Fig. 33-1

Overview: Life Without a Backbone

• Invertebrates are animals that lack a backbone (ansence NOT a good character)

• They account for 95% of known animal species

• Is it a natural (monopyletic group)?

• i.e. clade or grade?

Fig. 33-2

ANCESTRALPROTIST

Commonancestor ofall animals

Calcareaand Silicea

Cnidaria

Lophotrochozoa

Ecdysozoa

Deuterostomia

Major groups

“Porifera” (grade)

Fig. 33-3-1Calcarea and Silicea (5,500 species)

A sponge

Cnidaria (10,000 species)

A jelly

Acoela (400 species)

Acoel flatworms (LM)

1.5 mm

LOPHOTROCHOZOANS

Platyhelminthes (20,000 species)

A marine flatworm

Ectoprocta (4,500 species)

Ectoprocts A brachiopod

Brachiopoda (335 species)

Rotifera (1,800 species)

A rotifer (LM)

A ctenophore, or comb jelly

Ctenophora (100 species)

A placozoan (LM)

0.5 mm

Placozoa (1 species)

Fig. 33-3-2

An acanthocephalan (LM)

Acanthocephala (1,100 species)

Nemertea (900 species)

A ribbon worm

ECDYSOZOA

Loricifera (10 species) Priapula (16 species)

A loriciferan (LM) A priapulan

50 µm

Cycliophora (1 species)

Mollusca (93,000 species)

A cycliophoran (colorized SEM)

100 µm

An octopus

Annelida (16,500 species)

A marine annelid

Fig. 33-3-3Tardigrada (800 species)

Nematoda (25,000 species)

Onychophora (110 species)

Arthropoda (1,000,000 species)

DEUTEROSTOMIA

Hemichordata (85 species)

Echinodermata (7,000 species) Chordata (52,000 species)

Tardigrades (colorized SEM)

100 µm

A roundworm

An acorn worm

A sea urchin

An onychophoran

A scorpion(an arachnid)

A tunicate

Fig. 33-3a

A sponge

porifera

Fig. 33-3b

A jelly

Cnidaria

Fig. 33-3c

A placozoan (LM)

0.5 mm

Fig. 33-3d

A ctenophore, or comb jelly

Fig. 33-3e

Acoel flatworms (LM)

1.5 mm

Fig. 33-3f

A marine flatworm

Lophotrochozoa

Fig. 33-3g

A rotifer (LM)

Lophotrochozoa

Fig. 33-3h

Ectoprocts

Lophotrochozoa

Fig. 33-3i

A brachiopod

Lophotrochozoa

Fig. 33-3j

An acanthocephalan (LM)

Lophotrochozoa

Fig. 33-3k

A cycliophoran (colorized SEM)

100 µm

Lophotrochozoa

Fig. 33-3l

A ribbon worm

Lophotrochozoa

Fig. 33-3m

An octopus

Lophotrochozoa

Fig. 33-3n

A marine annelid

Lophotrochozoa

Fig. 33-3o

A loriciferan (LM)

50 µm

Ecdysozoa

Fig. 33-3p

A priapulan

Ecdysozoa

Fig. 33-3q

Tardigrades (colorized SEM)

100 µm

Ecdysozoa

Fig. 33-3s

A roundworm

Ecdysozoa

Fig. 33-3t

A scorpion(an arachnid)

Ecdysozoa

Fig. 33-3u

An acorn worm

Deuterostome

Fig. 33-3v

A sea urchin

Deuterostome

Fig. 33-3w

A tunicate

Deuterostome

Closer!

Concept 33.1: Sponges are basal animals that lack true tissues

• Sponges are sedentary animals from two phyla: Calcarea and Silicea

• They live in both fresh and marine waters

• Sponges lack true tissues and organs

Fig. 33-UN1

Calcarea and Silicea

CnidariaLophotrochozoa

Ecdysozoa

Deuterostomia

• Sponges are suspension feeders, capturing food particles suspended in the water that pass through their body

• Choanocytes, flagellated collar cells, generate a water current through the sponge and ingest suspended food

• Water is drawn through pores into a cavity and out through an opening

Fig. 33-4

Azure vase sponge (Callyspongiaplicifera)

Spongocoel

Osculum

Epidermis Waterflow

Mesohyl

Choanocyte

Flagellum

Collar

Food particlesin mucus

Choanocyte

AmoebocytePhagocytosis offood particles

Spicules

Amoebocytes

• Sponges consist of a noncellular mesohyl layer between two cell layers

• Amoebocytes are found in the mesohyl and play roles in digestion and structure

• Most sponges are hermaphrodites: Each individual functions as both male and female

Lophotrochozoans, a clade identified by molecular data, have the widest range of animal body forms

• Bilaterian animals have bilateral symmetry and triploblastic development

• The clade Bilateria contains Lophotrochozoa, Ecdysozoa, and Deuterostomia

• Look at your tree!

Fig. 33-UN3

Ecdysozoa

Deuterostomia

• The clade Lophotrochozoa was identified by molecular data

• Some develop a lophophore for feeding, others pass through a trochophore larval stage, and a few have neither feature

• Lophotrochozoa includes the flatworms, molluscs, and many other important phyla

Flatworms

• Members of phylum Platyhelminthes live in marine, freshwater, and damp terrestrial habitats

• They are flattened dorsoventrally and have a gastrovascular cavity

• Gas exchange takes place across the surface

Turbellarians

• Turbellarians are nearly all free-living and mostly marine

• The best-known turbellarians are commonly called planarians

Fig. 33-9

Fig. 33-10

Pharynx

Gastrovascularcavity

Eyespots

Ganglia Ventral nerve cords

Monogeneans and Trematodes

• Monogeneans and trematodes live as parasites in or on other animals

• They parasitize a wide range of hosts, and most have complex life cycles

• Trematodes that parasitize humans spend part of their lives in snail hosts

• Most monogeneans are parasites of fish

Human host

Motile larva

Snail host

Ciliated larva

Female

Blood fluke=schistomiasis

Tapeworms

• Tapeworms are parasites of vertebrates and lack a digestive system

• Tapeworms absorb nutrients from the host’s intestine

• Fertilized eggs, produced by sexual reproduction, leave the host’s body in feces

Fig. 33-12

Proglottids withreproductive structures

Sucker

Scolex

200 µm

Molluscs

• Phylum Mollusca includes snails and slugs, oysters and clams, and octopuses and squids

• Most molluscs are marine, though some inhabit fresh water and some are terrestrial

• Molluscs are soft-bodied animals, but most are protected by a hard shell

• Most molluscs have separate sexes with gonads located in the visceral mass

• The life cycle of many molluscs includes a ciliated larval stage called a trochophore

• There are four major classes of molluscs:

Chitons

• oval-shaped marine animals encased in an armor of eight dorsal plates

Fig. 33-16

Gastropods

• About three-quarters of all living species of molluscs are gastropods

Video: NudibranchsVideo: Nudibranchs

Fig. 33-17

(a) A land snail

(b) A sea slug

Fig. 33-17a

(a) A land snail

Fig. 33-17b

(b) A sea slug

• Most gastropods are marine, but many are freshwater and terrestrial species

• Most have a single, spiraled shell

• Slugs lack a shell or have a reduced shell

• The most distinctive characteristic of gastropods is torsion, which causes the animal’s anus and mantle to end up above its head

Fig. 33-18

Mantlecavity

Stomach Intestine

Bivalves

• Molluscs of class Bivalvia include many species of clams, oysters, mussels, and scallops

• They have a shell divided into two halves

Fig. 33-19

Fig. 33-20

Digestivegland

MantleHinge area

Coelom

Heart Adductormuscle

Excurrentsiphon

Waterflow

IncurrentsiphonGillGonad

Mantlecavity

Cephalopods

• Class Cephalopoda includes squids and octopuses, carnivores with beak-like jaws surrounded by tentacles of their modified foot

• Most octopuses creep along the sea floor in search of prey

Fig. 33-21

Octopus

Chambered nautilus

Fig. 33-21a

Octopus

• Squids use their siphon to fire a jet of water, which allows them to swim very quickly

Fig. 33-21b

• One small group of shelled cephalopods, the nautiluses, survives today

Fig. 33-21c

Chamberednautilus

Nematodes

• Nematodes, or roundworms, are found in most aquatic habitats, in the soil, in moist tissues of plants, and in body fluids and tissues of animals

• They have an alimentary canal, but lack a circulatory system

• Reproduction in nematodes is usually sexual, by internal fertilization

Video: Video: C.C. eleganselegans Crawling Crawling

Video: Video: C.C. eleganselegans Embryo Development (Time Lapse) Embryo Development (Time Lapse)

Fig. 33-25

25 µm

• Some species of nematodes are important parasites of plants and animals

Fig. 33-26

Encysted juveniles Muscle tissue 50 µm

Trichinella

Arthropods

• Two out of every three known species of animals are arthropods

• Members of the phylum Arthropoda are found in nearly all habitats of the biosphere

General Characteristics of Arthropods

• The appendages of some living arthropods are modified for many different functions

Video: Lobster Mouth PartsVideo: Lobster Mouth Parts

Fig. 33-29

Cephalothorax

Antennae(sensoryreception) Head

Thorax

Abdomen

Swimming appendages(one pair locatedunder eachabdominal segment)

Walking legs

Mouthparts (feeding)Pincer (defense)

• The body of an arthropod is completely covered by the cuticle, an exoskeleton made of layers of protein and the polysaccharide chitin (like fungus!)

• When an arthropod grows, it molts its exoskeleton

• Arthropods have an open circulatory system in which fluid called hemolymph is circulated into the spaces surrounding the tissues and organs

• A variety of organs specialized for gas exchange have evolved in arthropods

• Molecular evidence suggests that living arthropods consist of four major lineages that diverged early in the phylum’s evolution:

– Cheliceriforms (sea spiders, horseshoe crabs, scorpions, ticks, mites, and spiders)

– Myriapods (centipedes and millipedes)

– Hexapods (insects and relatives)

– Crustaceans (crabs, lobsters, shrimps, barnacles, and many others)

Cheliceriforms

• Cheliceriforms, subphylum Cheliceriformes, are named for clawlike feeding appendages called chelicerae

Fig. 33-30

• Most modern cheliceriforms are arachnids, which include spiders, scorpions, ticks, and mites

Fig. 33-31

Scorpion

Dust mite

Web-building spider

50 µm

Fig. 33-31a

Scorpion

Fig. 33-31b

Dust mite

50 µm

Fig. 33-31c

Web-building spider

Myriapods

• Subphylum Myriapoda includes millipedes and centipedes (many feet!)

Fig. 33-33

Fig. 33-34

Insects

• Subphylum Hexapoda, insects and relatives, has more species than all other forms of life combined

• They live in almost every terrestrial habitat and in fresh water

• The internal anatomy of an insect includes several complex organ systems

Fig. 33-35

Abdomen Thorax Head

Compound eye

Antennae

Dorsalartery Crop

Cerebral ganglion

Mouthparts

Nerve cords

Tracheal tubesOvary

Malpighiantubules

Vagina

• Insects diversified several times following the evolution of flight, adaptation to feeding on gymnosperms, and the expansion of angiosperms

• Insect and plant diversity declined during the Cretaceous extinction, but have been increasing in the 65 million years since

• Flight is one key to the great success of insects

• An animal that can fly can escape predators, find food, and disperse to new habitats much faster than organisms that can only crawl

• Many insects undergo metamorphosis during their development

• In incomplete metamorphosis, the young, called nymphs, resemble adults but are smaller and go through a series of molts until they reach full size

• Insects with complete metamorphosis have larval stages known by such names as maggot, grub, or caterpillar

• The larval stage looks entirely different from the adult stage

Video: Butterfly EmergingVideo: Butterfly Emerging

Fig. 33-36

(a) Larva (caterpillar)(b) Pupa

(c) Later-stage pupa (d) Emerging

(e) Adult

Fig. 33-36a

(a) Larva (caterpillar)

Fig. 33-36b

(b) Pupa

Fig. 33-36c

(c) Later-stage pupa

Fig. 33-36d

(d) Emerging adult

Fig. 33-36e

(e) Adult

• Most insects have separate males and females and reproduce sexually

• Individuals find and recognize members of their own species by bright colors, sound, or odors

• Some insects are beneficial as pollinators, while others are harmful as carriers of diseases, or pests of crops

• Insects are classified into more than 30 orders

Video: Bee PollinatingVideo: Bee Pollinating

Fig. 33-37

Crustaceans

• While arachnids and insects thrive on land, crustaceans, for the most part, have remained in marine and freshwater environments

• Crustaceans, subphylum Crustacea, typically have branched appendages that are extensively specialized for feeding and locomotion

• Most crustaceans have separate males and females

• Isopods include terrestrial, freshwater, and marine species

– Pill bugs are a well known group of terrestrial isopods

• Decapods are all relatively large crustaceans and include lobsters, crabs, crayfish, and shrimp

Fig. 33-38

(a) Ghost crab

(b) Krill (c) Barnacles

Fig. 33-38a

(a) Ghost crab

• Planktonic crustaceans include many species of copepods, which are among the most numerous of all animals

Fig. 33-38b

(b) Krill

• Barnacles are a group of mostly sessile crustaceans

• They have a cuticle that is hardened into a shell

Fig. 33-38c

(c) Barnacles

Concept 33.5: Echinoderms and chordates are deuterostomes

• Sea stars and other echinoderms, phylum Echinodermata, may seem to have little in common with phylum Chordata, which includes the vertebrates

• Shared characteristics define deuterostomes (Chordates and Echinoderms)

– Radial cleavage

– Formation of the mouth at the end of the embryo opposite the blastopore

Fig. 33-UN5

Ecdysozoa

Deuterostomia

Fig. 33-1

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