The aim of the present paper is to review theColeophoridae of India and neighbouring regions, i. e.the former ‘British India’. These were described byEdward Meyrick between 1917 and 1933. Since thenthere has been no further work on Indian Coleo-phoridae, and since the independence of India in 1948no more species have been recorded from that vast re-gion, which undoubtedly has a very interesting fauna.Recently a number of species have been recorded fromPakistan (Baldizzone 1994). These are not treated inthis paper (except for the check-list), in which onlyMeyrick’s species are revised. In total Meyrick de-scribed 20 species, 19 in Coleophora Hübner and onein Macrocorystis Meyrick, here synonymised withColeophora. Three species are transferred to other fam-ilies in Gelechioidea and one is synonymised, so thatstill 16 Coleophora species named by Meyrick arerecognised.

MATERIAL

All the Meyrick types studied are in the NaturalHistory Museum in London. Not all specimens werestudied, but representative syntype specimens (withsuggestions for lectotype selection) were selected byDr. K. Sattler or Mr. K. Tuck for study by the seniorauthor. From these eight lectotypes were selected.The identity of Coleophora species can only be estab-lished with certainty on the basis of genitalia charac-ters; therefore lectotype selection of specimens withgenitalia preparation is essential to prevent futureconfusion (ICZN article 74.7.3).

The label information of types is cited in full, ‘1/3’indicates the first specimen of a series of three of thatspecies in the Meyrick collection (not necessarily allof which are types).

1

GIORGIO BALDIZZONE* & HUGO W. VAN DER WOLF

Asti, Italy & Nuenen, The Netherlands

A TAXONOMIC REVIEW OF THE COLEOPHORIDAE

(LEPIDOPTERA) OF THE INDIAN SUBCONTINENT

AND SRI LANKA DESCRIBED BY EDWARD MEYRICK

*Contribution to the knowledge of the Coleophoridae, C

Baldizzone, G., & H. W. van der Wolf, 2003. A taxonomic review of the Coleophoridae (Lepi-doptera) of the Indian subcontinent and Sri Lanka described by Edward Meyrick. Contribu-tion to the knowledge of the Coleophoridae, C. – Tijdschrift voor Entomologie 146: 1-31, figs.1-89. [ISSN 0040-7496]. Published 1 June 2003.The present taxonomic knowledge of the Coleophoridae of the Indian subcontinent and SriLanka is reviewed. Nineteen species are treated here, all described by E. Meyrick; three of themare transferred from Coleophoridae because of external morphology and genitalia structure:Coleophora aphypnota Meyrick, 1917 is transferred to Batrachedridae, C. crossophanes Meyrick,1917 to Momphidae (provisional), and C. tarsocoma Meyrick, 1917 to Cosmopterigidae. Thefollowing new synonymies are established: the monotypic Macrocorystis Meyrick, 1931 is a ju-nior synonym of Coleophora Hübner, 1822, C. enchorda Meyrick, 1931 a junior synonym of C.versurella Zeller, 1849, and C. sarobiensis Toll & Amsel, 1967 a junior synonym of C. niphome-sta Meyrick, 1917. Coleophora byrsostola (Meyrick, 1931) is a new combination (from Macroco-rystis). Lectotypes are selected for 8 species. A checklist of the 34 Coleophoridae of the Indiansubcontinent and Sri Lanka is provided. Dr. G. Baldizzone, Via Manzoni, 24, I-14100 Asti, Italy. E-mail: bldz@libero.it [correspond-ing author]H. W. van der Wolf, Andriesplein 2, NL 5671 VS Nuenen, The Netherlands. E-mail: van-der.wolf@hccnet.nlKey words. – Coleophoridae; Batrachedridae; Momphidae; Cosmopterigidae; Indian subconti-nent; India; Sri Lanka; new synonymies.

Abbreviations for MuseumsDEI Deutsches Entomologisches Institut, Eber-

swalde, Germany.BMNH Natural History Museum, London, U.K.USNM United States National Museum, Smithson-

ian Institution, Washington, USA.

Checklist of the Coleophoridae of the Indiansubcontinent and Sri Lanka

Coleophora Hübner, 1822Macrocorystis Meyrick, 1931 syn. n.

crypsiphanes Meyrick, 1917; Sri Lankaaegra Meyrick, 1917; Pakistan, Afghanistanthiophaea Meyrick, 1917; Indiaalcyonipennella (Kollar, 1832); Palaearctic, Pakistan,

Australia (intr.), New Zealand (intr.)statherota Meyrick, 1917; Sri Lankaparaptarmica Toll & Amsel, 1967; Central Asia,

Pakistankabulensis Baldizzone, 1994; Afghanistan, Pakistansogdianae Baldizzone, 1994; Iran, Afghanistan, Paki-

stanswatensis Baldizzone, 1994; Iran, Afghanistan, Paki-

stansalviella Chrétien, 1917; North Africa, Central Asia,

Pakistanparthenica Meyrick, 1891; Palaearctic, Pakistan,

U.S.A.(intr.)galligena Falkovitsh, 1970; Central Asia, Jordan,

Pakistanklimeschiella, Toll, 1952; Palaearctic, Pakistan, U.S.A

(intr.)byrsostola (Meyrick, 1931)(Macrocorystis) comb. n.;

Indiacentrota Meyrick, 1917; Indialongiductella Baldizzone, 1989; Iran, Pakistantollamseliella Oudejans, 1971; Central Asia, Pakistancaelebipennella Zeller, 1839; Palaearctic, Pakistanargopleura Meyrick, 1917; Afghanistan, Kashmir,

Pakistancastalia Meyrick, 1930; Afghanistan, Kashmir, Pak-

istanversurella Zeller, 1849; Holarctic, India, Argentina

(intr.), Chile (intr.)enchorda Meyrick, 1931 syn. n.

aphrocrossa Meyrick, 1933; Kashmir polichomriensis Toll & Amsel, 1967; Afghanistan,

Pakistanabbasella Baldizzone, 1994; Iran, Pakistanpakistana Baldizzone, 1994; Pakistanechyropis Meyrick, 1930 ; Kashmirogmotona Meyrick, 1917; India, Sri Lankapercnoceros Meyrick, 1933; Kashmirniphomesta Meyrick, 1917; Turkmenistan, Iran,

Afghanistan, Pakistan

sarobiensis Toll & Amsel, 1967 syn. n.hydrella Baldizzone, 1994; Pakistangabralensis Baldizzone, 1994; Pakistangedrosiae Baldizzone, 1994; Iran, Pakistanchordoscelis Meyrick, 1917; Indialeucochares Meyrick, 1922; India

Species transferred from Coleophoridae:‘Coleophora’ aphypnota Meyrick, 1917 to Batrachedri-

dae‘Coleophora’ crossophanes Meyrick, 1917 to Momphi-

dae (provisional)‘Coleophora’ tarsocoma Meyrick, 1917 to Cosmopte-

rigidae

Genus Coleophora Hübner

Coleophora Hübner, 1822: 67. Type species (by monotypy):Tinea anatipennella Hübner, 1796: pl. 27, fig. 186.

Macrocorystis Meyrick, 1931: 49. Type species (by mono-typy): Macrocorystis byrsostola Meyrick, 1931. syn. n.

Remark. – The structure of the genitalia of Macro-corystes byrsostola are those of a Coleophora. ThusMacrocorystis is a junior subjective synonym, see fur-ther under C. byrsostola.

Coleophora crypsiphanes Meyrick (figs. 3, 21-24, 25-27).

Coleophora crypsiphanes Meyrick, 1917:70. Lectotype ?(here designated). SRI LANKA: – ‘LECTOTYPE’ (round blue-edged BMNH label), ‘Maskeliya, Ceylon, Pole, 11. 06 ‘,‘crypsiphanes Meyr.’ (Meyrick’s handwritten label),‘Coleophora crypsiphanes Meyr., 1/3, E. Meyrick det., inMeyrick coll.’, ‘Meyrick Coll., B.M. 1938-290’, ‘B.M.Genitalia Slide 24477 ?’, ‘Lectotype ?, Coleophora cryp-siphanes Meyr., teste K. Sattler, 1988’ (BMNH) [examined].

Coleophora crypsophanes Vives Moreno, 1988: 69. [Mis-spelling]

Additional material examined. – SRI LANKA:1/, paralec-totype ‘Maskeliya, Ceylon, Pole, 5.06’, ‘Coleophora cryp-siphanes Meyr. /, 2/3, E. Meyrick det., in Meyrick coll.’,‘Meyrick coll., BM. 1938-290’, Slide 24478 (BMNH).

Description. – The original description is correct:‘? 10 mm., / 14-16 mm. Head and thorax glossywhitish-grey or grey-whitish. Palpi and antennae sim-ple, grey-whitish. Abdomen whitish-grey. Forewingsnarrowly elongate-lanceolate; greyish-ochreous, withstrong silvery reflection, in / irrorated with dark grey,especially towards costa and posteriorly; second discalstigma rather large, blackish: cilia ochreous-grey-whitish, greyer towards tornus. Hindwings light grey;cilia whitish-grey.’ (Meyrick 1917: 70)

Male genitalia (fig. 21-22). – Spinose part ofgnathos globular. Tegumen compact. Transtilla nar-row and long, medially joined. Valvula indistinct. Cu-

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cullus long. Sacculus very simple, small, lateral marginending ventrally in a rounded, sclerotized point. Phal-lotheca conical. Vesica without cornuti, well sclero-tized on ventral margin.

Structure of abdominal supports (fig. 23). – Malewith posterior lateral struts, about half length of ante-rior ones. Transverse strut characterised by straightproximal margin and convex distal margin, mergingmedially with proximal one. Tergal disks with rathersmall conical spines, about 6 or 7 times longer thanwide. Female without posterior lateral struts, and withtransverse strut almost straight, both margins slenderand parallel. Tergal disks (3rd tergite) about 3 timeslonger than wide.

Female genitalia (fig. 25-26). – Papillae anales large,long. Apophyses posteriores about half length of ante-riores. Sterigma subtrapezoid, membranous, sclero-tized only at distal margin and at two lateral strips atostium bursae originating at base of apophyses anteri-ores and reaching proximal margin. Membranous os-tium bursae small, funnel-shaped. Short colliculum,beaker-shaped, with very small internal spines. Ductusbursae completely transparent, about twice length ofsterigma. Bursa copulatrix bag-shaped, very big andlong, with striking heavily sclerotized sword-shapedsignum.

Remark. – This species does not belong to anygroup of palaearctic Coleophoridae, but resemblescertain species of the afrotropical region, which willbe treated in a monograph by the authors.

Biology. – Unknown.Distribution. – Sri Lanka: Maskeliya.

Coleophora aegra Meyrick (figs. 4, 28-31)

Coleophora aegra Meyrick, 1917:71. Lectotype ? (here des-ignated), INDIA. – ‘LECTOTYPE’ (round blue-edged BMNH

label), ‘Abbottabad, NW – India, TBF. 10. 6. 16’, ‘aegraMeyr.’ (Meyrick’s handwritten label), ‘Coleophora aegraMeyr., 1/1, E. Meyrick det., in Meyrick coll.’, ‘MeyrickColl., BM 1938-290’, ‘B.M. Genitalia Slide ? 24452’,‘Lectotype ?, Coleophora aegra Meyr., teste K. Sattler,1987’ (BMNH) [examined].

Additional material examined. – AFGHANISTAN: 2?, 25km N Barikot, 1800 m, Nuristan, 12.-17. vii. 1963, Kasy &Vartian, slide Bldz 8180 (coll. Baldizzone). – PAKISTAN: 1?,N-Pakistan, Murree, 29.vii.1979, slide Patzak 2036 (DEI).

Description. – The original description is: ‘10-11mm. Head glossy grey, sides of crown whitish. Palpismooth, grey. Antennae simple, white, ringed abovewith grey. Thorax and abdomen dark grey. Forewingselongate-lanceolate; dark grey: cilia grey, sprinkledwith dark gey towards base. Hindwings dark grey; ciliagrey.’ It should be noted that the forewing is dull grey,speckled with darker scales.

Male genitalia (figs. 28-30). – Spinose part ofgnathos large, globular. Tegumen medially constrict-ed. Transtilla small, triangularly extended. Valvulasuboval, long, poorly delineated. Cucullus short,rounded. Sacculus characterised by sharp subtriangu-lar process. Phallotheca long, conical, slightly curved.A single cornutus in the form of a long spine.

Structure of abdominal supports (fig. 31). – Noposterior lateral struts. Transverse strut almost straightat proximal margin, slightly convex at distal margin,which is thicker near disks of second tergite. Tergaldisks (3rd tergite), with short conical spines, abouttwice as long as wide.

Remark. – This species, of which the female is un-known, belongs to the second group of Toll’s system(1953, 1962), section C. serratella (Linnaeus, 1761),and is near C. neviusiella Busck, 1904, a speciesknown from China and the oriental coast of Russia.

Biology. – Unknown.Distribution. – Northern Pakistan: Abbottabad

and Murree; Afghanistan: Nuristan.

Coleophora thiophaea Meyrick(figs. 5, 32-35)

Coleophora thiophaea Meyrick, 1917: 70. Lectotype ? (heredesignated), INDIA. – ‘LECTOTYPE’ (round blue-edgedBMNH label), ‘Dindigul Madras, R.M., 28. 9. 16’(Meyrick’s handwritten label), ‘thiophaea Meyr.’(Meyrick’s handwritten label), ‘Coleophora thiophaeaMeyr., 2/2, E. Meyrick det., in Meyrick Coll.’, ‘B.M. Gen-italia Slide No 24465 ?’, ‘Lectotype ?, Coleophora thio-phaea Meyr., teste K. Sattler 1988’ (BMNH) [examined].

Description. – The original description is correct:‘11 mm. Head and thorax whitish-ochreous. Palpi fer-ruginous-yellowish or yellow-whitish. Antennae stout,ochreous-whitish, basal joint shortly rough-scaled an-teriorly. Abdomen whitish-ochreous. Forewings elon-gate-lanceolate; light ochreous-yellow, deeper and fer-ruginous-tinged posteriorly; an undefined elongaterather dark fuscous patch extending along termen, suf-fused anteriorly with yellow-ferruginous: cilia greyish,on costa ochreous-yellow. Hindwings grey, tingedwith pale ochreous towards apex; cilia pale grey, oncosta tinged with ochreous.’

Male genitalia (figs. 32-35). – Spinose part ofgnathos oval, ventral margin enveloped by spinosestructure joined to gnathos arms. Tegumen mediallyconstricted, with two slightly widened arms. Transtil-la small, subtriangular, extended. Valvula large, heavi-ly sclerotized, ventral margin in the form of a teardrop,covered with bristles.; dorsal margin very thick, cover-ing cucullus margin towards ⅔ of its length. Cucullusshort, narrower at base. Sacculus narrow, long, ventralmargin curved, ending at dorsal angle in a slightlycurved subtriangular point. Phallotheca conical, long,

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more sclerotized at ventral margin, with some spinesof different lengths in distal part. Vesica short, with-out cornuti, ventrally more sclerotized.

Structure of abdominal supports (fig. 36). – Noposterior lateral struts; transverse strut slender, almoststraight. Oval tergal disks bear many conical spines.

Remark. – This species, of which the female is un-known, resembles those of the group of C. vacciniellaHerrich-Schäffer, 1861 because of the structures ofthe male genitalia, although the structure at the baseof the spinal part of the gnathos is extraordinary. Thehabitus of the species is characteristic because of theyellow patches on the forewings.

Biology. – Unknown.Distribution. – South-eastern India: Tamil Nadu,

Dindigul.

Coleophora statherota Meyrick(figs. 37-40, 42-44)

Coleophora statherota Meyrick, 1917 : 72. Lectotype ? (heredesignated), SRI LANKA. – ‘LECTOTYPE’ (round blue-edgedBMNH label), ‘Maskeliya, Ceylon, Pole, 1. 06 ‘, ‘statherotaMeyr.’ (Meyrick’s handwritten label), ‘Coleophorastatherota Meyr., 3/4, E. Meyrick det., in Meyrick coll.’,‘Meyrick Coll., B.M. 1938-290’, ‘B.M. Genitalia Slide?No 24474 ‘, ‘Lectotype ?, Coleophora statherota Meyr.,teste K. Sattler, 1988’ (BMNH) [examined].

Additional material examined. – SRI LANKA: 1/ paralecto-type, ‘Maskeliya, Ceylon, de Mowbray, 8.05’, ‘Coleophorastatherota Meyr., 2/4, E. Meyrick det., in Meyrick coll.’, slide24476 (BMNH); 1/, N.E. Distr. Hakgala Botanical Gardens,6000 feet, black light, 6-8.x.1976, G. F.Hevel, R. E.Dietz, S.Karunaratne, D. W. Balasooriya, slide Bldz 9209 (USNM).

Description. – The original description is correct:‘18-20 mm. Head whitish-ochreous, sides of crownwhitish. Palpi grey, white towards base beneath, scalesof second joint hardly projecting at apex beneath. An-tennae whitish, faintly and suffusedly ringed with fus-cous, basal joint shortly rough-scaled anteriorly. Tho-rax white, patagia more or less infuscated. Abdomenwhitish-grey. Forewings narrowly elongate-lanceolate;fuscous, suffusedly streaked with dark fuscous be-tween veins; a narrow white costal streak from base tonear apex; veins more or less marked with ill-definedirregular white lines; cilia pale greyish, at apex with asuffused white bar. Hindwings grey; cilia light grey.’

Male genitalia (figs. 37-38). – Spinose part ofgnathos large, globular. Tegumen long, medially con-stricted. Transtilla small, subtriangular. Valvula large,with rounded ventral margin. Cucullus long, club-shaped, very oblique. Sacculus narrow, long, charac-terised in the ventral angle by a long subtriangularpoint, in the dorsal angle by a long process attached tobase of cucullus. Phallotheca big, conical, almost trans-parent. 6 cornuti, in the form of spines of different

lengths.Structure of abdominal supports (figs. 39, 44). –

No lateral posterior struts; transverse strut straight,with rather reduced proximal margin. Tergal disks(3rd tergite) about 6-7 times longer than wide, withsmall conical spines.

Female genitalia (figs. 42-43). – Papillae anales large,narrow. Apophyses posteriores about 2.5 times longerthan anteriores. Sterigma subtrapezoid, membranous,sclerotized mainly anteriorly on two strips from base ofapophyses anteriores to distal margin. Ostium bursaesmall, arched, membranous. Colliculum forming along funnel. Ductus bursae narrow, with long medialline about ⅔ of its length from colliculum; then widen-ing with a convolution, ending in bursa copulatrix,which is small, oval, without signum.

Remark. – The species probably belongs to thegroup of C. conyzae Zeller, 1868 because of the struc-ture of the genitalia, but it is rather characteristic be-cause of the structure of the sacculus in the male, andof the ductus bursae and the absence of a signum inthe female.

Biology. – Unknown.Distribution. – Sri Lanka: central mountains.

Coleophora byrsostola Meyrick comb. n.(figs. 7, 45-48, 50-53 )

Macrocorystis byrsostola Meyrick, 1931: 49. Lectotype ? (heredesignated), INDIA. – ‘LECTOTYPE’ (round purple-edgedBMNH label), ‘Mahableshawar Bombay RM. Bred .6. 30’(Meyrick’s handwritten label), ‘Macrocorystis birsostola (sic)Meyr. 1/9, E. Meyrick det., in Meyrick Coll.’ ‘MeyrickColl. B.M. 1938-290’, ‘MACROCORYSTIS Meyr.’ (Meyrick’shandwritten label), ‘byrsostola’ (Meyrick’s handwritten la-bel), ‘Lectotype ?, Macrocorystis birsostola (sic) Meyr., testeK. Sattler, 1988’ (BMNH) [not examined].

Material examined. – INDIA: 1?, Mahableshwar, 8.v.30,R.Maxwell, slide 24468 (BMNH); 1/, Mahableshwar,8.vi.30, R. Maxwell, slide 24469 (BMNH); 1?, Didibi, N.Coorg, LN. vii.[19]07, [as Coleophora tarsocoma Meyr. 1/4],slide 24454 (BMNH).

Description. – The original description is correct:‘8-9 mm. Head, antennae whitish-ochreous. Palpi,thorax greyish-ochreous. Forewings elongate-lanceo-late; greyish-ochreous; second discal stigma dark fus-cous: cilia whitish-grey-ochreous. Hindwings lightgrey; cilia whitish-grey-ochreous.’

Male genitalia (fig. 45-48). – Spinose part ofgnathos globular. Tegumen medially constricted.Transtilla slender, long, medially joined. Valvula big,covered with bristles, rounded on the ventral margin.Cucullus short, stout, with broad base. Sacculusrather small, on the lateral margin ending in a moreor less long and rounded process. Phallotheca short,conical, more sclerotized on the ventral margin, end-

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ing in a beak-shaped point. Numerous cornuti, in theform of spines of different lengths.

Structure of abdominal supports (figs. 49, 53). –No posterior lateral struts; transverse strut slender,convex in the form of an obtuse angle. Tergal disks,with many conical spines, about 3 times longer thanwide (3rd tergite).

Female genitalia (figs. 50-52). – Papillae analeslarge, oval. Apophyses posteriores about half length ofanteriores. Sterigma subtrapezoidal, rather wide andshort. Ostium bursae arched. Colliculum beaker-shaped. Ductus bursae with a spinose, coiled sleeveonly in a short distal section; remainder of ductustransparent, slightly more sclerotized near the inser-tion of the ductus seminalis. Bursa copulatrix round-ed, with a big anchor-shaped signum.

Remark. – It is difficult to place the species in agroup of Toll’s system. It does not resemble anypalaearctic species, but it has links with someafrotropical species, which are still being studied.

Biology. – According to the original descriptionthe species was bred from leaves of Strobilanthes(Acanthaceae), ‘from larvae mining blotches in De-cember in leaf of Strobilanthes (Acanthaceae), severalin one leaf; pupa within blotch, forming a compactoval opaque capsule, which usually drops out as themined portion of leaf withers.’

Distribution. – Western India: Maharashstra, Ma-hableshwar; south-western India: Karnataka, Coorgplateau.

Coleophora centrota Meyrick(fig. 8)

Coleophora centrota Meyrick, 1917:71. Holotype ?, INDIA.– ‘HOLOTYPE’ (round purple-edged BMNH label), ‘Dibidi,N. Coorg., LN. 11. 13’, ‘centrota Meyr.’, (Meyrick’shandwritten label), ‘Coleophora centrota Meyr., 1/1, E.Meyrick det., in Meyrick coll.’, ‘Meyrick Coll., BM1938-290’, ‘Abdomen Missing’, (BMNH) [examined].

Description. – The original description is correct: ‘9mm. Head, palpi, antennae and thorax whitish-ochre-ous, antennae simple. Forewings lanceolate; paleochreous, slightly tinged with grey posteriorly; seconddiscal stigma rather large, blackish; cilia whitish-ochre-ous. Hindwings and cilia whitish-grey-ochreous.’.

Remark. –The holotype, the only specimenknown, has no abdomen. Therefore it is not possibleto give a diagnosis of this species. According to the ex-ternal morphology it is probably the same as C. byr-sostola.

Distribution. – South-western India: Karnataka,Coorg plateau.

Coleophora argopleura Meyrick(figs. 9, 54-57, 58-60)

Coleophora argopleura Meyrick, 1917:71. Holotype /, IN-DIA. – ‘HOLOTYPE’ (round purple-edged BMNH label),‘Abbottabad, N.W. India, TBF. 10. 6. 16’, ‘argopleuraMeyr.’ (Meyrick’s handwritten label), ‘Coleophora argo-pleura Meyr., 1/1, E. Meyrick det., in Meyrick coll.’,‘Meyrick Coll., BM 1938-290’, ‘B.M. Genitalia Slide /24451’, ‘Holotype /, Coleophora argopleura Meyr., testeK. Sattler, 1987’ (BMNH) [examined].

Additional material examined. – AFGHANISTAN: 3?, 25km N Barikot, 1800 m, Nuristan, 12-17.vii.1963, Kasy &Vartian, slides Bldz 7915, 12874 (coll. Baldizzone). – INDIA:1?, Kashmir, 10 km N Srinagar, 11.-20.vii.1982, M. & E.Arenberger, slide Bldz 9700, 1/, idem, slide Bldz 7897(coll. Baldizzone).

Description. – The original description is correct:‘18 mm. Head and thorax white. Palpi white, secondjoint with scales hardly projecting at apex beneath.Antennae white, basal joint roughly tufted anteriorlywith long scales, stalk clothed with long rough scalesabove towards base. Abdomen ochreous-whitish.Forewings lanceolate; ochreous-yellow, towards costasuffused with fuscous; a strong white costal streakfrom base to 5/6, extreme costal edge dark grey to-wards base; dorsum suffused with white anteriorly,dorsal edge white to beyond middle; a fine white linealong fold from base to middle of wing: cilia paleochreous. Hindwings dark grey; cilia light greyish-ochreous.’

Male genitalia (fig. 54-56). – Spinose part ofgnathos globular. Tegumen constricted at base ofgnathos arms. Transtilla rather small, subtriangular.Valvula with rounded ventral margin; dorsal marginwith short sclerotized bristle. Cucullus rather long,club-shaped, considerably constricted at base. Saccu-lus small, simple, with curved ventral margin, endingin triangular point on lateral margin. Phallothecaconical, dorsally partially sclerotized. Cornuti 5-6,stout spines, reunited into a chain.

Structure of abdominal supports (figs. 57, 60). –Posterior lateral struts about 2.5-3.0 times shorterthan anteriores. Transverse strut slightly convex, slen-der. Tergal disks, with many short conical spines,about twice longer than wide (3rd tergite).

Female genitalia (figs. 58-59). – Papillae analeslarge, oval. Apophyses posteriores about 2.5 timeslonger than anteriores. Sterigma subtriangular, itsproximal margin rather wide, distal margin long andmedially, near the arched ostium bursae, very spinose.Colliculum beaker-shaped. Ductus bursae ratherlong, a medial line occupying about 4/5 of its length,the distal spinose sleeve occupying 3/5. Bursa copula-trix ample, transparent, with leaf-shaped signum.

Remark. – The species belongs to the 18th group ofToll’s system; the genitalia do not allow a more de-

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tailed positioning.Biology. – Unknown.Distribution. – Northern Pakistan, Kashmir,

north-eastern Afghanistan.

Coleophora castalia Meyrick(figs. 10, 61-64, 65-67)

Coleophora castalia Meyrick, 1930:561. Holotype ?, INDIA.– ‘HOLOTYPE’ (round purple-edged BMNH label), ‘Yusi-marg, Kashmir, 7500’, TBF. 8. 23’, ‘castalia Meyr.’(Meyrick’s handwritten label), ‘Coleophora castalia Meyr.,1/1, E. Meyrick det., in Meyrick coll.’, ‘Meyrick Coll.,BM 1938-290’, ‘B.M. Genitalia Slide ? 24440’, ‘Holo-type ?, Coleophora argopleura Meyr., teste K. Sattler,1986’ (BMNH) [examined].

Additional material examined. – AFGHANISTAN: 1/, 25km N Barikot, 1800 m, Nuristan, 12-17.vii.1963, Kasy &Vartian, slide Bldz 7810 (coll. Baldizzone). – INDIA: 1/, As-sam, Khasi-Hills, vii.1977, Vollmer, slide Bldz 4596 (coll.Baldizzone); 1?, 1/, Kashmir, 15 km W Kangan, Wangan,16-17.vii.1982, M.& E. Arenberger, slides Bldz 7895, 7896(coll. Baldizzone). – PAKISTAN: 11/, Prov. Swat, Madyan,71°90’-35°70’, 1400 m, 19.vi.-4.vii.1971, Vartian, slideBldz 12916 (coll. Baldizzone).

Description. – The original description, based on asingle specimen, is correct: ‘18 mm. Head ochreous-white. Palpi whitish. Antennae whitish ringed grey-ish-ochreous, scape with rough projecting hairscalesanteriorly, basal sixth of stalk thickened with loosescales above. Forewings brownish, dorsal half verypale ochreous; a moderate attenuated white costalstreak from base to ¾; streaks of white suffusion con-fluent towards base along dorsum to middle of wing,and along fold; a slender streak of white suffusion be-neath brown area from middle of disc to beneathapex, broader posteriorly: cilia white, towards tornustinged pale ochreous. Hindwings grey; cilia ochreous-grey-whitish.’ It should be noted that there is a cer-tain variation in colour, more or less fuscous, accord-ing to the locality. The same holds good for the whitecostal line, which is more or less wide, and purewhite, or slightly suffused with brown.

Male genitalia (fig. 61-63). – Spinose part ofgnathos big, pear-shaped. Tegumen long, constrictedat gnathos base. Transtilla small, subtriangular. Valvu-la wide, rounded, covered with bristles. Sacculusshort, rounded and sclerotized on lateral margin, asmall triangular tooth on dorsal angle. Phallothecaconical, long, partially sclerotized on dorsal and later-al sides. Numerous cornuti, as small spines, most ofthem reunited into long strip.

Structure of abdominal supports (figs. 64, 67). –Posterior lateral struts, poorly indicated, are about halflength of anteriores. The almost straight transversestrut slender. Tergal disks, covered with strong conical

spines, about twice longer than wide (3rd tergite).Female genitalia (figs. 65-66). – Papillae anales

oval. Apophyses posteriores about 3 times length ofthe anteriores. Sterigma subtrapezoidal, sclerotizedmainly in central anterior part. Ostium bursae arched.Colliculum beaker-shaped. Ductus bursae with medi-al line over 4/5 of its length, covered by a sleeve withsmall spines over half its length; ductus transparent,with central convolutions ending in bursa copulatrix,which is round, with large leaf-shaped signum.

Remark. – The species belongs to the 18th group ofToll’s system but is difficult to place near any otherspecies because of the shape of the sacculus in themale, and of the sterigma and ductus bursae in the fe-male.

Biology. – Unknown.Distribution. – North-western Afghanistan, north-

ern Pakistan, Kashmir, north-eastern India: Assam,Khasi-Jaintia Hills.

Coleophora versurella Zeller

Coleophora versurella Zeller, 1849: 352.Coleophora enchorda Meyrick, 1931:180. Holotype /, IN-

DIA. – ‘HOLOTYPE’ (round purple-edged BMNH label),‘Pusa, Bihar, TBF, bred 9. 17’, ‘enchorda Meyr.’(Meyrick’s handwritten label), ‘Coleophora enchordaMeyr. 1/1, E. Meyrick det., in Meyrick coll.’, ‘B.M. Gen-italia Slide / 24444’, ‘Holotype /, Coleophora enchordaMeyr., teste K. Sattler, 1986’ (BMNH) [examined]. syn. n.

Remark. – The structure of the genitalia of theholotype of C. enchorda, shows that this is a juniorsynonym of Coleophora versurella Zeller, 1849.

Biology. – According to Meyrick’s description thespecimen was bred from Amaranthus. In fact, the lar-va of C. versurella lives on several Amaranthaceae.

Distribution. – A species with a Holarctic distribu-tion. Also known from South America.

Coleophora aphrocrossa Meyrick(figs. 11, 68-70)

Coleophora aphrocrossa Meyrick, 1933:364. Holotype /, IN-DIA. – ‘HOLOTYPE’ (round purple-edged BMNH label),‘Gulmarg, Kashmir, TBF. 8800’ J. 31’, ‘aphrocrossa Meyr.’(Meyrick’s handwritten label), ‘Coleophora aphrocrossaMeyr., 1/1, E. Meyrick det., in Meyrick coll.’, ‘MeyrickColl., BM 1938-290’, ‘B.M. Genitalia Slide /24441’,‘Holotype /, Coleophora aphrocrossa Meyr., teste K. Sat-tler, 1986’ (BMNH) [examined].

Description. – The original description is correct:‘12 mm. Head light greyish, crown fuscous exceptsides. Palpi white, oblique apical half of second jointgrey becoming fuscous anteriorly, with slight apicalprojection. Antennae white ringed dark fuscous,scape loosely scaled. Thorax grey. Forewings grey;

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costal edge ochreous-white from near base to apex:cilia whitish-grey, on costa ochreous-whitish. Hind-wings light grey; cilia whitish-grey.’

Female genitalia (figs. 68-69). – Papillae analeslong, very slender. Apophyses posteriores about 5times longer than anteriores. Sterigma subtrape-zoidal, excavated in distal margin near ostium bursae,which is arched. Colliculum narrow, long, rather scle-rotized. Ductus bursae with medial line in distal half,which is covered by sleeve with small spines over ⅔ ofits length; proximal half of ductus almost transparent,with speckled sclerotization. Bursa copulatrix ample,with small leaf-shaped signum.

Structure of abdominal supports (fig. 70). – Noposterior lateral struts; transverse strut slightly con-vex. Tergal disks, with small conical spines, about 1,5times longer than wide (3rd tergite).

Remark. – The species, of which the male is un-known, belongs to the 30th group of Toll’s system,and can be placed in the section of C. versurellaZeller, 1849.

Biology. – Unknown.Distribution. – Western Kashmir: Gulmarg.

Coleophora echyropis Meyrick(fig. 6)

Coleophora echyropis Meyrick, 1930 : 562. Holotype ?, IN-DIA. – ‘HOLOTYPE’ (round purple-edged BMNH label),‘Yusimarg, Kashmir, TBF, 7500’, 8.’, ‘echyropis Meyr.’(Meyrick’s handwritten label), ‘Coleophora echyropisMeyr., 1/1, E. Meyrick det., in Meyrick coll.’, ‘HolotypeColeophora echyropis Meyr., teste K. Sattler 1988’, ‘Ab-domen missing’ (BMNH) [examined].

Original description: ‘?, 14 mm. (Head and tho-rax discoloured.) Antennae white ringed fuscous,scape rather rough-scaled anteriorly. Palpi white, sec-ond joint with a fuscous lateral streak, scales shortlyprojecting at apex, terminal joint ⅔ of second.Forewings light greyish-ochreous; a broad whitestreak with two or three black specks along costa frombase to ⅔, suffused posteriorly; apical third of wingsuffusedly mixed white, with a very few black specksbetween veins; a slender whitish-grey streak in poste-rior half of cell, with a few black scales; a greyish linewith some black scales along fold: cilia whitish-greymixed white at base, on costa white. Hindwings grey;cilia pale grey.’

Remark. – The holotype is in bad condition: theright wings and the abdomen are missing. Thereforeit is impossible to identify the specimen. Judgingfrom the remaining forewing, the species might be-long to the group of C. vestianella (Linnaeus, 1758).

Biology. – Unknown.Distribution. – Kashmir.

Coleophora ogmotona Meyrick(figs. 15, 71-74)

Coleophora ogmotona Meyrick, 1917: 72. Holotype ?, SRI

LANKA. – ‘HOLOTYPE’ (round purple-edged BMNH label),‘Puttalam, Ceylon, Pole, .6. 04’, ‘ogmotona Meyr.’(Meyrick’s handwritten label), ‘Coleophora ogmotonaMeyr., 3/4, E. Meyrick det., in Meyrick coll.’, ‘B.M. Geni-talia Slide ? No 24467’, ‘Holotype ?, Coleophora ogmo-tona Meyr., teste K. Sattler, 1988’ (BMNH) [examined].

Additional material examined. – INDIA: 1?, Kanara,Dharwar, R.M.[axwell] 1.ii.[19]16, slide 24471; 1?, Dhar-war, (?)Annimbhari, R. Maxwell, slide 24470(BMNH).

Description. – The original description is correct: ‘8mm. Head and palpi fuscous-whitish. Antennae stouttowards base, grey obscurely ringed with whitish.Thorax fuscous, with four whitish longitudinal lines.Abdomen grey. Forewings elongate-lanceolate; fus-cous, obscurely darker-sprinkled; veins marked byslender whitish lines: cilia light grey. Hindwings grey;cilia light grey.’

Male genitalia (figs. 71-73). – Globular part ofgnathos small, rounded. Tegumen small, mediallywith an Y-shaped reinforcement. Transtilla small,subtriangular. Valvula small, well sclerotized, extend-ed on the ventral margin. Cucullus long, apicallyslightly wider than basally. Sacculus with complexstructure: from the ventral margin rises a long processin the form of an acute horn, reaching base of dorsalangle; a short, curved protuberance, on lateral marginnear ventral angle, and a slightly longer one on dorsalangle. Phallotheca with two sclerotized, almost sym-metrical rods, apically with curved, acute point. Cor-nutus small, like a spine.

Structure of abdominal supports (fig. 74). – Noposterior lateral struts; transverse strut straight, moresclerotized on proximal margin. Tergal disks, withsmall conical spines, about twice longer than wide(3rd tergite)

Remark. – The species, of which the female is un-known, belongs to the 30th group of Toll’s system.The structure of the sacculus resembles more or lessthat of C. gymnocarpella Walsingham, 1907, whosegenitalia were illustrated by Baldizzone (1994).

Biology. – Unknown.Distribution. – South-western India: Dharwar

plateau; Sri-Lanka: western region, Puttalam.

Coleophora percnoceros Meyrick(figs. 12, 75-78)

Coleophora percnoceros Meyrick, 1933: 365. Holotype ?, IN-DIA. – ‘HOLOTYPE’ (round purple-edged BMNH label),‘Killanmarg, Kashmir, 10500’ TBF, 7. 31’, ‘percnocerosMeyr.’ (Meyrick’s handwritten label), ‘Coleophora percno-ceros Meyr., 1/1, E. Meyrick det., in Meyrick coll.’,

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‘Meyrick Coll., B.M. 1938-290’, ‘B.M. Genitalia Slide ?No 24439 ‘, ‘Holotype ?, Coleophora percnoceros Meyr.,teste K. Sattler, 1986’ (BMNH) [examined].

Description. – The original description is correct:‘12 mm. Head, thorax light grey slightly mixed white.Palpi without projection, grey, internally white. An-tennae dark grey, scape smooth. Forewings light ash-grey; costal edge suffused whitish from base to beyondmiddle; veins indicated by faint streaks of whitish suf-fusion: cilia pale grey, on costa whitish except nearapex of wing. Hindwings light grey; cilia pale grey.’

Male genitalia (figs. 75-77). – Spinose part ofgnathos suboval. Tegumen with two long, externallydilated arms. Transtilla slender, long. Valvula round-ed on ventral margin. Cucullus compact, ear-shaped.Sacculus big, narrow, characterised by small triangu-lar tooth in ventral angle, a curved one in dorsal an-gle, and a smaller one on lateral margin halfway be-tween preceding ones. Phallotheca with two robustrods, the longer terminating in a conical point, theother with small tooth dorsally on apex. Two smallcornuti, basally joined, forming a spine.

Structure of abdominal supports (fig. 78). – Noposterior lateral struts; transverse strut straight, wide,with thicker proximal margin, distal margin moreslender and convex. Tergal disks, with small conicalspines, about 5 times longer than wide (3rd tergite).

Remark. – The species, of which the female is un-known, belongs to the 30th group of Toll’s system,and resembles C. leonensis Baldizzone & van derWolf, 2000, although the structure of the sacculus isdifferent: longer on the lateral margin.

Biology. – Unknown.Distribution.- Kashmir.

Coleophora niphomesta Meyrick(figs. 14, 79-83, 84-86)

Coleophora niphomesta Meyrick, 1917 : 73. Holotype /, IN-DIA. – ‘HOLOTYPE’ (round purple-edged BMNH label), ‘Pe-shawar, N. W. India, TBF. 5. 16’, ‘niphomesta Meyr.’(Meyrick’s handwritten label), ‘Coleophora niphomestaMeyr. 1/1, E. Meyrick det., in Meyrick coll.’, ‘B.M. Gen-italia Slide /24453’, ‘Holotype /, Coleophora niphomes-ta Meyr., teste K. Sattler, 1987’ (BMNH) [examined].

Coleophora sarobiensis Toll & Amsel, 1967: 14. syn. n.

Description. – The very detailed original descrip-tion is based on a single specimen : ‘11 mm. Headand thorax white. Palpi white, externally with a greyline, scales hardly projecting at apex of second jointbeneath. Antennae white ringed with pale fuscous,basal joint somewhat roughened with scales anterior-ly. Abdomen whitish-grey. Posterior tibiae white witha fuscous longitudinal line. Forewings lanceolate;whitish-ochreous; a white streak along costa frombase to ⅔, one suffused along dorsum from base to ⅔,

and one along fold throughout; two or three suffusedwhite streaks on veins towards costa posteriorly, andone from middle of disc somewhat downward-sinu-ate to termen beneath apex: cilia pale greyish, withwhite scales at base. Hindwings pale grey; cilia ochre-ous-whitish-grey.’ When studying several specimensfrom Iran and Afghanistan we noticed that some-times the forewings have no white lines along theveins, but are almost unicolorous, white dirty-ochre-ous sprinkled with some darker scales.

Male genitalia (figs. 79-82). – Spinose part ofgnathos small, globular. Tegumen with two longarms. Transtilla slender, extended. Valvula small,sclerotized, with rounded ventral margin. Cucullusclub-shaped. Sacculus characterised by large roundedprotuberance in ventral angle, and in dorsal angle abig curved horn, with triangular tooth on inner side.Phallotheca with two curved almost symmetricalrods, with a tooth on dorsal side in variable positionaccording to the specimen, normally at ⅓ beforeapex; another smaller subtriangular more or less acutetooth on apex. Cornuti rather small, 2-3, joined intospine-like formation.

Structure of abdominal supports (fig. 83, 86). – Noposterior lateral struts; transverse strut almost straight,with two clear margins in male, and merged ones in fe-male. Tergal disks, covered with small conical spines,2.5 times longer than wide in male, and 3.5 times infemale (3rd tergite).

Female genitalia (figs. 84-85). – Papillae analesoval, long. Apophyses posteriores about twice lengthof anteriores. Sterigma subtrapezoidal, with roundeddistal margin. Ostium bursae arched. Colliculum awidened beaker. Ductus bursae with spinose sleeve indistal section, which is about ⅛ of its total length;other part of ductus transparent, ending in bag-shaped bursa copulatrix containing a leaf-likesignum.

Remark. – The species belongs to the 30th group ofToll’s system, near C. beduina Baldizzone, 1987.

Biology. – The larva lives on Aerva persica (Ama-ranthaceae), the plant on which F. Kasy found it inIran (Baldizzone 1994: 173).

Distribution. – Turkmenistan, Iran, Pakistan,Afghanistan.

Coleophora chordoscelis Meyrick(figs. 13, 87-89)

Coleophora chordoscelis Meyrick, 1917:73. Lectotype /(here designated), INDIA. – ‘LECTOTYPE’ (round blue-edged BMNH label), ‘Dindigul, Madras, RM. 28.9.16’,‘chordoscelis Meyr.’ (Meyrick’s handwritten label),‘Coleophora chordoscelis Meyr., 1/4, E. Meyrick det., inMeyrick coll.’, ‘Meyrick Coll., BM 1938-290’, ‘B.M.Genitalia Slide / 24474’, ‘Lectotype /, Coleophora chor-doscelis Meyr., teste K. Sattler, 1988’ (BMNH) [examined].

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Description. – The original description is correct:‘?, /. 12 mm. Head and thorax glossy light grey, ex-treme lateral edge of crown whitish. Palpi smooth,light grey, edges white. Antennae stout towards base,white suffusedly ringed with grey. Abdomen light grey.Posterior tibiae white with a fine median longitudinalfuscous line. Forewings narrow-lanceolate; glossy lightgreyish-ochreous suffused with light grey; costal edgewhite from base to about ¾: cilia pale greyish, on costawhitish. Hindwings and cilia light grey.’

Female genitalia (figs. 87-88). – Papillae anales nar-row, long. Apophyses posteriores about twice lengthof anteriores. Sterigma subtrapezoidal, very simple.Ostium bursae large, arched. Colliculum a longbeaker. Ductus bursae, about 4 times longer thanlength of sterigma, totally transparent, with speckledsclerotization on two small coils joining bursa copula-trix, which is oval, with horn-shaped signum.

Structure of abdominal supports (fig. 89). – Noposterior lateral struts; transverse strut slender, slightlyconvex. Tergal disks, with some small conical spines,8-9 times longer than wide (3rd tergite).

Remark. – The species, whose male is unknown,belongs to the 30th group of Toll’s system, but thestructure of the female genitalia does not allow a moreprecise position near other species. It should be notedthat Meyrick in his description mentions four speci-mens of both sexes, while in the BMNH collectionthere are only females.

Biology. – Unknown.Distribution. – South-eastern India: Tamil Nadu,

Madras.

Coleophora leucochares Meyrick(fig. 41)

Coleophora leucochares Meyrick, 1922:556. Holotype ?, IN-DIA. – ‘HOLOTYPE’ (round purple-edged BMNH label),‘Shillong, Assam, TBF, 9. 20’, ‘leucochares Meyr.’(Meyrick’s handwritten label), ‘Coleophora leucocharesMeyr. 2/2, E. Meyrick det., in Meyrick coll.’, ‘Abdomenmissing’, ‘Holotype ?, Coleophora leucochares Meyr.,teste K. Sattler, 1988’ (BMNH) [examined].

Description. – The very simple original descriptioncorresponds well with the only specimen known: ‘10mm. Head, palpi, antennae, thorax white. Forewingsnarrow, acute-pointed; shining white: cilia white.Hindwings pale grey; cilia grey-whitish.’

Remark. – The specimen on which the descriptionis based is in poor condition and the abdomen ismissing. For this reason it is not possible to give a pre-cise diagnosis.

Biology. – Unknown.Distribution. – North-eastern India: Assam, Shil-

long.

SPECIES EXCLUDED FROM COLEOPHORIDAE

‘Coleophora’ aphypnota Meyrick(figs. 1, 17-18)

Coleophora aphypnota Meyrick,1917:73 Lectotype ? (heredesignated), SRI LANKA. – ‘LECTOTYPE’ (round blue-edgedBMNH label), ‘Puttalam, Ceylon, Pole, 8. 04’, ‘aphypnotaMeyr.’ (Meyrick’s handwritten label), ‘Coleophora aphyp-nota Meyr., 1/4, E. Meyrick det., in Meyrick coll.’,‘Meyrick Coll., BM 1938-290’, ‘B.M. Genitalia Slide ?24473’, ‘Lectotype ?, Coleophora aphypnota Meyr., testeK. Sattler, 1988’ (BMNH) [examined].

Description. – The material examined correspondswell with the original description.

Male genitalia (fig. 17). Structure of abdominal supports (fig. 18).Remark. – This species does not belong to

Coleophoridae, but to Batrachedridae, as shown bythe male genitalia (S. Yu. Sinev in litt.). The female isnot known.

Biology. – Unknown.Distribution. – Sri Lanka: western region, Puttalam.

‘Coleophora’ crossophanes Meyrick(figs. 2, 19-20)

Coleophora crossophanes Meyrick, 1917: 70. Lectotype ? (heredesignated), SRI LANKA. – ‘LECTOTYPE’ (round blue-edgedBMNH label), Maskeliya, Ceylon, Pole.10.05’; ‘Coleophoracrossophanes Meyr., 2/3, E. Meyrick det., in MeyrickColl.’; ‘Meyrick Coll., B.M. 1938-290.’; ‘crossophanesMeyr.’ (Meyrick’s handwritten label) [not examined].

Additional material examined. – SRI LANKA: 1?, paralec-totype ‘Maskeliya, Ceylon, Pole, 5. 06’, ‘Coleophora crosso-phanes Meyr. ?, 3/3, E. Meyrick det., in Meyrick coll.’,‘Meyrick coll., BM. 1938-290’, slide 24479 (BMNH)

Description. – The original description corre-sponds well with the specimen examined.

Male genitalia (fig. 19). Structure of abdominal supports (fig. 20).Remark. – It should be noted that among the three

original specimens there are only males and no females,as indicated by Meyrick in his description. Based onexternal morphology and male genitalia it is clear thatthis species does not belong to Coleophoridae. It couldbelong to the Momphidae (K. Sattler in litt.).

Biology. – Unknown.Distribution. – Sri Lanka: Maskeliya.

‘Coleophora’ tarsocoma Meyrick(fig. 16)

Coleophora tarsocoma Meyrick, 1917:71. Holotype ?, IN-DIA. – ‘HOLOTYPE’ (round purple-edged BMNH label),‘Khasi Hills, Assam, – 4. 1907’, ‘tarsocoma Meyr.’(Meyrick’s handwritten label), ‘Coleophora tarsocoma

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Meyr., 3/4, E. Meyrick, in Meyrick coll.’, ‘Holotype ?,Coleophora tarsocoma Meyr., teste K. Sattler, 1987’, ‘Ab-domen missing’ (BMNH) [examined].

Description. – The original description is correct.Remark. – The holotype is the only specimen in

the BMNH. A second specimen labelled ‘Coleophoratarsocoma Meyr., 1/4, E. Meyrick, in Meyrick coll.’ isactually Coleophora byrsostola (Meyrick, 1931) [seethere]. Based on the external morphology of the holo-type it can be said that tarsocoma does not belong tothe Coleophoridae. According to K. Sattler (in litt.) itcould belong to the Cosmopterigidae.

Biology. – Unkown.Distribution. – North-eastern India: Assam, Khasi-

Jaintia Hills, near Shillong.

CONCLUSIONS

The study of the species of the Indian region, al-though very interesting, does not yield a great amountof information, mainly because of the small number ofspecimens known. If one tries to separate the speciesaccording to zoogeographical regions, it is clear that allspecies of the northern sector (C. aegra, C. argopleura,C. castalia, C. versurella, C. aphrocrossa, C. echyropis, C.percnoceros, C. niphomesta, C. leucochares) correspondmore or less with the palaearctic groups (followingToll’s classification (1953, 1962, pending a modern re-vision). A certain number of new species from Nepal,Kashmir and Northern India are still to be added toMeyrick’s material; they all belong to the palaearcticfauna. Among them is also a species close to C. argo-pleura, a noteworthy species, which belongs to a groupwhich appears to be characteristic for that region.

The species of the southern region and of Sri-Lanka(C. crypsiphanes, C. thiophaea, C. statherota, C. byrsos-tola, C. centrota, C. ogmotona, C. chordoscelis) are moredifficult to interpret. C. thiophaea could belong to thegroup of C. vacciniella Herrich-Schäffer, although itshabitus is rather peculiar and its affinities are not fullyunderstood, as the female is not known. As to C.statherota, the male resembles species of the C. conyzaeZeller group, but the female, without signum, is near-er to some species of South Africa. The other species,particularly C. crypsiphanes, have no palaearctic equiv-alents and resemble some afrotropical species, most ofwhich still await description. It is therefore essentialthat many more specimens should be studied beforewe can gather a more satisfactory insight in the Indianfauna of Coleophoridae.

ACKNOWLEDGEMENTS

We are indebted to Dr. Klaus Sattler and Mr.Kevin Tuck (BMNH), who have entrusted us with

Meyrick’s type material, and who have generously as-sisted us with photocopies, information and advice;and to Dr. S. Yu. Sinev (St. Petersburg) for informa-tion on ‘Coleophora’ aphypnota. We also express ourgratitude to Dr. Don Davis of USNM for material hehas entrusted us with. Also a word of thanks to ananonymous referee for advice resulting in improve-ments in this publication.

REFERENCES

Baldizzone, G., 1987. Contributions à la connaissance desColeophoridae, XLIII. Sur quelques espèces deColeophoridae de Libye. (Lepidoptera). – Alexanor l5(1): 3-16, 29 figs. (Suppl.).

Baldizzone G., 1994. Contribuzioni alla conoscenza deiColeophoridae. LXXV. Coleophoridae dell’Area Irano-Anatolica e regioni limitrofe (Lepidoptera). –AssociazioneNaturalistica Piemontese Memorie 3: 1-424, 699 figs.

Baldizzone G., & H.W. van der Wolf, 2000. Correctionsand additions to the checklist of European Coleophori-dae (Lepidoptera: Coleophoridae). – SHILAP Revista delepidopterología 28 (112): 395-428, 51 figs.

Hübner, J. G., 1796-[1836]. Sammlung europäischerSchmetterlinge, 8. Tineae. – Augsburg. pls. 1-71, 78 pp.

Hübner, J. G., 1822. Systematisch-alphabetisches Verzeich-niss aller bisher bey den Fürbildungen zur Sammlung eu-ropäischer Schmetterlinge angegebene Gattungsbenen-nungen. – Augsburg, vi+81 pp.

Meyrick, E., 1917. [no title.] – Exotic Microlepidoptera 2(2/3): 33-96.

Meyrick, E., 1922. [no title.] – Exotic Microlepidoptera 2(16/19): 481-608.

Meyrick, E., 1930. [no title.] – Exotic Microlepidoptera 3(18/20): 545-640.

Meyrick, E., 1931 [no title.] – Exotic Microlepidoptera 4(2/6): 33-192.

Meyrick, E., 1933 [no title.] – Exotic Microlepidoptera 4(12/14): 353 448.

Toll, S., [1953]. Rodzina Eupistidae (Coleophoridae) Pols-ki. – Materialy do fizjografii kraju 32: 1-292, 38 pls.

Toll, S., 1962. Materialien zur Kenntnis der paläarktischenArten der Familie Coleophoridae (Lepidoptera). – Actazoologica cracoviensia 7 (16): 577-720, 133 pls.

Toll, S., & Amsel, H. G., 1967. Coleophoriden ausAfghanistan (Lepidoptera: Coleophoridae). – Beiträgezur naturkundlichen Forschung in Südwestdeutschland,26 (3): 5-16, 5 pls.

Vives Moreno, A., 1988. Catalogo mundial sistematico y dedistribucion de la familia Coleophoridae Hübner, [1825](Insecta, Lepidoptera). – Boletin de Sanidad Vegetal 12:1-196.

Zeller, P. C., 1849. Beitrag zur Kenntnis der Coleophoren.– Linnaea Entomologica 4: 191- 416.

Received: 2 March 2002Accepted: 31 May 2002

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Figs. 1-2. Males of ‘Coleophora’ spp., now excluded from Coleophoridae. – 1, C. aphypnota, lectotype; 2, C. crossophanes, para-lectotype. – Figs. 3-8. Males of Coleophora spp. 3, C. crypsiphanes, lectotype; 4, C. aegra, Afghanistan, Barikot; 5, C. thiophaea,lectotype; 6, C. echyropis, holotype; 7, C. byrsostola, Mahableshawar, 8.5.30; 8, C. centrota, holotype.

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1 2

3 4

5 6

7 8

Figs. 9-15. Adults of Coleophora spp. 9, C. argopleura, ?, Afghanistan, Barikot; 10, C. castalia, ?, Kashmir; 11, C. aphro-crossa, holotype /, 12, C. percnoceros, holotype ?, 13, C. chordoscelis, lectotype /, 14, C. niphomesta, ?, Iran, Bandar-Ab-bas; 15, C. ogmotona, holotype ?. – Fig. 16 ‘Coleophora’ tarsocoma, holotype ?, excluded from Coleophoridae (photo BMNH).

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9 10

11 12

13 14

15 16

Figs. 17-18. ‘Coleophora’ aphypnota, lectotype, slide BMNH 24473. – 17, Male genitalia; 18, abdomen. – Figs. 19-20.‘Coleophora’ crossophanes, paralectotype, slide BMNH 24479. – 19, Male genitalia; 20, abdomen.

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17 19

18 20

Figs. 21-24. Coleophora crypsiphanes, lectotype, slide BMNH 24477. – 21, Male genitalia; 22, detail of genitalia at high magni-fication; 23, abdomen; 24, labels.

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21 22

23 24

Figs. 25-27. Coleophora crypsiphanes, paralectotype, slide BMNH 24478. – 25, Female genitalia; 26, sterigma at high magnifi-cation; 27, abdomen.

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26

25 27

Figs. 28-31. Coleophora aegra, lectotype, slide BMNH 24452. – 28, Male genitalia; 29, detail of genitalia at high magnification;30, detail, slide Bldz 8180; 31, abdomen.

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28 29

31 30

Figs. 32-36. Coleophora thiophaea, lectotype, slide BMNH 24465. – 32, Male genitalia; 33, detail of genitalia at high magnifi-cation; 34, detail of distal part of gnathos at high magnification; 35, detail of distal part of phallotheca at high magnification;36, abdomen.

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32 33

34

36 35

Figs. 37-39. Coleophora statherota, lectotype, slide BMNH 24475. – 37, Male genitalia; 38, detail of genitalia at high magnifica-tion; 39, abdomen. – Figs. 40-41. Coleophora spp. 40, C. statherota Meyrick, lectotype; 41, C. leucochares Meyrick, holotype.

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37 38

39 41

40

Figs. 42-44. Coleophora statherota, paralectotype, slide BMNH 24476, – 42, Female genitalia; 43, sterigma at high magnifica-tion; 44, abdomen.

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43

42 44

Figs. 45-48. Coleophora byrsostola, slide BMNH 24454. – 45 Male genitalia; 46, detail of genitalia at high magnification; 47,detail, slide BMNH 24456; 48, cornuti at high magnification, slide BMNH 24456; 49, abdomen, slide BMNH 24454.

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45

49

46

48

47

Figs. 50-53. Coleophora byrsostola, slide BMNH 24469. – 50; Female genitalia; 51, sterigma at high magnification; 52, slideBMNH 24455; 53, abdomen.

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50 51

53 52

Figs. 54-57. Coleophora argopleura, slide Bldz 12874. – 54, Male genitalia; 55, detail of genitalia at high magnification; 56,cornuti at high magnification; 57, abdomen.

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54

55

56

57

Figs. 58-60. Coleophora argopleura, slide BMNH 24451, holotype. – 58, Female genitalia; 59, sterigma at high magnification;60, abdomen.

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59

58 60

Figs. 61-64. Coleophora castalia, slide Bldz 7895. – 61, Male genitalia; 62, detail of genitalia at high magnification; 63, cor-nuti at high magnification; 64, abdomen.

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61

62

64 63

Figs. 65-67. Coleophora castalia, slide Bldz 12916. –65, Female genitalia; 66, sterigma at high magnification; 67, abdomen.

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66

65 67

Figs. 68-70. Coleophora aphrocrossa, slide BMNH 24441, holotype. – 68, Female genitalia; 69, sterigma at high magnification;70, abdomen.

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69

68 70

Figs. 71-74. Coleophora ogmotona, slide BMNH 24470. – 71, Male genitalia; 72, detail of genitalia at high magnification; 73,cornutus at high magnification; 74, abdomen.

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71 72

74 73

Figs. 75-78. Coleophora percnoceros, slide BMNH 24439, holotype. – 75, Male genitalia; 76, detail of genitalia at high magni-fication; 77, cornuti at high magnification; 78, abdomen.

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75 76

78 77

Figs. 79-83. Coleophora niphomesta, slide Bldz 8696. – 79, Male genitalia; 80, detail of genitalia at high magnification; 81,cornutus at high magnification, slide Bldz 9608; 82, cornutus, slide Bldz 9234; 83, abdomen.

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79 80

81

83 82

Figs. 84-86. Coleophora niphomesta, slide Bldz 8699. – 84, Female genitalia; 85, sterigma at high magnification; 86, abdomen.

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85

84 86

Figs. 87-89. Coleophora chordoscelis, slide BMNH 24474, lectotype. – 87, Female genitalia; 88, sterigma at high magnification;89, abdomen.

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88

87 89