabb
itbr
ush
Nat
ive
Pres
Jul–
Sep
xx
Com
posi
tae
(Ast
erac
eae)
Chry
soth
amnu
s vi
scid
ifl o
rus
var.
visc
idifl
orus
gree
n ra
bbit
brus
hN
ativ
eRe
pJu
l–Se
px
Com
posi
tae
(Ast
erac
eae)
Gut
ierr
ezia
mic
roce
phal
ath
read
sna
kew
eed
Nat
ive
Pres
Jul–
Oct
xx
Com
posi
tae
(Ast
erac
eae)
Gut
ierr
ezia
sar
othr
aebr
oom
sna
kew
eed
Nat
ive
Pres
Jul–
Oct
xx
Cupr
essa
ceae
Juni
peru
s os
teos
perm
aU
tah
juni
per
Nat
ive
Pres
NA
xx
Elae
agna
ceae
Shep
herd
ia r
otun
difo
liaro
undl
eaf
buff
alob
erry
Nat
ive
Pres
Mar
–May
xx
Ephe
drac
eae
Ephe
dra
torr
eyan
aTo
rrey
’s ep
hedr
aN
ativ
ePr
esN
Ax
Ephe
drac
eae
Ephe
dra
viri
dis
var.
viri
dis
gree
n ep
hedr
aN
ativ
ePr
esN
Ax
Ephe
drac
eae
Ephe
dra
viri
dis
var.
visc
ida
Cutl
er’s
ephe
dra
Nat
ive
Pres
NA
x
Faga
ceae
Que
rcus
tur
bine
llatu
rbin
ella
live
-oak
Nat
ive
Pres
x
Labi
atae
(Lam
iace
ae)
Salv
ia d
orri
i ssp
. dor
rii
Dor
r’s s
age
Nat
ive
Pres
May
–Jun
x
Poly
gona
ceae
Erio
gonu
m c
orym
bosu
m v
ar.
aure
umgo
lden
wild
buc
kwhe
atN
ativ
eRe
pA
ug–O
ctx
Poly
gona
ceae
Erio
gonu
m c
orym
bosu
m v
ar.
cory
mbo
sum
cris
plea
f w
ild b
uckw
heat
Nat
ive
Pres
Jul–
Oct
Rosa
ceae
Am
elan
chie
r ut
ahen
sis
Uta
h se
rvic
eber
ryN
ativ
ePr
esM
ay–J
unx
Rosa
ceae
Fallu
gia
para
doxa
Apa
che
plum
eN
ativ
ePr
esA
pr–A
ugx
Rosa
ceae
Prun
us a
mer
ican
aA
mer
ican
plu
mIn
tro
Pres
Cult
ivat
ed
Rosa
ceae
Purs
hia
mex
ican
a va
r. st
ansb
urya
nacl
iff-
rose
Nat
ive
Pres
May
–Jun
xx
Rosa
ceae
Rosa
woo
dsii
Woo
ds’ r
ose
Nat
ive
Pres
May
–Jul
xx
Salic
acea
eSa
lix e
xigu
a va
r. st
enop
hylla
coyo
te w
illow
Nat
ive
Pres
Apr
–Aug
x
50 Annotated Checklist of Vascular Flora: Pipe Spring National Monument
App
endi
x D
. Con
fi rm
ed, h
isto
rica
l, an
d re
port
ed t
axa
of P
ipe
Spri
ng N
atio
nal M
onum
ent
orga
nize
d by
life
for
m a
nd c
omm
on n
ame
wit
h ec
olog
ical
not
es, c
ont.
Fam
ilySp
ecie
s na
me
Com
mon
nam
eN
ativ
ity
Park
st
atus
Flow
er
tim
eD
esSh
Pin
Jun
Wet
Com
men
ts
Shru
bs, c
ont.
Saxi
frag
acea
e (G
ross
ular
iace
ae)
Ribe
s au
reum
go
lden
cur
rant
Nat
ive
Pres
Mar
–Jul
Dis
turb
ed a
reas
Saxi
frag
acea
e (G
ross
ular
iace
ae)
Ribe
s sa
tivu
mga
rden
cur
rant
Intr
oRe
pM
ayCu
ltiv
ated
Sola
nace
aeLy
cium
pal
lidum
pale
wol
fber
ryN
ativ
ePr
esM
ar–M
ayx
x
Tam
aric
acea
eTa
mar
ix c
hine
nsis
fi ve
-sta
men
tam
aris
kIn
tro
Rep
Apr
–Sep
x
Zygo
phyl
lace
aeLa
rrea
tri
dent
ata
creo
sote
bus
hN
ativ
ePr
esA
pr–J
unx
Pere
nnia
l For
bs
Apo
cyna
ceae
Am
soni
a jo
nesi
iJo
nes’
am
soni
aN
ativ
eH
ist
Apr
–Jun
xx
Apo
cyna
ceae
Am
soni
a to
men
tosa
var
. st
enop
hylla
tom
ento
se a
mso
nia
Nat
ive
His
tA
pr–M
ayx
Apo
cyna
ceae
Apo
cynu
m c
anna
binu
mco
mm
on d
ogba
neN
ativ
eRe
pJu
n–A
ugx
Asc
lepi
adac
eae
Asc
lepi
as la
tifo
liabr
oadl
eaf
milk
wee
dN
ativ
eH
ist
Jun–
Aug
xx
Asc
lepi
adac
eae
Asc
lepi
as s
ubve
rtic
illat
aw
horl
ed m
ilkw
eed
Nat
ive
Pres
May
–Sep
xD
istu
rbed
are
as
Bora
gina
ceae
Cryp
tant
ha c
onfe
rtifl
ora
basi
n ye
llow
cry
ptan
thN
ativ
ePr
esA
pr–J
unx
x
Bora
gina
ceae
Cryp
tant
ha fl
ava
plat
eau
yello
w c
rypt
anth
Nat
ive
Rep
Apr
–Jun
x
Bora
gina
ceae
Cryp
tant
ha f
ulvo
cane
scen
spl
atea
u cr
ypta
nth
Nat
ive
His
tM
ay–J
unx
Cact
acea
eEc
hino
cere
us e
ngel
man
nii v
ar.
vari
egat
usG
len
Cany
on h
edge
hog
cact
usN
ativ
eRe
pM
ar–J
ulx
Cact
acea
eEc
hino
cere
us t
rigl
ochi
diat
us
var.
mel
anac
anth
uscl
aret
cup
Nat
ive
Pres
Apr
–Jun
x
Cact
acea
eEc
hino
cere
us t
rigl
ochi
diat
us
var.
moj
aven
sis
Moh
ave
clar
etcu
pN
ativ
eH
ist
Apr
–Jun
xx
Cact
acea
eO
punt
ia e
rina
cea
var.
aure
aPi
pe S
prin
g pr
ickl
ypea
rN
ativ
ePr
esM
ay–J
unx
x
Cact
acea
eO
punt
ia e
rina
cea
var.
utah
ensi
sU
tah
pric
klyp
ear
Nat
ive
Pres
May
–Jun
xx
Cact
acea
eO
punt
ia f
ragi
lisbr
ittl
e pr
ickl
y-pe
arN
ativ
eRe
pJu
n–Ju
lx
x
Cact
acea
eO
punt
ia p
haea
cant
ha v
ar.
maj
orla
rge
pric
klyp
ear
Nat
ive
Pres
Apr
–Jul
xx
Cact
acea
eO
punt
ia p
inka
vae
Pink
ava’
s pr
ickl
ypea
rN
ativ
ePr
esM
ay–J
unx
Cact
acea
eSc
lero
cact
us w
hipp
lei v
ar.
rose
ussm
all-fl
ow
er fi
shho
ok
cact
usN
ativ
ePr
esA
pr–M
ayx
x
Appendices 51
App
endi
x D
. Con
fi rm
ed, h
isto
rica
l, an
d re
port
ed t
axa
of P
ipe
Spri
ng N
atio
nal M
onum
ent
orga
nize
d by
life
for
m a
nd c
omm
on n
ame
wit
h ec
olog
ical
not
es, c
ont.
Fam
ilySp
ecie
s na
me
Com
mon
nam
eN
ativ
ity
Park
st
atus
Flow
er
tim
eD
esSh
Pin
Jun
Wet
Com
men
ts
Pere
nnia
l For
bs, c
ont.
Com
posi
tae
(Ast
erac
eae)
Art
emis
ia c
ampe
stri
s va
r. sc
oule
rian
asa
nd w
orm
woo
dN
ativ
eRe
pJu
l–Se
px
Com
posi
tae
(Ast
erac
eae)
Art
emis
ia d
racu
ncul
us s
sp.
glau
cata
rrag
onN
ativ
eRe
pJu
l–O
ctx
Com
posi
tae
(Ast
erac
eae)
Art
emis
ia lu
dovi
cian
a va
r. al
bula
Loui
sian
a w
orm
woo
dN
ativ
ePr
esJu
l–O
ctx
x
Com
posi
tae
(Ast
erac
eae)
Ast
er h
espe
rius
Sisk
iyou
ast
erN
ativ
ePr
esJu
l–Se
px
Com
posi
tae
(Ast
erac
eae)
Chae
topa
ppa
eric
oide
sro
se-h
eath
Nat
ive
Pres
May
–Sep
xx
Com
posi
tae
(Ast
erac
eae)
Erig
eron
div
erge
ns v
ar.
dive
rgen
ssp
read
ing
dais
yN
ativ
ePr
esM
ay–S
epx
Com
posi
tae
(Ast
erac
eae)
Erig
eron
pum
ilus
var.
conc
innu
sN
avaj
o fl
eaba
neN
ativ
eRe
pM
ay–A
ugx
Com
posi
tae
(Ast
erac
eae)
Gai
llard
ia p
inna
tifi
daH
opi b
lank
etfl
ower
Nat
ive
Pres
Apr
–Aug
xx
Com
posi
tae
(Ast
erac
eae)
Gri
ndel
ia s
quar
rosa
var
. se
rrul
ata
curl
ycup
gum
wee
dN
ativ
eRe
pJu
l–Se
pD
istu
rbed
are
as
Com
posi
tae
(Ast
erac
eae)
Hym
enop
appu
s fi
lifol
ius
var.
cine
reus
com
mon
hya
lineh
erb
Nat
ive
Pres
May
–Oct
x
Com
posi
tae
(Ast
erac
eae)
Iva
axill
aris
po
vert
ywee
dN
ativ
ePr
esM
ay–S
epx
Com
posi
tae
(Ast
erac
eae)
Mac
haer
anth
era
cane
scen
s va
r. ca
nesc
ens
hoar
y as
ter
Nat
ive
Pres
May
–Sep
xx
x
Com
posi
tae
(Ast
erac
eae)
Ono
pord
um a
cant
hium
Scot
ch t
hist
leIn
tro
Rep
Jun–
Sep
x
Com
posi
tae
(Ast
erac
eae)
Psilo
stro
phe
spar
sifl
ora
gree
nste
m p
aper
fl ow
erN
ativ
eH
ist
May
–Sep
Com
posi
tae
(Ast
erac
eae)
Sene
cio
doug
lasi
i var
. lo
ngilo
bus
Dou
glas
’ gro
unds
elN
ativ
ePr
esM
ay–O
ctx
Com
posi
tae
(Ast
erac
eae)
Solid
ago
sp.
gold
enro
dN
ativ
eRe
pJu
n–O
ctx
Com
posi
tae
(Ast
erac
eae)
Step
hano
mer
ia p
auci
fl or
abr
own-
plum
e w
ire
lett
uce
Nat
ive
Pres
May
–Oct
xx
Com
posi
tae
(Ast
erac
eae)
Thel
espe
rma
subn
udum
var
. su
bnud
umsc
apos
e gr
eent
hrea
dN
ativ
ePr
esM
ay–A
ugx
x
Com
posi
tae
(Ast
erac
eae)
Tow
nsen
dia
inca
naho
ary
tow
nsen
dia
Nat
ive
Pres
Apr
–Jul
xx
Com
posi
tae
(Ast
erac
eae)
Trag
opog
on d
ubiu
sye
llow
sal
sify
Intr
oPr
esM
ay–J
ulx
Conv
olvu
lace
aeCo
nvol
vulu
s ar
vens
isfi
eld
bind
wee
dIn
tro
His
tJu
n–A
ugD
istu
rbed
are
as
Cruc
ifer
ae (B
rass
icac
eae)
Ara
bis
pere
nnan
s va
r. pe
renn
ans
pere
nnia
l roc
kcre
ssN
ativ
eRe
pM
ar–J
ulx
52 Annotated Checklist of Vascular Flora: Pipe Spring National Monument
App
endi
x D
. Con
fi rm
ed, h
isto
rica
l, an
d re
port
ed t
axa
of P
ipe
Spri
ng N
atio
nal M
onum
ent
orga
nize
d by
life
for
m a
nd c
omm
on n
ame
wit
h ec
olog
ical
not
es, c
ont.
Fam
ilySp
ecie
s na
me
Com
mon
nam
eN
ativ
ity
Park
st
atus
Flow
er
tim
eD
esSh
Pin
Jun
Wet
Com
men
ts
Pere
nnia
l For
bs, c
ont.
Cruc
ifer
ae (B
rass
icac
eae)
Lepi
dium
mon
tanu
m v
ar.
jone
sii
Jone
s’ p
eppe
rwor
tN
ativ
ePr
esA
pr–S
epx
x
Cruc
ifer
ae (B
rass
icac
eae)
Lepi
dium
mon
tanu
m v
ar.
stel
lae
Stel
la’s
pepp
erw
ort
Nat
ive
His
tM
ay–J
unx
Cruc
ifer
ae (B
rass
icac
eae)
Nas
turt
ium
offi
cin
ale
wat
ercr
ess
Intr
oPr
esM
ay–S
epx
Cruc
ifer
ae (B
rass
icac
eae)
Stan
leya
pin
nata
prin
ce’s-
plum
eN
ativ
ePr
esA
pr–S
epx
x
Cucu
rbit
acea
eCu
curb
ita
foet
idis
sim
abu
ffal
o go
urd
Nat
ive
Pres
May
–Aug
xx
Dis
turb
ed a
reas
Euph
orbi
acea
eCh
amae
syce
alb
omar
gina
tara
ttle
snak
e-w
eed
Nat
ive
Rep
Apr
–Sep
x
Euph
orbi
acea
eCh
amae
syce
fen
dler
iFe
ndle
r’s s
purg
eN
ativ
ePr
esM
ay–S
epx
x
Fum
aria
ceae
Cory
dalis
aur
ea v
ar.
occi
dent
alis
gold
en c
oryd
alis
Nat
ive
His
tx
x
Gen
tian
acea
eSw
erti
a al
bom
argi
nata
whi
te-m
argi
ned
swer
tia
Nat
ive
His
tM
ay–J
ulx
x
Gen
tian
acea
eSw
erti
a ut
ahen
sis
Uta
h sw
erti
aN
ativ
eRe
pJu
n–Ju
lx
x
Irid
acea
eSi
syri
nchi
um d
emis
sum
blue
-eye
d gr
ass
Nat
ive
Rep
Apr
–Aug
x
Labi
atae
(Lam
iace
ae)
Mar
rubi
um v
ulga
reco
mm
on h
oreh
ound
Intr
oPr
esM
ay–O
ctx
xD
istu
rbed
are
as
Labi
atae
(Lam
iace
ae)
Men
tha
spic
ata
spea
rmin
tIn
tro
Pres
Jun–
Aug
x
Legu
min
osae
(Fab
acea
e)A
stra
galu
s am
phio
xys
var.
vesp
erti
nus
Shel
don’
s m
ilkve
tch
Nat
ive
His
tM
ar–J
unx
x
Legu
min
osae
(Fab
acea
e)A
stra
galu
s ce
ram
icus
var
. ce
ram
icus
pain
ted
milk
vetc
hN
ativ
ePr
esA
pr–J
unx
Legu
min
osae
(Fab
acea
e)A
stra
galu
s pr
aelo
ngus
var
. pr
aelo
ngus
stin
king
milk
vetc
hN
ativ
eRe
pA
pr–J
ulx
x
Legu
min
osae
(Fab
acea
e)G
lycy
rrhi
za le
pido
ta
Nut
tall’
s lic
oric
eN
ativ
eRe
pM
ay–J
ulx
Legu
min
osae
(Fab
acea
e)M
edic
ago
sati
vaal
falf
aIn
tro
Rep
May
–Sep
Dis
turb
ed a
reas
Legu
min
osae
(Fab
acea
e)M
elilo
tus
offi
cina
lisye
llow
sw
eetc
love
rIn
tro
Rep
May
–Oct
xD
istu
rbed
are
as
Legu
min
osae
(Fab
acea
e)Pe
teri
a th
omps
onia
eTh
omps
on’s
pete
ria
Nat
ive
Pres
Apr
–Jun
x
Lilia
ceae
(The
mid
acea
e)A
ndro
step
hium
bre
vifl
orum
pink
fun
nel-l
ilyN
ativ
eH
ist
Apr
–May
Lilia
ceae
(C
aloc
hort
acea
e)Ca
loch
ortu
s fl
exuo
sus
sinu
ous
mar
ipos
aN
ativ
ePr
esA
pr–J
unx
Loas
acea
eM
entz
elia
inte
gra
Vir
gin
stic
klea
fN
ativ
eH
ist
Apr
–Oct
x
Loas
acea
eM
entz
elia
mul
tifl
ora
dese
rt s
tick
leaf
Nat
ive
Pres
Apr
–Sep
x
Mal
vace
aeA
ltha
ea r
osea
holly
hock
Intr
oRe
pJu
n–O
ctCu
ltiv
ated
Appendices 53
App
endi
x D
. Con
fi rm
ed, h
isto
rica
l, an
d re
port
ed t
axa
of P
ipe
Spri
ng N
atio
nal M
onum
ent
orga
nize
d by
life
for
m a
nd c
omm
on n
ame
wit
h ec
olog
ical
not
es, c
ont.
Fam
ilySp
ecie
s na
me
Com
mon
nam
eN
ativ
ity
Park
st
atus
Flow
er
tim
eD
esSh
Pin
Jun
Wet
Com
men
ts
Pere
nnia
l For
bs, c
ont.
Mal
vace
aeSp
haer
alce
a am
bigu
a va
r. am
bigu
aM
ohav
e gl
obem
allo
wN
ativ
ePr
esM
ar–S
epx
x
Mal
vace
aeSp
haer
alce
a pa
rvif
olia
smal
l-lea
f gl
obem
allo
wN
ativ
eH
ist
Mar
–Oct
xx
Nyc
tagi
nace
aeA
bron
ia f
ragr
ans
frag
rant
san
d-ve
rben
aN
ativ
ePr
esA
pr–S
ep
Nyc
tagi
nace
aeM
irab
ilis
mul
tifl
ora
sh
owy
four
-o’c
lock
Nat
ive
Pres
Apr
–Jul
x
Ole
acea
eM
enod
ora
scab
raro
ugh
men
odor
aN
ativ
ePr
esA
pr–A
ugx
Ona
grac
eae
Epilo
bium
cili
atum
nort
hern
will
ow-h
erb
Nat
ive
Rep
Jun–
Aug
x
Ona
grac
eae
Gau
ra c
occi
nea
scar
let
gaur
aN
ativ
eH
ist
May
–Jul
xx
Ona
grac
eae
Oen
othe
ra c
aesp
itos
a va
r. na
vajo
ensi
sPa
ria
even
ing
prim
rose
Nat
ive
Pres
Apr
–Jun
x
Ona
grac
eae
Oen
othe
ra c
alif
orni
caCa
lifor
nia
even
ing
prim
rose
Nat
ive
Pres
May
–Jul
x
Ona
grac
eae
Oen
othe
ra p
allid
a va
r. pa
llida
pale
eve
ning
pri
mro
seN
ativ
ePr
esM
ay–J
ulx
Oro
banc
hace
aeO
roba
nche
ludo
vici
ana
Coop
er’s
broo
mra
peN
ativ
ePr
esM
ay–J
ulx
Plan
tagi
nace
aePl
anta
go la
nceo
lata
Engl
ish
plan
tain
Intr
oRe
pM
ay–A
ugD
istu
rbed
are
as
Plan
tagi
nace
aePl
anta
go m
ajor
com
mon
pla
ntai
nIn
tro
Pres
May
–Aug
xD
istu
rbed
are
as
Pole
mon
iace
aeIp
omop
sis
long
ifl or
alo
ng-fl
ow
ered
gili
aN
ativ
ePr
esM
ay–A
ugx
Poly
gala
ceae
Poly
gala
sub
spin
osa
cush
ion
milk
wor
tN
ativ
eH
ist
May
–Jun
x
Poly
gona
ceae
Erio
gonu
m in
fl at
um v
ar.
infl
atum
dese
rt t
rum
pet
Nat
ive
Pres
Mar
–Oct
xx
Poly
gona
ceae
Erio
gonu
m m
icro
thec
um v
ar.
sim
pson
iisl
ende
r w
ild b
uckw
heat
Nat
ive
Pres
Jun–
Oct
x
Poly
gona
ceae
Poly
gonu
m a
mph
ibiu
mw
ater
sm
artw
eed
Nat
ive
Rep
Jun–
Sep
x
Poly
gona
ceae
Rum
ex c
risp
uscu
rly
dock
Intr
oRe
pM
ay–S
epx
Poly
gona
ceae
Rum
ex h
ymen
osep
alus
cana
igre
doc
kN
ativ
ePr
esA
pr–J
unx
Ranu
ncul
acea
e (H
elle
bora
ceae
)D
elph
iniu
m a
nder
soni
i var
. sc
apos
umpa
le la
rksp
urN
ativ
eH
ist
Apr
–Jun
xx
Rosa
ceae
Pote
ntill
a gr
acili
s va
r. el
mer
ico
mb-
leaf
cin
quef
oil
Nat
ive
His
tJu
n–A
ugx
Scro
phul
aria
ceae
Cast
illej
a ch
rom
osa
dese
rt p
aint
brus
hN
ativ
ePr
esA
pr–J
unx
x
Scro
phul
aria
ceae
Cast
illej
a lin
ariif
olia
Wyo
min
g pa
intb
rush
Nat
ive
Pres
Jul–
Aug
xx
Scro
phul
aria
ceae
Pens
tem
on la
evis
smoo
th p
enst
emon
Nat
ive
Rep
May
–Jun
x
Scro
phul
aria
ceae
Pens
tem
on p
alm
eri v
ar.
egla
ndul
osus
Palm
er’s
pens
tem
onN
ativ
ePr
esM
ay–A
ugx
x
54 Annotated Checklist of Vascular Flora: Pipe Spring National Monument
App
endi
x D
. Con
fi rm
ed, h
isto
rica
l, an
d re
port
ed t
axa
of P
ipe
Spri
ng N
atio
nal M
onum
ent
orga
nize
d by
life
for
m a
nd c
omm
on n
ame
wit
h ec
olog
ical
not
es, c
ont.
Fam
ilySp
ecie
s na
me
Com
mon
nam
eN
ativ
ity
Park
st
atus
Flow
er
tim
eD
esSh
Pin
Jun
Wet
Com
men
ts
Pere
nnia
l For
bs, c
ont.
Scro
phul
aria
ceae
Pens
tem
on u
tahe
nsis
Uta
h pe
nste
mon
Nat
ive
His
tA
pr–J
unx
Sola
nace
aeCh
amae
sara
cha
coro
nopu
sfa
lse
nigh
tsha
deN
ativ
ePr
esA
pr–J
ulx
Sola
nace
aePh
ysal
is c
rass
ifol
iath
ick-
leav
ed g
roun
d-ch
erry
Nat
ive
Rep
Mar
–May
x
Sola
nace
aePh
ysal
is h
eder
ifol
ia v
ar.
palm
eri
Palm
er’s
grou
nd-c
herr
yN
ativ
eRe
pM
ay–O
ctx
Um
belli
fera
e (A
piac
eae)
Beru
la e
rect
a va
r. in
cisa
cutl
eaf
wat
er-p
arsn
ipN
ativ
ePr
esJu
n–Se
px
Um
belli
fera
e (A
piac
eae)
Cym
opte
rus
mul
tine
rvat
uspu
rple
-ner
ved
spri
ng-
pars
ley
Nat
ive
Pres
Mar
–Apr
x
Ver
bena
ceae
Ver
bena
bra
ctea
tapr
ostr
ate
verv
ain
Nat
ive
Pres
May
–Sep
xD
istu
rbed
are
as
Vis
cace
aePh
orad
endr
on ju
nipe
rinu
mju
nipe
r m
istl
etoe
Nat
ive
Pres
Jul–
Sep
x
Ann
ual F
orbs
Am
aran
thac
eae
Am
aran
thus
blit
oide
spr
ostr
ate
pigw
eed
Nat
ive
Pres
Jun–
Sep
Dis
turb
ed a
reas
Am
aran
thac
eae
Am
aran
thus
hyp
ocho
ndri
acus
grai
n am
aran
thIn
tro
Pres
Jun–
Oct
Cult
ivat
ed
Bora
gina
ceae
Cryp
tant
ha c
ircu
msc
issa
cush
ion
cryp
tant
hN
ativ
eH
ist
Apr
–Jul
x
Bora
gina
ceae
Cryp
tant
ha c
rass
isep
ala
var.
elac
hant
hath
ick-
sepa
led
cryp
tant
hN
ativ
eH
ist
Apr
–Jun
xx
Bora
gina
ceae
Cryp
tant
ha g
raci
lissl
ende
r cr
ypta
nth
Nat
ive
His
tA
pr–J
ulx
x
Bora
gina
ceae
Cryp
tant
ha m
icra
ntha
redr
oot
cryp
tant
hN
ativ
eH
ist
Mar
–May
x
Bora
gina
ceae
Cryp
tant
ha p
tero
cary
a va
r. cy
clop
tera
win
gnut
cry
ptan
thN
ativ
eH
ist
Apr
–Jul
x
Bora
gina
ceae
Cryp
tant
ha p
tero
cary
a va
r. pt
eroc
arya
win
gnut
cry
ptan
thN
ativ
ePr
esA
pr–J
ulx
x
Bora
gina
ceae
Hel
iotr
opiu
m c
onvo
lvul
aceu
mbi
ndw
eed
helio
trop
eN
ativ
eH
ist
May
–Sep
x
Bora
gina
ceae
Lapp
ula
occi
dent
alis
var
. cu
pula
tacu
psee
d st
icks
eed
Nat
ive
Pres
Apr
–Jun
Dis
turb
ed a
reas
Bora
gina
ceae
Plag
iobo
thry
s jo
nesi
iJo
nes’
pop
corn
-fl o
wer
Nat
ive
Pres
Apr
–May
x
Capp
arac
eae
(Cle
omac
eae)
Cleo
me
lute
aye
llow
bee
plan
tN
ativ
ePr
esA
pr–S
epx
Dis
turb
ed a
reas
Chen
opod
iace
aeA
trip
lex
arge
ntea
var
. ar
gent
easi
lver
ora
chN
ativ
ePr
esJu
n–Se
pD
istu
rbed
are
as
Chen
opod
iace
aeCh
enop
odiu
m a
lbum
var
. al
bum
lam
bsqu
arte
rIn
tro
Pres
Jun–
Sep
Dis
turb
ed a
reas
Appendices 55
App
endi
x D
. Con
fi rm
ed, h
isto
rica
l, an
d re
port
ed t
axa
of P
ipe
Spri
ng N
atio
nal M
onum
ent
orga
nize
d by
life
for
m a
nd c
omm
on n
ame
wit
h ec
olog
ical
not
es, c
ont.
Fam
ilySp
ecie
s na
me
Com
mon
nam
eN
ativ
ity
Park
st
atus
Flow
er
tim
eD
esSh
Pin
Jun
Wet
Com
men
ts
Ann
ual F
orbs
, con
t.
Chen
opod
iace
aeCh
enop
odiu
m f
rem
onti
i var
. fr
emon
tii
Frem
ont’s
goo
sefo
otN
ativ
ePr
esA
ug–O
ctx
Chen
opod
iace
aeCh
enop
odiu
m le
ptop
hyllu
mna
rrow
leaf
goo
sefo
otN
ativ
eRe
pJu
l–Se
px
Chen
opod
iace
aeSa
lsol
a tr
agus
Russ
ian
this
tle
Intr
oPr
esJu
n–Se
px
xD
istu
rbed
are
as
Chen
opod
iace
aeSu
aeda
tor
reya
na v
ar.
torr
eyan
aTo
rrey
’s se
epw
eed
Nat
ive
Pres
Jun–
Sep
x
Com
posi
tae
(Ast
erac
eae)
Am
bros
ia a
cant
hica
rpa
bur
ragw
eed
Nat
ive
Pres
Jul–
Oct
x
Com
posi
tae
(Ast
erac
eae)
Am
bros
ia a
rtem
isiif
olia
com
mon
rag
wee
dN
ativ
ePr
esJu
l–O
ctx
Dis
turb
ed a
reas
Com
posi
tae
(Ast
erac
eae)
Chae
nact
is c
arph
oclin
iaG
ray’
s ch
aena
ctis
Nat
ive
Pres
Apr
–Jun
x
Com
posi
tae
(Ast
erac
eae)
Cony
za c
anad
ensi
s va
r. gl
abra
taCa
nadi
an h
orse
wee
dN
ativ
eRe
pJu
l–Se
px
Com
posi
tae
(Ast
erac
eae)
Gai
llard
ia a
rizo
nica
Ari
zona
bla
nket
fl ow
erN
ativ
eRe
pA
pr–M
ayx
Com
posi
tae
(Ast
erac
eae)
Hel
iant
hus
annu
us s
sp.
lent
icul
aris
com
mon
sun
fl ow
erN
ativ
ePr
esJu
n–Se
px
Com
posi
tae
(Ast
erac
eae)
Lact
uca
serr
iola
pric
kly
lett
uce
Intr
oPr
esJu
l–Se
px
Dis
turb
ed a
reas
Com
posi
tae
(Ast
erac
eae)
Layi
a gl
andu
losa
tidy
tips
Nat
ive
Pres
Apr
–Jul
xx
Com
posi
tae
(Ast
erac
eae)
Mac
haer
anth
era
tana
ceti
folia
tans
ylea
f as
ter
Nat
ive
Pres
May
–Sep
xD
istu
rbed
are
as
Com
posi
tae
(Ast
erac
eae)
Mal
acot
hrix
gla
brat
afi
lifor
m d
eser
t-da
ndel
ion
Nat
ive
Pres
Apr
–Jul
x
Com
posi
tae
(Ast
erac
eae)
Mal
acot
hrix
son
choi
des
sow
this
tle
dese
rt-
dand
elio
nN
ativ
eH
ist
Apr
–Jun
x
Com
posi
tae
(Ast
erac
eae)
Sonc
hus
aspe
rsp
iny-
leaf
sow
-thi
stle
Intr
oPr
esJu
l–O
ctx
Com
posi
tae
(Ast
erac
eae)
Sonc
hus
oler
aceu
sco
mm
on s
owth
istl
eIn
tro
Rep
Jul–
Oct
x
Com
posi
tae
(Ast
erac
eae)
Step
hano
mer
ia e
xigu
aw
hite
-plu
me
wir
e le
ttuc
eN
ativ
ePr
esM
ay–S
ep
Com
posi
tae
(Ast
erac
eae)
Ver
besi
na e
ncel
ioid
es v
ar.
exau
ricu
lata
gold
en c
row
nbea
rdN
ativ
ePr
esM
ay–O
ctx
Dis
turb
ed a
reas
Com
posi
tae
(Ast
erac
eae)
Xan
thiu
m s
trum
ariu
m v
ar.
cana
dens
ero
ugh
cock
lebu
rN
ativ
eRe
pA
pr–O
ctx
Dis
turb
ed a
reas
Cruc
ifer
ae (B
rass
icac
eae)
Caps
ella
bur
sa-p
asto
ris
shep
herd
’s-pu
rse
Intr
oH
ist
Apr
–Sep
Dis
turb
ed a
reas
Cruc
ifer
ae (B
rass
icac
eae)
Des
cura
inia
pin
nata
var
. gl
abra
wes
tern
tan
sym
usta
rdN
ativ
eH
ist
Mar
–Aug
xx
Cruc
ifer
ae (B
rass
icac
eae)
Des
cura
inia
pin
nata
var
. in
term
edia
wes
tern
tan
sym
usta
rdN
ativ
eH
ist
Mar
–Aug
xx
56 Annotated Checklist of Vascular Flora: Pipe Spring National Monument
App
endi
x D
. Con
fi rm
ed, h
isto
rica
l, an
d re
port
ed t
axa
of P
ipe
Spri
ng N
atio
nal M
onum
ent
orga
nize
d by
life
for
m a
nd c
omm
on n
ame
wit
h ec
olog
ical
not
es, c
ont.
Fam
ilySp
ecie
s na
me
Com
mon
nam
eN
ativ
ity
Park
st
atus
Flow
er
tim
eD
esSh
Pin
Jun
Wet
Com
men
ts
Ann
ual F
orbs
, con
t.
Cruc
ifer
ae (B
rass
icac
eae)
Des
cura
inia
pin
nata
var
. os
mia
rum
wes
tern
tan
sym
usta
rdN
ativ
ePr
esM
ar–A
ugx
x
Cruc
ifer
ae (B
rass
icac
eae)
Des
cura
inia
sop
hia
fl ix
wee
dIn
tro
Pres
Mar
–Aug
xD
istu
rbed
are
as
Cruc
ifer
ae (B
rass
icac
eae)
Dit
hyre
a w
isliz
eni
spec
tacl
epod
Nat
ive
Pres
Apr
–Sep
x
Cruc
ifer
ae (B
rass
icac
eae)
Erys
imum
rep
andu
msp
read
ing
wal
lfl ow
erIn
tro
His
tA
pr–J
un
Cruc
ifer
ae (B
rass
icac
eae)
Lepi
dium
den
sifl
orum
var
. ra
mos
umpr
airi
e pe
pper
wor
tN
ativ
eH
ist
May
–Jul
x
Cruc
ifer
ae (B
rass
icac
eae)
Lepi
dium
per
folia
tum
clas
ping
pep
perw
ort
Intr
oH
ist
May
–Jul
Dis
turb
ed a
reas
Cruc
ifer
ae (B
rass
icac
eae)
Sisy
mbr
ium
alt
issi
mum
tum
ble
mus
tard
Intr
oPr
esM
ay–A
ugx
xD
istu
rbed
are
as
Cruc
ifer
ae (B
rass
icac
eae)
Stre
ptan
thel
la lo
ngir
ostr
islo
ng-b
eak
fi dd
lem
usta
rdN
ativ
eH
ist
Mar
–Jun
xx
Cype
race
aeCy
peru
s sp
.fl
at-s
edge
Intr
oRe
pJu
n–Se
pD
istu
rbed
are
as
Euph
orbi
acea
eCh
amae
syce
gly
ptos
perm
ari
dge-
seed
ed s
purg
eN
ativ
ePr
esJu
n–A
ugx
Dis
turb
ed a
reas
Euph
orbi
acea
eCh
amae
syce
pro
stra
tapr
ostr
ate
spur
geIn
tro
Pres
May
–Sep
Dis
turb
ed a
reas
Ger
ania
ceae
Erod
ium
cic
utar
ium
stor
k’s-
bill
Intr
oPr
esFe
b–O
ctx
xD
istu
rbed
are
as
Hyd
roph
ylla
ceae
Phac
elia
cre
nula
ta v
ar.
corr
ug
ata
corr
ugat
e ph
acel
iaN
ativ
eH
ist
May
–Jun
x
Hyd
roph
ylla
ceae
Phac
elia
cre
nula
ta v
ar.
cren
ulat
acr
enul
ate
phac
elia
Nat
ive
Pres
May
–Jun
x
Hyd
roph
ylla
ceae
Phac
elia
ives
iana
Ives
’ pha
celia
Nat
ive
Pres
Apr
–Jun
xx
Hyd
roph
ylla
ceae
Phac
elia
pul
chel
la v
ar.
good
ding
iiG
oodd
ing’
s pr
etty
ph
acel
iaN
ativ
eH
ist
Apr
–May
x
Legu
min
osae
(Fab
acea
e)Lu
pinu
s pu
sillu
s va
r. pu
sillu
sdw
arf
lupi
neN
ativ
eH
ist
Apr
–Jul
x
Legu
min
osae
(Fab
acea
e)M
elilo
tus
alba
whi
te s
wee
tclo
ver
Intr
oRe
pM
ay–O
ctx
xx
Dis
turb
ed a
reas
Lina
ceae
Linu
m a
rist
atum
broo
mfl
axN
ativ
ePr
esA
pr–J
unx
Loas
acea
eM
entz
elia
alb
icau
lisw
hite
-ste
m b
lazi
ngst
arN
ativ
ePr
esA
pr–A
ugx
Mal
vace
aeM
alva
neg
lect
aco
mm
on m
allo
wIn
tro
Pres
Apr
–Oct
Dis
turb
ed a
reas
Ona
grac
eae
Cam
isso
nia
cham
aene
rioi
des
slen
der-
pod
cam
isso
nia
Nat
ive
Pres
Apr
–Jun
x
Ona
grac
eae
Cam
isso
nia
parr
yiPa
rry’
s ca
mis
soni
aN
ativ
eH
ist
May
–Jun
x
Ona
grac
eae
Cam
isso
nia
parv
ula
tiny
cam
isso
nia
Nat
ive
His
tM
ay–J
unx
x
Ona
grac
eae
Gau
ra p
arvi
fl or
asm
all-fl
ow
ered
gau
raN
ativ
ePr
esM
ay–A
ugD
istu
rbed
are
as
Ona
grac
eae
Oen
othe
ra a
lbic
aulis
whi
te-s
tem
eve
ning
pr
imro
seN
ativ
eH
ist
Mar
–Jun
x
Appendices 57
App
endi
x D
. Con
fi rm
ed, h
isto
rica
l, an
d re
port
ed t
axa
of P
ipe
Spri
ng N
atio
nal M
onum
ent
orga
nize
d by
life
for
m a
nd c
omm
on n
ame
wit
h ec
olog
ical
not
es, c
ont.
Fam
ilySp
ecie
s na
me
Com
mon
nam
eN
ativ
ity
Park
st
atus
Flow
er
tim
eD
esSh
Pin
Jun
Wet
Com
men
ts
Ann
ual F
orbs
, con
t.
Ona
grac
eae
Oen
othe
ra p
rim
iver
is v
ar.
bufo
nus
earl
y ev
enin
g pr
imro
seN
ativ
eRe
pM
ar–M
ayx
Papa
vera
ceae
Esch
scho
lzia
min
utifl
ora
pygm
y go
lden
-pop
pyN
ativ
ePr
esFa
b–M
ayx
Sand
y so
il
Pole
mon
iace
aeEr
iast
rum
ere
mic
umM
ohav
e er
iast
rum
Nat
ive
His
tA
pr–J
un
Pole
mon
iace
aeG
ilia
inco
nspi
cua
shy
gilia
Nat
ive
Pres
Apr
–Jul
xx
Pole
mon
iace
aeG
ilia
lept
omer
ia v
ar.
lept
omer
iaco
mm
on g
ilia
Nat
ive
His
tM
ay–J
unx
x
Pole
mon
iace
aeIp
omop
sis
poly
clad
onsp
read
ing
gilia
Nat
ive
His
tA
pr–J
unx
Pole
mon
iace
aeLa
nglo
isia
set
osis
sim
aM
ohav
e la
nglo
isia
Nat
ive
His
tM
ar–M
ayx
Poly
gona
ceae
Cent
rost
egia
thu
rber
iTh
urbe
r’s s
pine
fl ow
erN
ativ
eH
ist
Mar
–Jul
x
Poly
gona
ceae
Erio
gonu
m c
ernu
um v
ar.
cern
uum
nodd
ing
wild
buc
kwhe
atN
ativ
eH
ist
Apr
–Oct
xx
Poly
gona
ceae
Erio
gonu
m d
efl e
xum
skel
eton
wee
d w
ild
buck
whe
atN
ativ
eH
ist
Apr
–Oct
x
Poly
gona
ceae
Erio
gonu
m p
alm
eria
num
Palm
er’s
wild
buc
kwhe
atN
ativ
ePr
esM
ar–O
ctx
Poly
gona
ceae
Poly
gonu
m a
vicu
lare
yard
kno
twee
dIn
tro
Pres
May
–Oct
Dis
turb
ed a
reas
Port
ulac
acea
ePo
rtul
aca
oler
acea
com
mon
pur
slan
eIn
tro
Pres
Jun–
Sep
xD
istu
rbed
are
as
Scro
phul
aria
ceae
Cord
ylan
thus
par
vifl
orus
smal
lfl ow
er b
ird’
s-be
akN
ativ
ePr
esA
ug–S
epx
Sola
nace
aeD
atur
a w
righ
tii
ange
l’s t
rum
pet
Nat
ive
Pres
Apr
–Oct
xD
istu
rbed
are
as
Sola
nace
aeN
icot
iana
tri
gono
phyl
lade
sert
tob
acco
Nat
ive
His
tM
ar–O
ctx
Sola
nace
aeSo
lanu
m s
arra
choi
des
grou
nd-c
herr
y ni
ghts
hade
Intr
oRe
pM
ay–O
ctD
istu
rbed
are
as
Zygo
phyl
lace
aeTr
ibul
us t
erre
stri
spu
nctu
revi
neIn
tro
Pres
May
–Sep
Dis
turb
ed a
reas
Pere
nnia
l Gra
min
oids
Cype
race
aeCa
rex
nebr
asce
nsis
Neb
rask
a se
dge
Nat
ive
Pres
x
Cype
race
aeCa
rex
prae
grac
ilis
blac
kcre
eper
sed
geN
ativ
ePr
esM
ay–A
ugx
Cype
race
aeEl
eoch
aris
par
ishi
iPa
rish
’s sp
iker
ush
Nat
ive
Pres
Apr
–Sep
x
Cype
race
aeSc
irpu
s pu
ngen
s va
r. lo
ngis
pica
tus
thre
e-sq
uare
bul
rush
Nat
ive
Pres
May
–Aug
x
Gra
min
eae
(Poa
ceae
)A
grop
yron
cri
stat
umcr
este
d w
heat
gras
sIn
tro
Rep
Jun–
Aug
xCu
ltiv
ated
Gra
min
eae
(Poa
ceae
)A
gros
tis
stol
onif
era
redt
opIn
tro
Pres
Jun–
Sep
x
Gra
min
eae
(Poa
ceae
)A
rist
ida
purp
urea
purp
le t
hree
-aw
nN
ativ
ePr
esM
ar–S
epx
xD
istu
rbed
are
as
Gra
min
eae
(Poa
ceae
)Bo
utel
oua
curt
ipen
dula
side
oats
gra
ma
Nat
ive
Rep
Jul–
Sep
x
58 Annotated Checklist of Vascular Flora: Pipe Spring National Monument
App
endi
x D
. Con
fi rm
ed, h
isto
rica
l, an
d re
port
ed t
axa
of P
ipe
Spri
ng N
atio
nal M
onum
ent
orga
nize
d by
life
for
m a
nd c
omm
on n
ame
wit
h ec
olog
ical
not
es, c
ont.
Fam
ilySp
ecie
s na
me
Com
mon
nam
eN
ativ
ity
Park
st
atus
Flow
er
tim
eD
esSh
Pin
Jun
Wet
Com
men
ts
Pere
nnia
l Gra
min
oids
, con
t.
Gra
min
eae
(Poa
ceae
)Bo
utel
oua
erio
poda
blac
k gr
ama
Nat
ive
Pres
Jun–
Aug
x
Gra
min
eae
(Poa
ceae
)Bo
utel
oua
grac
ilis
blue
gra
ma
Nat
ive
Pres
Jul–
Sep
xx
Gra
min
eae
(Poa
ceae
)Br
omus
iner
mis
var
. ine
rmis
smoo
th b
rom
eIn
tro
Rep
Jun–
Aug
x
Gra
min
eae
(Poa
ceae
)D
isti
chlis
spi
cata
de
sert
sal
tgra
ssN
ativ
eRe
pJu
n–Se
px
Gra
min
eae
(Poa
ceae
)El
ymus
elo
ngat
us
tall
whe
atgr
ass
Intr
oRe
pJu
n–A
ugx
Gra
min
eae
(Poa
ceae
)El
ymus
ely
moi
des
squi
rrel
tail
Nat
ive
Rep
May
–Aug
xx
Gra
min
eae
(Poa
ceae
)El
ymus
his
pidu
s in
term
edia
te w
heat
gras
sIn
tro
Rep
Jun–
Aug
Dis
turb
ed a
reas
Gra
min
eae
(Poa
ceae
)El
ymus
sm
ithi
iw
este
rn w
heat
gras
sN
ativ
eRe
pJu
n–A
ugx
Gra
min
eae
(Poa
ceae
)El
ymus
tra
chyc
aulu
s sl
ende
r w
heat
gras
sN
ativ
eH
ist
Jun–
Aug
x
Gra
min
eae
(Poa
ceae
)Er
ione
uron
pul
chel
lum
fl uf
fgra
ssN
ativ
eRe
pA
pr–A
ugx
x
Gra
min
eae
(Poa
ceae
)Fe
stuc
a ar
undi
nace
ata
ll fe
scue
Intr
oRe
pM
ay–J
ulx
Gra
min
eae
(Poa
ceae
)H
ilari
a ja
mes
iiga
lleta
Nat
ive
Pres
May
–Aug
xx
Gra
min
eae
(Poa
ceae
)H
orde
um ju
batu
mfo
xtai
l bar
ley
Nat
ive
Rep
Jun–
Aug
x
Gra
min
eae
(Poa
ceae
)M
uhle
nber
gia
aspe
rifo
liasc
ratc
hgra
ssN
ativ
ePr
esJu
l–A
ugx
Gra
min
eae
(Poa
ceae
)M
uhle
nber
gia
port
eri
bush
muh
lyN
ativ
ePr
esJu
l–O
ctx
Gra
min
eae
(Poa
ceae
)Po
a fe
ndle
rian
am
utto
ngra
ssN
ativ
eRe
pM
ay–A
ugx
x
Gra
min
eae
(Poa
ceae
)Po
a pr
aten
sis
Ken
tuck
y bl
uegr
ass
Intr
oRe
pM
ay–A
ugx
Gra
min
eae
(Poa
ceae
)Po
lypo
gon
sem
iver
tici
llatu
sw
ater
pol
ypog
onIn
tro
Rep
Jul–
Aug
xD
istu
rbed
are
as
Gra
min
eae
(Poa
ceae
)Sp
orob
olus
air
oide
s va
r. ai
roid
esal
kali
saca
ton
Nat
ive
Rep
Jun–
Aug
xx
Gra
min
eae
(Poa
ceae
)Sp
orob
olus
con
trac
tus
spik
e dr
opse
edN
ativ
ePr
esJu
n–Se
px
x
Gra
min
eae
(Poa
ceae
)Sp
orob
olus
cry
ptan
drus
sand
dro
psee
dN
ativ
ePr
esJu
n–A
ugx
Gra
min
eae
(Poa
ceae
)Sp
orob
olus
fl ex
uosu
sm
esa
drop
seed
Nat
ive
Pres
Jun–
Sep
xx
Gra
min
eae
(Poa
ceae
)St
ipa
com
ata
var.
com
ata
need
le-a
nd-t
hrea
dN
ativ
ePr
esM
ay–J
ulx
x
Gra
min
eae
(Poa
ceae
)St
ipa
hym
enoi
des
Indi
an r
iceg
rass
Nat
ive
Pres
May
–Jul
xx
Gra
min
eae
(Poa
ceae
)St
ipa
spec
iosa
dese
rt n
eedl
egra
ssN
ativ
ePr
esA
pr–J
unx
x
Junc
acea
eJu
ncus
arc
ticu
sBa
ltic
rus
hN
ativ
ePr
esM
ay–J
ulx
Junc
acea
eJu
ncus
ens
ifol
ius
var.
brun
nesc
ens
Trac
y’s
rush
Nat
ive
His
tJu
n–A
ugx
Junc
acea
eJu
ncus
tor
reyi
Torr
ey’s
rush
Nat
ive
Pres
Jun–
Aug
x
Appendices 59
App
endi
x D
. Con
fi rm
ed, h
isto
rica
l, an
d re
port
ed t
axa
of P
ipe
Spri
ng N
atio
nal M
onum
ent
orga
nize
d by
life
for
m a
nd c
omm
on n
ame
wit
h ec
olog
ical
not
es, c
ont.
Fam
ilySp
ecie
s na
me
Com
mon
nam
eN
ativ
ity
Park
st
atus
Flow
er
tim
eD
esSh
Pin
Jun
Wet
Com
men
ts
Ann
ual G
ram
inoi
ds
Gra
min
eae
(Poa
ceae
)Bo
utel
oua
barb
ata
sixw
eeks
gra
ma
Nat
ive
Pres
Aug
–Oct
x
Gra
min
eae
(Poa
ceae
)Br
omus
dia
ndru
sri
pgut
bro
me
Intr
oPr
esA
pr–J
unx
xD
istu
rbed
are
as
Gra
min
eae
(Poa
ceae
)Br
omus
rub
ens
red
brom
eIn
tro
Pres
Mar
–Jun
x
Gra
min
eae
(Poa
ceae
)Br
omus
tec
toru
mch
eatg
rass
Intr
oPr
esM
ay–J
ulx
xx
Dis
turb
ed a
reas
Gra
min
eae
(Poa
ceae
)Ce
nchr
us lo
ngis
pinu
sfi
eld
sand
bur
Nat
ive
Rep
Jul–
Sep
xD
istu
rbed
are
as
Gra
min
eae
(Poa
ceae
)Ec
hino
chlo
a cr
us-g
alli
barn
yard
gras
sIn
tro
Rep
Aug
–Sep
Dis
turb
ed a
reas
Gra
min
eae
(Poa
ceae
)Er
agro
stis
cili
anen
sis
stin
kgra
ssIn
tro
Rep
Jul–
Sep
Dis
turb
ed a
reas
Gra
min
eae
(Poa
ceae
)Er
agro
stis
mex
ican
aM
exic
an lo
vegr
ass
Intr
oRe
pJu
l–O
ctD
istu
rbed
are
as
Gra
min
eae
(Poa
ceae
)Er
agro
stis
pec
tina
cea
tuft
ed lo
vegr
ass
Nat
ive
Pres
Jul–
Aug
Dis
turb
ed a
reas
Gra
min
eae
(Poa
ceae
)H
orde
um m
urin
umra
bbit
bar
ley
Intr
oPr
esM
ay–J
ulx
Dis
turb
ed a
reas
Gra
min
eae
(Poa
ceae
)M
unro
a sq
uarr
osa
fals
e bu
ffal
ogra
ssN
ativ
ePr
esJu
n–Se
px
Gra
min
eae
(Poa
ceae
)Po
lypo
gon
mon
spel
iens
isra
bbit
sfoo
tgra
ssIn
tro
Pres
Jun–
Aug
xD
istu
rbed
are
as
60 Annotated Checklist of Vascular Flora: Pipe Spring National Monument
The Department of the Interior protects and manages the nation’s natural resources and cultural heritage; provides scientifi c and other information about those resources; and honors its special responsibilities to American Indians, Alaska Natives, and affi liated Island Communities.
NPS D-222, October 2008
Natural Resource Program Center 1201 Oak Ridge Drive, Suite 150 Fort Collins, Colorado 80525
www.nature.nps.gov
National Park ServiceU.S. Department of the Interior
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