Animal Population Dynamics

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    ANIMAL POPULATION DYNAMICS

    The constituents that influence the rate of change in

    the numbers of a particular species found in the

    wild include such density-dependent factors.

    Environmental aspects such as weather,

    temperature, flooding, snowfall, all of which are

    density-independent will also affect populationdynamics.

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    FIVE BASIC COMPONENTS

    Birth

    Death

    Gender Ratio

    Age Structure

    Dispersal

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    A number of components affect a populations

    birth rate:

    The amount and quantity of food Age at first reproduction

    The birth interval

    The average number of young born per pregnancy

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    BIRTHor NATALITY RATE

    Is the total number of births per 1000 of apopulation each year.

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    DEATH or MORTALITY RATE

    Is defined as the number of animals thatdie per unit time divided by the number

    of animals alive at the beginning of that

    time period.

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    GENDER RATIO

    Is the proportion of males to

    females within a population.

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    AGE STRUCTURE

    Percentage of the population at

    each age level in a population.

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    DISPERSAL

    Is defined as the movement of an animal

    from the location of its birth to a new area

    where it lives and reproduces.

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    The simplest (exponential) model assumes that the

    resources necessary for population growth are unlimited.

    Therefore, the population grows at an exponential rate,which is the maximum rate possible for that particular

    species:

    = rN (Eq.1)

    where

    -the change in numbers of animals within a

    certain population per unit timerthe specific rate of change

    Nnumber of animals within a certain population

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    If Nois the no. of organisms attime zero, the no. of organisms

    at any particular time, N can be

    determined by integrating Eq.1over that particular time period:

    N = Noexp(r) Noer

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    Growth occurs in discrete intervals describedby the term .

    (+)() = =

    where N(+1) - population after (+1) no. in yearsNpopulation after years

    rthe specific growth rate (net new organisms per

    unit time)

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    Example 1: Use the ff. data along with the exponential

    model to determine the population of the predicted

    eastern gray wolf in the state of Wisconsin in the year

    2005. Compare that result with that obtained with the

    geometric model.

    Year 1975 1980 1990 1995 1996 1997 1998 1999

    Number 8 22 45 83 99 148 180 200

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    Solution:

    N() = N(0)e0.123()= 8e0.123(30)

    = 320 wolves

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    For year 0 to year 1: 8 x = a

    For year 1 to year 2: a x = b

    For year 2 to year 3: b x = c

    And so on until we get

    year 4 to year 5: x = d

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    8 x 5

    = N(5)

    where N(5) is the no. of

    individuals 5 years later; and

    is equal to 22. So

    8 x 5= 22

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    Solving for yields = 1.224

    =

    where is the year of calculation and 0is the firstyear for which data is available.

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    YEAR NUMBER 0 8

    5 22 1.224

    15 45 1.122

    20 83 1.124

    21 99 1.127

    22 148 1.142

    23 180 1.145

    24 200 1.144

    Average: 1.147

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    N(30) = N(24) x (1.1476)

    = 455 individuals

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    The population change can be represented

    by the model:

    = rN[

    ]

    N() =

    +()

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    Example 2: Assume that the population of the greater

    roadrunner in the Guadelope Desert was 200 per

    hectare at the beginning of 1999. If the carryingcapacity, K, is 600 and r = 0.25 . year-1, what is the no.

    of roadrunners one, five & ten years? What happens

    when the no. of roadrunners equals K?

    SOLUTION:

    N(1) =

    +()^(.)

    = 234 roadrunners

    N(5) = 381 roadrunners

    N(10) = 515 roadrunners

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    Numerous more complex models exist. These include

    phenomena known as monotonic damping, damped oscillations,

    limiting cycles, or chaotic dynamics. A number of these models

    can also be used to describe plant population dynamics.

    Much more complex models describe the interactions between

    species by considering predator-prey relationships. These

    models show how the interactions between two species result

    in periodic behavior.

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    The models use the following two differential equations to describe

    the numbers of predators, K, and prey, P:

    =

    =

    where a = growth rate of the prey

    b = mortality parameter of the prey

    c = growth rate of the predator

    d = mortality parameter of the prey

    These equations are often referred to as the LOTKA-VOLTERRA

    MODEL.

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    HUMAN POPULATION DYNAMICS

    Human population dynamics also depend on birth, death,

    gender ratio, age structure, and dispersal. In humans,

    cultural factors play a significant role. In human populations,dispersalis referred to as immigrationand emigration.

    The effect of cultural differences can be easily illustrated

    using population pyramids. Population pyramids display theage by gender data of a community at one point in time.

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    Population pyramids

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    Assuming an exponential growth rate, the population can be

    predicted using the equation:

    P(t) = Po

    where P(t) - the population at time, t

    Po- population at time, 0

    r - rate of growthttime

    The growth rate can be determined as a function of the birth rate

    (b), death rate (d), immigration rate (i), and emigration rate (m):

    r = b - d + im

    where the rates are all expressed as some value per unit time.

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    Example 2: A population of humanoids on the island of

    Huronth on the planet Szacak has a net birth rate (b) of 1.0

    individuals/(individual x year) and a net death rate (d) of 0.9

    individuals/(individual x year). Assume that the netimmigration rate is equal to the net emigration rate. How many

    years are required for the population to double? If in year zero,

    the population on the island is 85, what is the population 50

    years later?Solution:

    r = 1.00.9 =.

    tdouble=

    =

    .

    = 6.93 years

    No= 85

    t = 50 years

    r = 0.1N50= No

    = 85 . ()

    = 12,615 humanoids

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    TOTAL FERTILITY RATE

    Is the number of children a woman will have over herentire lifetime.