2 Artificial Selection

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    Artificial Selection

    Domestication & BreedDevelopment

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    To Succeed with Domestication

    Adapt to environment

    Provide nutrition needs

    Meet social requirementsMust consider typical social organization of species

    How to young learn to be adults

    Problems

    Vices, Aggression Impotence, Infanticide

    Able to reproduce

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    Domestication of the Horse

    23,000 B.C.Southern France archeological site

    Food cracked bones of as many as 100,000horses

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    Start of Primitive Agriculture

    4500-3500 B.C. Less nomadic Depletion of local herds Domestication

    Means of dependable food supply Taming by-product of hunting Orphaned foal 2-12 months of age Adopt, raise and put to work

    Ex. Turkestan Turkmenes Lived on mares milk and horse meat - Raised fillies/ate colts Tethered mares in season for wild stallions to breed Earliest selection criteria Milk Production

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    Domestication of the Horse

    Ass, Onager & Camel

    domesticated before the

    horse 3000 B.C. drawings of

    Onager being ridden with a

    nose ring and jaw strap

    Horse domesticated in

    numerous areas duringsimilar time period 3000-2300 B.C.

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    Signs of Domestication

    Caves of Sumeria (Perisian Gulf) 2800 B.C. First signs of selective breeding

    Drawings of horse heads in 4 horizontal rows Different head profile types (convex, straight,

    concave) and mane types (erect, hanging and

    maneless)

    Central Asia 2800 B.C. Drawings of horses hitched wearing snaffle-type bits

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    Earliest Selective Breeding

    TemperamentThose that refused to obey returned to wild or

    used for meatMore agreeable animals remained in

    comparative safety of captivity

    Bred to other docile animals to produce good-

    tempered foals

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    Earliest Selection Criteria

    Learning Ability - Trainability Horse transferred reliance to man

    Food, protection and companionship Learned to overcome fears hunting lions, tigers, etc.

    Xenophon 430 B.C. first recorded advocate of

    training by reward.

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    Other Traits

    Speed and SurvivabilityWithhold water for several days

    Release herd pointed in the direction of water

    First to arrive were selected for breeding and

    training

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    Man Alters the Horse

    Orthopedic Problems1000 B.C. Xenophon first asserted no foot,

    no horse

    500 B.C. Persians develop first horseshoes heavy cloth Other problemsOccurrence of genetic abnormalities

    May select beauty over functionality Small, dainty hooves

    Selection criteria for performance reducedversatility

    Breeds

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    Breed Development

    Breed = group with distinctivecharacteristics that are transmitted to their

    offspring. Initial development through natural

    crosses and artificial selectionEnvironment important roleAdaptations of the 4 original types

    Results in many breeding groups

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    Breed Development

    Distinctive Characteristics Extremes in Climate, Topography and Vegetation Conformation

    Color Performance

    Skills and Abilities well promoted by breed advocates

    Intelligence

    Disposition Suitable to Mans Purpose gaits

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    Formal Development

    Man bred early types to fix certaindesirable characteristicsLittle change if type was suitable

    pony breeds are relatively pure almost directdescendants of one early type

    Some breeds took centuries Outcrossing with numerous other breeds

    Easiest to develop if trait can be fixed (set)

    Get desirable and undesirable characteristics set One sire may have strong prepotence good and bad

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    Genetic Evidence

    DNA of ancient wild horses found in permafrost Sweden & Estonia 2000yr old remains, Alaska

    28000 yr old remains Analyzed the mitochondria DNA

    Mutations occur at a known rate = molecular clock

    Maternal ancestry traced

    Tested against 600 living horses From 25 breeds worldwide

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